Reptilia - Testudines - Testudinidae

Full reference: S. Gmira, F. de Lapparent de Broin, D. Geraads, D. Levefre, A. Mohib and J.-P. Raynal. 2013. Les tortues du Pliocène d’Ahl al Oughlam (Casablanca, Maroc) et de localités mio-pliocènes avoisinantes. Geodiversitas 35(3):691-733

Belongs to Testudo according to G. L. Georgalis et al. 2021

See also Gmira et al. 2013 and Lujan et al. 2016

Sister taxa: Testudo (Chersine), Testudo (Stylemys), Testudo amberiacensis, Testudo amiatae, Testudo antakyensis, Testudo antiqua, Testudo atlas, Testudo bosporica, Testudo brevitesta, Testudo brontops, Testudo carinata, Testudo cautleyi, Testudo cernovi, Testudo changshanensis, Testudo chienfutungensis, Testudo costarricensis, Testudo craverii, Testudo denizoti, Testudo despotti, Testudo emys, Testudo escheri, Testudo eurysternum, Testudo floweri, Testudo geometrica, Testudo globosa, Testudo graeca, Testudo hellenica, Testudo hermanni, Testudo hexagonata, Testudo hipparionum, Testudo honanensis, Testudo houzei, Testudo hypercostata, Testudo hypselonota, Testudo ibera, Testudo indica, Testudo kalksburgensis, Testudo kenitrensis, Testudo kleinmanni, Testudo lamanonii, Testudo larteti, Testudo lohanica, Testudo lunanensis, Testudo lutaria, Testudo margae, Testudo marginata, Testudo marmorum, Testudo nanus, Testudo neraudii, Testudo pecorinii, Testudo phosphoritarum, Testudo promarginata, Testudo pusilla, Testudo scabra, Testudo shansiensis, Testudo sharanensis, Testudo shensiensis, Testudo shilouensis, Testudo sloanei, Testudo sphaerica, Testudo steinheimensis, Testudo tarakliensis, Testudo terrestris, Testudo tunhuanensis, Testudo ulanensis, Testudo weissingeri, Testudo yunnanensis

Type specimen: INSAP. AaO-272, a shell. Its type locality is Ahl al Oughlam - general, which is in a Pliocene coastal limestone/conglomerate in Morocco.

Ecology: ground dwelling herbivore-herbivore

Distribution: found only at Ahl al Oughlam - general

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Specimen images are retrieved through the ePANDDA API.