Full reference: A. G. Ponomarenko. 1969. Istoricheskoe Razvitie Zhestkokrylykh-Arkhostemat [Historical Development of the Archostomate Beetles]. Trudy Akademiya Nauk SSSR 125:1-240
Sister taxa: none
Type specimen: PIN 2069/1237, an exoskeleton. Its type locality is Madygen, PIN collection 2069, Dzhailoucho area, which is in a Carnian lacustrine - small siliciclastic in the Madygen Formation of Kyrgyzstan.
Distribution: found only at Madygen, PIN collection 2069, Dzhailoucho area
Specimen images are retrieved through the ePANDDA API.
Click image to enlarge. Click to access iDigBio record.