Full reference: A. G. Ponomarenko. 1969. Istoricheskoe Razvitie Zhestkokrylykh-Arkhostemat [Historical Development of the Archostomate Beetles]. Trudy Akademiya Nauk SSSR 125:1-240
See also Ponomarenko 1969
Sister taxa: none
Type specimen: PIN 2496/7, an exoskeleton. Its type locality is Kenderlyk, Akkolka River (PIN collection 2496), which is in a Norian terrestrial siliciclastic in the Tologoi Formation of Kazakhstan.
Distribution: found only at Kenderlyk, Akkolka River (PIN collection 2496)
Specimen images are retrieved through the ePANDDA API.
Click image to enlarge. Click to access iDigBio record.