Insecta - Coleoptera - Scarabaeidae
Alternative combination: Proteroscarabaeus crassus
Full reference: A. G. Ponomarenko. 1990. Beetles, Scarabaeida, in Pozdne-Mezozoyskie Nasekomye Vostochnogo Zabaykal'ya. Akademiya Nauk SSSR, Trudy Paleontologicheskogo Instituta 239:39-82
Belongs to Lithanomala according to G. V. Nikolajev 2007
See also Krell 2000, Nikolajev 1992 and Ponomarenko 1990
Sister taxa: Lithanomala oblonga, Lithanomala sibirica
Type specimen: PIN 4085/12, an exoskeleton. Its type locality is Leskovo (PIN collection 4085), which is in a Barremian lacustrine - small horizon in the Leskovskaya Formation of the Russian Federation.
Ecology:
Distribution: found only at Leskovo (PIN collection 4085)
Specimen images are retrieved through the ePANDDA API.
Click image to enlarge. Click to access iDigBio record.