Mammalia - Carnivora - Ursidae
Parent taxon: Ursidae according to R. M. Hunt 1998
See also Bjork 1970, Jin et al. 2007 and Kurten and Anderson 1980
Sister taxa: Agriotheriinae, Ailuropodinae, Amphicynodontinae, Eoarctos, Harpaleocyon, Hemicyoninae, Metarctos, Myadina, Phoberocyoninae, Proarctotherium, Taxina, Tremarctinae, Ursulus
Subtaxa: Arctotheriini Aurorarctos Ballusia Melursus Protarctos Tremarctini Ursavini Ursini
Ecology: ground dwelling herbivore-carnivore
Distribution:
• Quaternary of Argentina (9 collections), Austria (1), Belgium (2), Bolivia (4), Brazil (6), Canada (58: British Columbia, Manitoba, Newfoundland, Northwest Territories, Nunavut, Ontario, Quebec, Saskatchewan, Yukon), Chile (2), China (20), Croatia (2), Czechia (3), Denmark (1), France (25), Georgia (2), Germany (21), Greece (16), Greenland (33), Hungary (6), India (2), Indonesia (1), Israel (3), Italy (37), Japan (4), Kazakhstan (1), Laos (1), Libya (1), Malaysia (2), Mexico (6), Monaco (3), Mongolia (1), Morocco (4), the Netherlands (4), North Atlantic (1), North Pacific (1), Norway (8), Portugal (4), Romania (5), the Russian Federation (10), Spain (24), Svalbard and Jan Mayen (1), Sweden (10), Taiwan (1), Thailand (2), Türkiye (3), the United Kingdom (13), United States (168: Alabama, Alaska, Arkansas, California, Colorado, Florida, Georgia, Idaho, Illinois, Indiana, Iowa, Kansas, Kentucky, Louisiana, Maine, Maryland, Massachusetts, Michigan, Mississippi, Missouri, Montana, Nebraska, Nevada, New Mexico, New York, Ohio, Oregon, Pennsylvania, South Carolina, South Dakota, Tennessee, Texas, Utah, Virginia, Washington, West Virginia, Wyoming), Venezuela (3), Vietnam (1)
• MN 16 of France (2), Italy (1)
• Villafranchian of Greece (2), Hungary (4), Spain (1)
• Blancan of El Salvador (1), Mexico (1), United States (14: Arizona, California, Florida, Idaho, Nevada, New Mexico, Texas, Washington)
• Pliocene to Pleistocene of China (1), France (2), Greece (2), Hungary (1), Italy (2), Türkiye (1), Venezuela (1)
• Pliocene of Bulgaria (1), Canada (1: Northwest Territories), China (1), France (1), Georgia (2), Greece (4), Hungary (2), Italy (1), Moldova (3), Morocco (1), Poland (1), Romania (3), the Russian Federation (3), Spain (1), Tajikistan (4), Ukraine (2), the United Kingdom (2)
• Turolian of China (4), Türkiye (1)
• Hemphillian of United States (7: California, Florida, Indiana, Nebraska, Oregon, Tennessee)
• Miocene of Bosnia and Herzegovina (1), Canada (1: Saskatchewan), China (1), France (10), Germany (2), Greece (1), Hungary (1), Italy (1), Poland (1), Slovakia (2), Switzerland (1), Türkiye (2), United States (9: Colorado, Nebraska, Nevada, Oregon, South Dakota)
• Neogene of Italy (1)
• Arikareean of United States (1: Oregon)
Total: 652 collections including 727 occurrences