Insecta - Coleoptera - Carabidae
Alternative spellings: Pterostichii, Pterostichinae
Full reference: F. A. Bonelli. 1810. Observations Entomologiques. Première Partie (Cicindélètes et Portion des Carabiques) 1-58
Parent taxon: Harpalinae according to V. Tamutis et al. 2011
See also Bouchard et al. 2011, Hayashi et al. 2005 and LeConte 1861
Sister taxa: Abacetini, Aethocarabus, Chlaeniini, Dryptini, Galeritini, Harpalini, Helluonini, Lebiini, Licinini, Odacanthini, Panagaeini, Pseudomorphini, Sinis, Sphodrini, Zabrini, Zuphiini, Oodini
Subtaxa: Cyclotrachelus Pterostichina
Type: Pterostichus
Distribution:
• Quaternary of Canada (4: Nunavut, Ontario collections), Greenland (8), Japan (7), New Zealand (1), the Russian Federation (49), Ukraine (1), the United Kingdom (10), United States (21: Alaska, California, Colorado, Ohio, Pennsylvania, Washington, Wisconsin)
• Pliocene of Japan (2), the Russian Federation (1)
• Miocene of Germany (2), the Russian Federation (2), United States (2: Alaska, Washington)
• Oligocene of France (2), Germany (1)
• Eocene of Germany (1), the United Kingdom (1), United States (4: Colorado)
Total: 119 collections including 248 occurrences