PBDB Taxon

Basic info	Taxonomic history	Classification	Included Taxa
Morphology	Ecology and taphonomy	External Literature Search	Age range and collections

Protocupoides

Insecta - Coleoptera - Permocupedidae

Taxonomy

Protocupoides was named by Rohdendorf (1961). Its type is Protocupoides plavilstshikovi.

It was assigned to Cupidae by Rohdendorf (1961); to Permocupedidae by Pinto (1987) and Carpenter (1992); and to Permocupedinae by Kirejtshuk (2020).

Species

P. acer, P. carinatus, P. elongatus, P. esini, P. plavilstshikovi (type species), P. rohdendorfi, P. sharovi

Synonyms

Tomiocupes was named by Rohdendorf (1961).

It was synonymized subjectively with Protocupoides by Pinto (1987) and Kirejtshuk (2020).

It was assigned to Cupidae by Rohdendorf (1961).
Tricupes was named by Rohdendorf (1961).

It was synonymized subjectively with Protocupoides by Pinto (1987) and Kirejtshuk (2020).

It was assigned to Cupidae by Rohdendorf (1961).
Protocupoides rohdendorfi was named by Pinto (1987). Its type specimen is UFRGS MP-I-5266, an elytra, and it is an impression. Its type locality is Road Cut on Road BR 290 near Minas do Leao, which is in a Kungurian lacustrine - large siltstone in the Irati Formation of Brazil.

It was considered a nomen dubium by Kirejtshuk (2020).

Synonymy list

Year	Name and author
1961	Protocupoides Rohdendorf p. 408
1961	Tomiocupes Rohdendorf p. 408
1961	Tricupes Rohdendorf p. 410
1987	Protocupoides Pinto p. 8
1987	Protocupoides rohdendorfi Pinto p. 9 figs. 2; Pl I fig 2
1992	Protocupoides Carpenter
2020	Protocupoides Kirejtshuk p. 16

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Rank	Name	Author
kingdom	Animalia	()
—	Bilateria
—	Eubilateria	Ax 1987
—	Protostomia
—	Ecdysozoa
—	Panarthropoda
phylum	Arthropoda	Latreille 1829
superclass	Hexapoda	Latreille 1825
class	Insecta
—	Dicondylia
—	Paranotalia
subclass	Pterygota	()

Rank	Name	Author
—	Neopterygota	Crampton 1924
infraclass	Neoptera	Martynov 1923
—	Holometabola
—	Aparaglossata
—	Neuropteriformia
—	Coleopterida
order	Coleoptera
—	Mesocoleoptera
family	Permocupedidae
subfamily	Permocupedinae
genus	Protocupoides

If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.

G. †Protocupoides Rohdendorf 1961

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†Protocupoides acer Rohdendorf 1961

†Protocupoides carinatus Rohdendorf 1961

†Protocupoides elongatus Ponomarenko 2013

†Protocupoides esini Ponomarenko 2013

†Protocupoides plavilstshikovi Rohdendorf 1961

†Protocupoides sharovi Rohdendorf 1961

Invalid names: Protocupoides rohdendorfi Pinto 1987 [nomen dubium], Tomiocupes Rohdendorf 1961 [synonym], Tricupes Rohdendorf 1961 [synonym]

Diagnosis

No diagnoses are available

Measurements

No measurements are available

Composition:

chitinc

Environment:

terrestrialc

Locomotion:

actively mobilep

Created:

2017-04-17 09:56:18

Modified:

2017-04-17 09:56:18

Source: c = class, p = phylum

References: Kiessling 2004, Bush and Bambach 2015

Age range: base of the Kungurian to the top of the Wuchiapingian or 279.30000 to 254.17000 Ma

Collections (5 total)

Time interval	Ma	Country or state	Original ID and collection number
Kungurian	279.3 - 272.3	Brazil (Rio Grande do Sul)	P. rohdendorfi (116121)
Roadian	272.3 - 268.8	Russian Federation (Kemerovo)	P. plavilstshikovi, Tomiocupes sharovi (104704)
Wordian	266.9 - 264.28	Russian Federation (Udmurtia)	P. elongatus (103502)
Wuchiapingian	259.9 - 254.17	Russian Federation	Tomiocupes carinatus, Tricupes acer (124608)
Wuchiapingian	259.9 - 254.17	Russian Federation (Vologda)	P. esini (104754)