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Marginocephalia
Taxonomy
Marginocephalia was named by Sereno (1986) [Listed as "Hypoorder"]. It is not extant. It was considered monophyletic by Butler et al. (2008) and Butler and Sullivan (2009).
It was assigned to Heterodontosauria by Olshevsky and Ford (1994); to Neornithischia by Sereno (1997), Sereno (1998) and Sereno (1999); to Ornithischia by Lambert et al. (2001); to Chasmatopia by Buchholz (2002); to Heterodontosauriformes by Xu et al. (2006); to Neornithischia by Arbour and Evans (2019); to Cerapoda by Sereno (1986), Benton (1990), Weishampel (2004), Butler et al. (2007), Butler et al. (2008), Butler and Sullivan (2009), Han et al. (2012), Madzia et al. (2018), Herne et al. (2019) and Dieudonné et al. (2020); and to Ornithischia by Dodson (1997), Rauhut (2003) and Tanaka et al. (2024).
It was assigned to Heterodontosauria by Olshevsky and Ford (1994); to Neornithischia by Sereno (1997), Sereno (1998) and Sereno (1999); to Ornithischia by Lambert et al. (2001); to Chasmatopia by Buchholz (2002); to Heterodontosauriformes by Xu et al. (2006); to Neornithischia by Arbour and Evans (2019); to Cerapoda by Sereno (1986), Benton (1990), Weishampel (2004), Butler et al. (2007), Butler et al. (2008), Butler and Sullivan (2009), Han et al. (2012), Madzia et al. (2018), Herne et al. (2019) and Dieudonné et al. (2020); and to Ornithischia by Dodson (1997), Rauhut (2003) and Tanaka et al. (2024).
Synonymy list
| Year | Name and author |
|---|---|
| 1986 | Marginocephalia Sereno |
| 1990 | Marginocephalia Benton p. 19 |
| 1994 | Marginocephalia Olshevsky and Ford p. 91 |
| 1997 | Marginocephalia Dodson p. 474 |
| 1997 | Marginocephalia Sereno p. 443 |
| 1998 | Marginocephalia Sereno p. 62 |
| 1999 | Marginocephalia Sereno p. 2138 fig. 2 |
| 2001 | Marginocephalia Lambert et al. p. 7 |
| 2002 | Marginocephalia Buchholz p. 20 |
| 2003 | Marginocephalia Rauhut p. 16 |
| 2004 | Marginocephalia Weishampel p. 323 |
| 2006 | Marginocephalia Xu et al. p. 2139 fig. 3 |
| 2007 | Marginocephalia Butler et al. p. 2044 |
| 2008 | Marginocephalia Butler et al. p. 22 figs. 2-4 |
| 2009 | Marginocephalia Butler and Sullivan p. 22 |
| 2012 | Marginocephalia Han et al. p. 1391 fig. 14 |
| 2018 | Marginocephalia Madzia et al. p. 973 fig. 4 |
| 2019 | Marginocephalia Arbour and Evans |
| 2019 | Marginocephalia Herne et al. p. 573 |
| 2020 | Marginocephalia Dieudonné et al. |
| 2024 | Marginocephalia Tanaka et al. p. 5 |
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If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.
Unr. †Marginocephalia Sereno 1986
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Unr. †Ceratopsia Marsh 1890 [ceratopsian]
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G. †Albalophosaurus Ohashi and Barrett 2009
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†Albalophosaurus yamaguchiorum Ohashi and Barrett 2009
Fm. †Ceratopsidae Marsh 1888 [ceratopsid]
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Subfm. †Centrasaurinae Lambe 1915
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Tr. †Albertaceratopsini Loewen et al. 2024
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G. †Albertaceratops Ryan 2007
G. †Lokiceratops Loewen et al. 2024
G. †Medusaceratops Ryan et al. 2010
Subfm. †Centrosaurinae Lambe 1915
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Unr. †Centrosaurini Ryan et al. 2017
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G. †Centrosaurus Lambe 1905
G. †Coronosaurus Ryan et al. 2012
G. †Rubeosaurus McDonald and Horner 2010
G. †Spinops Farke et al. 2011
G. †Styracosaurus Lambe 1913
G. †Monoclonius Cope 1876
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Invalid names: Monoclonius fissus Cope 1889 [nomen dubium], Monoclonius recurvicornis Cope 1889 [nomen dubium], Monoclonius sphenocerus Cope 1889 [nomen dubium]
Unr. †Nasutoceratopsini Ryan et al. 2017
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G. †Avaceratops Dodson 1986
G. †Crittendenceratops Dalman et al. 2018
G. †Furcatoceratops Ishikawa et al. 2023
G. †Nasutoceratops Sampson et al. 2013
G. †Yehuecauhceratops Rivera-Sylva et al. 2017
Unr. †Pachyrhinosaurini Fiorillo and Tykoski 2012
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G. †Achelousaurus Sampson 1994
G. †Einiosaurus Sampson 1994
G. †Pachyrhinosaurus Sternberg 1950
Invalid names: Pachyrostra Fiorillo and Tykoski 2012 [empty]
Invalid names: Brachyceratops Gilmore 1914 [nomen dubium]
Subfm. †Chasmosaurinae Lambe 1915
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G. †Agujaceratops Lucas et al. 2006 [Aguja horned face]
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†Agujaceratops mariscalensis Lehman 1989
†Agujaceratops mavericus Lehman et al. 2017
G. †Anchiceratops Brown 1914
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†Anchiceratops ornatus Brown 1914
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Invalid names: Anchiceratops longirostris Sternberg 1926 [synonym]
G. †Chasmosaurus Lambe 1914
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†Chasmosaurus belli Lambe 1902
†Chasmosaurus russelli Sternberg 1940
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Invalid names: Mojoceratops perifania Longrich 2010 [synonym]
Invalid names: Chasmosaurus brevirostris Lull 1933 [nomen dubium], Chasmosaurus kaiseni Brown 1933 [nomen dubium], Eoceratops Lambe 1915 [synonym], Mojoceratops Longrich 2010 [synonym], Monoclonius canadensis Lambe 1902 [nomen dubium], Protorosaurus Lambe 1914 [synonym]
G. †Navajoceratops Fowler and Freedman Fowler 2020
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†Navajoceratops sullivani Fowler and Freedman Fowler 2020
G. †Pentaceratops Osborn 1923
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†Pentaceratops sternbergii Osborn 1923
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Invalid names: Pentaceratops fenestratus Wiman 1930 [synonym]
G. †Terminocavus Fowler and Freedman Fowler 2020
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†Terminocavus sealeyi Fowler and Freedman Fowler 2020
Unr. †Triceratopsini Longrich 2011
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G. †Eotriceratops Wu et al. 2007
G. †Regaliceratops Brown and Henderson 2015
G. †Titanoceratops Longrich 2011
G. †Torosaurus Marsh 1891
G. †Triceratops Marsh 1889
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Invalid names: Nedoceratops Ukrainsky 2007 [synonym], Ojoceratops Sullivan and Lucas 2010 [synonym], Sterrholophus Marsh 1891 [synonym], Tatankaceratops Ott and Larson 2010 [synonym]
G. †Vagaceratops Sampson 2010
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†Vagaceratops irvinensis Holmes et al. 2001
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Invalid names: Mercuriceratops gemini Ryan et al. 2014 [synonym], Pentaceratops aquilonius Longrich 2014 [synonym]
Invalid names: Mercuriceratops Ryan et al. 2014 [synonym]
Invalid names: Ceratops Marsh 1888 [nomen dubium]
Invalid names: Agathaumas Cope 1872 [nomen dubium], Agathaumidae Cope 1889 [synonym], Bison alticornis Marsh 1887 [nomen dubium], Ceratopsinae Abel 1919 [empty], Monocloniinae Nopcsa 1923 [empty], Polyonax Cope 1874 [nomen dubium], Triceratops galeus Marsh 1889 [nomen dubium], Triceratops ingens Lull 1915 [nomen dubium], Triceratops maximus Brown 1933 [nomen dubium], Ugrosaurus CoBabe and Fastovsky 1987 [nomen dubium]
Infraor. †Neoceratopsia Sereno 1986
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G. †Archaeoceratops Dong and Azuma 1997
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†Archaeoceratops oshimai Dong and Azuma 1997
†Archaeoceratops yujingziensis You et al. 2010
Fm. †Chaoyangsauridae Zhao 1983
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G. †Chaoyangsaurus Zhao et al. 1999
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†Chaoyangsaurus youngi Zhao et al. 1999
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Invalid names: Chaoyangosaurus liaosiensis Zhao and Cheng 1983 [replaced]
Invalid names: Chaoyangosaurus Zhao and Cheng 1983 [replaced]
Invalid names: Xuanhuasaurus niei Zhao 1985 [nomen dubium]
Unr. †Coronosauria Sereno 1986
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Superfm. †Ceratopsoidea Hay 1902
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G. †Ajkaceratops Osi et al. 2010
G. †Turanoceratops Nessov et al. 1989
G. †Zuniceratops Wolfe and Kirkland 1998
Fm. †Protoceratopsidae Granger and Gregory 1923
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G. †Bagaceratops Maryanska and Osmólska 1975
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Invalid names: Gobiceratops Alifanov 2008 [synonym], Lamaceratops Alifanov 2003 [synonym], Magnirostris You and Dong 2003 [synonym], Platyceratops Alifanov 2003 [synonym]
G. †Breviceratops Kurzanov 1990
G. †Protoceratops Granger and Gregory 1923
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Invalid names: Bainoceratops Tereshchenko and Alifanov 2003 [synonym]
Invalid names: Asiaceratopsinae Nessov et al. 1989 [empty]
Invalid names: Bagaceratopsidae Alifanov 2003 [empty]
Fm. †Leptoceratopsidae Nopcsa 1923 [leptoceratopsid]
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Invalid names: Archaeoceratopsidae Dong and Azuma 1997 [empty], Ceratopsomorpha Wolfe and Kirkland 1998 [empty], Protoceratopoidea Tereshchenko 2007 [empty]
Fm. †Psittacosauridae Osborn 1923
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G. †Xuanhuasaurus Zhao 1985
Invalid names: Protiguanodontidae Osborn 1923 [synonym], Stenopelyxidae Nopcsa 1927 [synonym]
G. †Psittacosaurus Osborn 1923
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†Psittacosaurus amitabha Napoli et al. 2019
†Psittacosaurus gobiensis Sereno et al. 2010
†Psittacosaurus lujiatunensis Zhou et al. 2006
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Invalid names: Psittacosaurus houi You et al. 2003 [synonym], Psittacosaurus major Sereno et al. 2007 [synonym]
†Psittacosaurus meileyingensis Sereno et al. 1988
†Psittacosaurus mongoliensis Osborn 1923
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Invalid names: Protiguanodon mongoliense Osborn 1923 [synonym], Psittacosaurus guyangensis Cheng 1982 [synonym], Psittacosaurus osborni Young 1932 [synonym], Psittacosaurus protiguanodonensis Young 1958 [synonym], Psittacosaurus tingi Young 1932 [synonym]
†Psittacosaurus neimongoliensis Russell and Zhao 1996
†Psittacosaurus sibiricus Voronkevich and Averianov 2000
†Psittacosaurus xinjiangensis Sereno and Chao 1988
Invalid names: Hongshanosaurus You et al. 2003 [synonym], Protiguanodon Osborn 1923 [synonym], Psittacosaurus mazongshanensis Xu 1997 [nomen dubium], Psittacosaurus ordosensis Russell and Zhao 1996 [nomen dubium]
G. †Serendipaceratops Rich and Vickers-Rich 2003
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†Serendipaceratops arthurcclarkei Rich and Vickers-Rich 2003
Invalid names: Chaoyangosauridae Dong 1992 [empty], Dysganus Cope 1876 [nomen dubium], Eoceratopsinae Lambe 1915 [empty], Kulceratops Nessov 1995 [nomen dubium], Kulceratops kulensis Nessov 1995 [nomen dubium], Microceratus Mateus 2008 [nomen dubium], Pachyrhinosauridae Sternberg 1950 [empty], Ponderopoda Huene 1948 [synonym], Psittacosauroidea Zhao 1983 [empty], Psittacosaurus sattayaraki Buffetaut and Ingavat 1992 [nomen dubium]
Unr. †Pachycephalosauria Maryańska and Osmólska 1974
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Unr. †Eupachycephalosauria Dieudonné et al. 2020
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G. †Homalocephale Maryańska and Osmólska 1974
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†Homalocephale calathocercos Maryańska and Osmólska 1974
G. †Pachycephalosaurus Brown and Schlaikjer 1943
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†Pachycephalosaurus wyomingensis Gilmore 1931
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Invalid names: Dracorex hogwartsia Bakker et al. 2006 [synonym], Pachycephalosaurus grangeri Brown and Schlaikjer 1943 [synonym], Pachycephalosaurus reinheimeri Brown and Schlaikjer 1943 [synonym], Stygimoloch spinifer Galton and Sues 1983 [synonym], Tylosteus ornatus Leidy 1872 [synonym]
Invalid names: Dracorex Bakker et al. 2006 [synonym], Stygimoloch Galton and Sues 1983 [synonym], Tylosteus Leidy 1872 [synonym]
G. †Stegoceras Lambe 1902
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†Stegoceras novomexicanum Jasinski and Sullivan 2011
Invalid names: Hanssuessia Sullivan 2003 [synonym], Ornatotholus Galton and Sues 1983 [synonym]
Fm. †Pachycephalosauridae Sternberg 1945
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Subfm. †Pachycephalosaurinae Sereno 1997
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G. †Sphaerotholus Williamson and Carr 2003
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†Sphaerotholus buchholtzae Williamson and Carr 2003
†Sphaerotholus edmontonensis Brown and Schlaikjer 1943
†Sphaerotholus goodwini Williamson and Carr 2003
†Sphaerotholus lyonsi Woodruff et al. 2023
†Sphaerotholus triregnum Woodruff et al. 2023
Unr. †Pachycephalosaurini Sullivan 2003
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Invalid names: Gravitholus Wall and Galton 1979 [nomen dubium], Texacephale Longrich et al. 2010 [nomen dubium], Tholocephalidae Sereno 1986 [synonym]
Invalid names: Homalocephalidae Dong 1978 [empty], Pachycephalosauroidea Zhao 1983 [empty], Tianchungosauroidea Zhao 1983 [empty]
Diagnosis
| Reference | Diagnosis | |
|---|---|---|
| P. Dodson 1997 | ...A common character being the presence of a frill, formed by the squamosal and parietal bones, that overhangs the back of the skull...Both groups lack the obturator process, a bony tab on the ischium that supports the pubis. |