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Holocephali (chimaera)

Chondrichthyes

Synonyms
Synonymy list
YearName and author
1832Holocephali Bonaparte
1859Holocephali Owen p. 119
1873Holocephali Leidy p. 350
1898Holocephali Gadow p. 6
1902Holocephala Hay p. 322
1911Holocephala Jaekel p. 52
1934Holocephala Säve-Söderbergh p. 17
1946Holocephali Rapp p. 511
1958Bradyodonti Arambourg and Bertin
1963Holocephali Jordan p. 605
1964Bradyodonti Obruchev p. 362
1966Holocephali Cassier p. 98
1977Paraselachii Lund
1981Holocephali Duffin p. 470
1981Holocephali Thurmond and Jones p. 77
1981Bradyodonti Thurmond and Jones p. 78
1981Holocephali Zangerl p. 46
1982Bradyodonti Lund p. 939
1993Holocephali Cvancara and Hoganson p. 12
1997Euchondrocephali Lund and Grogan
2000Euchondrocephali Grogan and Lund p. 220
2000Paraselachii Grogan and Lund p. 220
2001Holocephali Purdy et al. p. 161
2002Holocephali Delsate et al. p. 25
2002Holocephali Sepkoski
2003Holocephali Kriwet and Gazdizicki p. 34
2004Paraselachiamorpha Goto and Okura p. 219
2006Paraselachimorpha Nelson p. 46
2008Holocephali Darras et al. p. 301
2011Holocephali Long p. 242
2015Holocephali Gouiric-Cavalli et al. p. 4
2016Holocephali Nelson et al. p. 48
2020Holocephali Popov et al. p. 5
2022Holocephali Ebersole et al. p. 373

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RankNameAuthor
kingdomAnimalia()
Bilateria
EubilateriaAx 1987
Life
Deuterostomia
phylumChordataHaeckel 1874
RankNameAuthor
subphylumVertebrata
superclassGnathostomata
classChondrichthyes
subclassHolocephali(Bonaparte 1832)
subclassHolocephali(Bonaparte 1832)

If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.

Subcl. Holocephali Bonaparte 1832 [chimaera]
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Fm. †Debeeriidae Grogan and Lund 2000
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G. †Debeerius Grogan and Lund 2000
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Debeerius ellefseni Grogan and Lund 2000
G. †Delphyodontos Lund 1980
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G. †Edaphodus Egerton 1836
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Invalid names: Passalodon Buckland 1838 [synonym]
Or. †Eugeneodontiformes Zangerl 1981
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Superfm. †Caseodontoidea Zangerl 1981
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G. †Campodus de Koninck 1844
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Campodus agassizianus Koninck 1844
Campodus corrugatus Newberry and Worthen 1870
Campodus scitulus Saint John and Worthen 1875
Campodus virginianus Saint John and Worthen 1875
Fm. †Caseodontidae Zangerl 1981
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G. †Caseodus Zangerl 1981
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Caseodus basalis Cope 1894
Caseodus eatoni Zangerl 1981
Caseodus varidentis Mutter and Neuman 2008
G. †Erikodus Nielsen 1952
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Erikodus groenlandicus Nielsen 1932
G. †Fadenia Nielsen 1932
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Fadenia crenulata Nielsen 1932
Fadenia gigas Eaton 1962
Fadenia uroclasmato Mutter and Neuman 2008
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Ornithoprion hertwigi Zangerl 1966
G. †Pirodus Lebedev 2001
G. †Romerodus Zangerl 1981
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Romerodus orodontus Zangerl 1981
G. †Chiastodus Trautschold 1879
Fm. †Eugeneodontidae Zangerl 1981
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G. †Bobbodus Zangerl 1981
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Bobbodus schaefferi Zangerl 1981
Bobbodus xerxesi Hampe et al. 2013
G. †Eugeneodus Zangerl 1981
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Eugeneodus richardsoni Zangerl 1981
G. †Gilliodus Zangerl 1981
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Gilliodus orvellei Zangerl 1981
Gilliodus peyeri Zangerl 1981
Invalid names: Orodus corrugatus Romer 1942 [nomen dubium]
G. †Tiaraju Richter 2007
Superfm. †Edestoidea Hay 1930
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Fm. †Edestidae Jaekel 1899
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G. †Edestus Leidy 1855
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Edestus crenulatus Hay 1909
Edestus giganteus Newberry 1888
Edestus heinrichi Newberry and Worthen 1870
Edestus karpinskii Missuna 1908
Edestus minisculus Hay 1909
Edestus minor Newberry 1866
Edestus minusculus Hay 1909
Edestus mirus Hay 1912
Edestus newtoni Woodward 1916
Edestus serratus Hay 1909
Edestus triserratus Newton 1904
Edestus vorax Leidy 1855
G. †Euctenius Traquair 1881
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Euctenius unilateralis Barkas 1874
G. †Lestrodus
G. †Prospiraxis Williams 1887
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Prospiraxis major Newberry 1885
Prospiraxis randallii Newberry 1885
G. †Protopirata Trautschold 1888
G. †Spiraxis Newberry 1885
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Spiraxis bivalvis Ward 1885
G. †Syntomodus Obruchev 1964
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Syntomodus abbreviatus Obruchev 1964
G. †Paredestus Mutter and Neuman 2008
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Paredestus bricircum Mutter and Neuman 2008
Fm. †Helicampodontidae Itano and Lebedev 2023
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G. †Helicampodus Branson 1935
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Helicampodus egloni Obruchev 1965
Helicampodus kokeni Branson 1935
G. †Hunanohelicoprion Liu 1994
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G. †Parahelicampodus Nielsen 1952
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Parahelicampodus sparcki Nielsen 1952
G. †Sinohelicoprion Liu and Chang 1963
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Sinohelicoprion changhsingensis Liu and Chang 1963
Fm. †Helicoprionidae Karpinsky 1911
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G. †Helicoprion Karpinsky 1899
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Helicoprion bessonowi Karpinsky 1899
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Invalid names: Helicoprion nevadensis Wheeler 1939 [synonym]
Helicoprion davisii Woodward 1886
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Invalid names: Helicoprion jingmenense Chen et al. 2007 [synonym], Helicoprion sierrensis Wheeler 1939 [synonym], Lissoprion ferrieri Hay 1907 [synonym]
Helicoprion ergassaminon Bendix-Almgreen 1966
Invalid names: Lissoprion Hay 1907 [synonym]
G. †Karpinskiprion Lebedev and Itano 2023
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Karpinskiprion ivanovi Karpinsky 1924
Invalid names: Agassizodontidae Zangerl 1981 [synonym]
Invalid names: Campyloprion Eastman 1902 [nomen dubium], Campyloprion annectans Eastman 1902 [nomen dubium], Edistidae Jaekel 1899 [empty]
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G. †Helodus Agassiz 1838
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Helodus angulatus Newberry and Worthen 1866
Helodus biformis Newberry and Worthen 1866
Helodus carbonarius Newberry and Worthen 1866
Helodus coniculus Newberry and Worthen 1866
Helodus consolidatus Newberry and Worthen 1866
Helodus crenulatus Newberry and Worthen 1866
Helodus denshumani Newberry and Worthen 1866
Helodus denticulatus Newberry and Worthen 1866
Helodus didymus Agassiz 1843
Helodus elytra Newberry and Worthen 1866
Helodus gibberulus Agassiz 1836
Helodus gibbus Leidy 1857
Helodus incisus Eastman 1903
Helodus laevis Newberry 1879
Helodus limax Newberry and Worthen 1866
Helodus mammillaris Agassiz 1843
Helodus planus Agassiz 1843
Helodus politus Newberry and Worthen 1866
Helodus rudis M'Coy 1848
Helodus rugosus Newberry and Worthen 1870
Helodus simplex Agassiz 1838
Helodus subpolitus Branson 1916
Helodus subteres Agassiz 1836
Helodus sulcatus Newberry and Worthen 1866
Helodus undulatus Newberry and Worthen 1866
Helodus wortheni Hay 1899
Superor. †Holocephalomorpha Lund 1986
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Or. Chimaeriformes Obruchev 1953 [chimaera]
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G. †Arctacanthus Nielsen 1932
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Arctacanthus exiguus Yamagishi 2004
Arctacanthus multisectus Branson 1933
Arctacanthus uncinatus Nielsen 1932
Arctacanthus wyomingensis Branson 1934
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Invalid names: Dolophonodus uncinatus Branson 1933 [replaced]
Invalid names: Ancistriodus Branson 1933 [synonym], Dolophonodus Branson 1933 [synonym], Hamatus Branson 1930 [synonym]
Fm. Callorhynchidae Garman 1901 [plownose chimaera]
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G. †Bathytheristes Duffin 1995
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Bathytheristes gracilis Duffin 1995
Subfm. Callorhynchinae Garman 1901
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G. †Ptyktoptychion Lees 1986
G. Callorhynchus Cuvier 1829
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Callorhynchus hectori Newton 1876
G. †Edaphadon Buckland 1838
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Edaphadon mirificus Leidy 1856
G. †Elasmodus Egerton
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Elasmodus avirostrus Averianov 1999
Elasmodus greenoughi Agassiz 1838
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Invalid names: Aptychus crassus Hebert 1855 [synonym], Elasmodus crassus Leriche 1906 [synonym], Elasmodus planus Leriche 1929 [synonym]
Elasmodus hunteri Egerton 1843
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Invalid names: Psaliodus compressus Agassiz 1843 [synonym]
Elasmodus kempi Ward 1977
Elasmodus khosatzkyi Averianov 1999
Elasmodus rossicus Averianov 1999
Elasmodus sinzovi Averianov and Glickman 1994
Elasmodus tsheganicus Averianov 2001
Elasmodus zharyk Averianov 1999
Invalid names: Psaliodus Agassiz 1843 [synonym]
G. †Ischyodus Egerton 1843
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Ischyodus avitus Meyer 1800
Ischyodus beaumonti Egerton 1843
Ischyodus bifurcatus Case 1978
Ischyodus bohemicus Bayer 1905
Ischyodus dufrenoyi Egerton 1843
Ischyodus egertoni Buckland 1835
Ischyodus emarginatus Egerton 1843
Ischyodus incisus Newton 1878
Ischyodus latus Newton 1878
Ischyodus mortoni Chapman and Pritchard 1907
Ischyodus panus Newton 1878
Ischyodus quenstedti Wagner 1857
Ischyodus rayhaasi Hoganson and Erickson 2005
Ischyodus thurmanni Pictet and Campiche 1878
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Invalid names: Ischyodus brevirostris Newton 1876 [synonym]
Ischyodus townsendi Buckland 1835
Ischyodus yanschini Averianov 1991
Ischyodus zinsmeisteri Applegate 1975
Invalid names: Edaphodontinae Stahl 1999 [empty]
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G. †Bryactinus Cope 1875
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Bryactinus amorphus Cope 1875
G. †Canadodus Popov et al. 2020
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Canadodus suntoki Popov et al. 2020
G. Chimaera Linnaeus 1758
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Subg. †Chimaera (Ischyodon) Agassiz 1843
Subg. †Chimaera (Psittacodon) Agassiz 1843
Chimaera furcata Fritsch 1878
Chimaera pliocenica Woodward 1891
Chimaera venablesae Cassier 1966
G. †Elasmodectes Woodward 1888
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Elasmodectes willetti Newton 1878
G. †Hedronchus Cope 1876
G. †Hydrolagus Gill 1862
G. †Isotaenia Cope 1875
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G. †Leptomylus Cope 1869
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Leptomylus cookii Cope 1870
Leptomylus densus Cope 1869
Leptomylus forfex Cope 1875
G. †Mylognathus Leidy 1856
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Mylognathus priscus Leidy 1856
G. †Similihariotta Zangerl 1979
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G. †Sphagepoea Cope 1869
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Sphagepoea aciculata Cope 1869
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G. †Anteliochimaera Tanaka 1909
Fm. Callorhinchidae Garman 1901 [plownose chimaera]
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G. †Brachymylus Smith Woodward 1894
G. Callorhinchus Lacépède 1798
G. †Ganodus Agassiz 1843
G. †Pachymylus Woodward 1892
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G. †Moskovirhynchus Popov and Shapovalov 2021
G. †Chalcodus Zittel 1887
Superfm. Chimaeroidea
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G. †Chimaerotheca Brown 1946
Fm. †Edaphodontidae Owen 1846
G. †Eomanodon Ward and Duffin 1989
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Eomanodon simmsi Ward and Duffin 1989
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G. †Amylodon Storms 1894
G. †Harriotta Goode and Bean 1895
G. †Rhinochimaera Garman 1901
G. †Crassidonta Branson 1916
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Crassidonta stuckenbergi Branson 1916
Crassidonta subcrenulata Teichert 1943
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G. †Echinochimaera Lund 1977
Subor. †Myriacanthoidei Patterson 1965
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G. †Chimaeropsis Zittel 1887
Fm. †Myriacanthidae Woodward 1889
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G. †Acanthorhina Fraas 1910
G. †Agkistracanthus Duffin and Furrer 1981
G. †Alethodontus Duffin 1983
G. †Halonodon Duffin 1984
G. †Metopacanthus Zittel 1887
G. †Myriacanthus Agassiz 1836
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Invalid names: Prognathodus Egerton 1872 [synonym]
G. †Oblidens Duffin and Milàn 2017
G. †Recurvacanthus Duffin 1981
Invalid names: Acanthorhinidae Patterson 1965 [synonym]
Invalid names: Chimeroidei Patterson 1965 [empty], Myracanthiodei [empty], Squalorajoidei Patterson 1965 [empty]
Or. †Chondrenchelyiformes Moy-Thomas 1939
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Fm. †Chondrenchelyidae Berg 1940
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G. †Chondrenchelys Traquair 1888
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Chondrenchelys problematica Traquair 1888
G. †Harpagofututor Lund 1982
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G. †Platyxystrodus Hay 1899
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Platyxystrodus bellulus St John and Worthen 1875
Platyxystrodus imitatus St John and Worthen 1875
Platyxystrodus inconditus Saint John and Worthen 1883
Platyxystrodus occidentalis St. John 1870
Platyxystrodus simplex St John and Worthen 1875
Platyxystrodus verus John and Worthen 1883
Invalid names: Xystrodus Morris and Roberts 1862 [replaced]
Or. †Cochliodontiformes Obruchev 1953
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Fm. †Cochliodontidae Owen 1867
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G. †Aspidodus Newberry and Worthen 1866
G. †Chitonodus Stjohn and Worthen 1883
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Chitonodus antiquus Saint John and Worthen 1883
Chitonodus liratus Saint John and Worthen 1883
Chitonodus springeri Saint John and Worthen 1883
G. †Cochliodus Agassiz 1838
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Cochliodus acutus Agassiz 1843
Cochliodus contortus Agassiz 1837
Cochliodus latus Leidy 1857
Cochliodus leidyi Saint John and Worthen 1883
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Invalid names: Deltodus stellatus Newberry and Worthen 1866 [synonym]
Cochliodus vanhornii Saint John and Worthen 1883
G. †Cranodus Trautschold 1879
G. †Cyrtonodus Davis 1884
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Cyrtonodus brevis Goto and Okura 2004
Cyrtonodus fukujiensis Goto and Okura 2004
Cyrtonodus hornei Woodward 1889
Cyrtonodus nishioi Goto and Okura 2004
G. †Deltodopsis St John and Worthen 1883
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Deltodopsis affinis Saint John and Worthen 1883
Deltodopsis angustus Newberry and Worthen 1870
Deltodopsis bialveatus Saint John and Worthen 1883
Deltodopsis convolutus Saint John and Worthen 1883
Deltodopsis exornatus Saint John and Worthen 1883
Deltodopsis inflexus Saint John and Worthen 1883
Deltodopsis sanctiludovici Saint John and Worthen 1883
G. †Deltodus Morris and Roberts 1862
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Deltodus alatus Newberry and Worthen 1870
Deltodus angularis Newberry and Worthen 1866
Deltodus cinctulus Saint John and Worthen 1883
Deltodus cinctus Newberry 1879
Deltodus cingulatus Newberry and Worthen 1866
Deltodus concha Trautschold 1874
Deltodus contortus Stjohn and Worthen 1883
Deltodus costatus Newberry and Worthen 1870
Deltodus fasciatus Newberry and Worthen 1870
Deltodus gibbus Woodward 1889
Deltodus grandis Newberry 1879
Deltodus inornatus Newberry 1897
Deltodus intermedius Saint John and Worthen 1883
Deltodus latior Saint John and Worthen 1883
Deltodus littoni Newberry and Worthen 1870
Deltodus mercurei Newberry 1876
Deltodus obliquus Saint John and Worthen 1883
Deltodus parvus Saint John and Worthen 1883
Deltodus planidens Cope 1894
Deltodus powellii Saint John and Worthen 1883
Deltodus propinquus Saint John and Worthen 1883
Deltodus rugosus Woodward 1889
Deltodus spatulatus Newberry and Worthen 1866
Deltodus sublaevis Agassiz 1843
Deltodus trilobus Saint John and Worthen 1883
Deltodus undulatus Newberry and Worthen 1866
G. †Dichelodus Giebel 1857
G. †Diplacodus Davis 1884
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Diplacodus bulboides Davis 1883
G. †Erismacanthus M'Coy 1848
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Erismacanthus barbatus Eastman 1903
Erismacanthus formosus Eastman 1902
Erismacanthus jonesii M'Coy 1848
Erismacanthus maccoyanus Stjohn and Worthen 1875
Erismacanthus major Davis 1883
G. †Helodopsis Waagen 1879
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Helodopsis abbreviata Waagen 1879
Helodopsis elongata Waagen 1879
G. †Icanodus Miller 1892
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Icanodus convexus Davis 1883
Icanodus limitaris St John and Worthen 1875
Invalid names: Tomodus Davis 1883 [replaced]
G. †Orthopleurodus St John and Worthen 1880
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Orthopleurodus convexus Saint John and Worthen 1883
Orthopleurodus novomexicanus Saint John and Worthen 1883
G. †Oxytomodus Trautschold 1880
G. †Paecilodus Agassiz 1843
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Paecilodus gibbosus Davis 1883
Paecilodus jonesii Mcoy 1855
G. †Periplectrodus St John and Worthen 1875
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Periplectrodus compressus Saint John and Worthen 1875
Periplectrodus expansus Saint John and Worthen 1875
Periplectrodus warreni Saint John and Worthen 1875
G. †Platyodus Newberry 1875
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Platyodus lineatus Newberry 1875
G. †Pleurodus Agassiz 1843
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Pleurodus rankinii Agassiz 1843
G. †Pleuroplax Woodward 1889
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Pleuroplax attheyi Barkas 1874
Pleuroplax rankinei Hancock and Atthey 1843
Pleuroplax woodi Davis 1883
G. †Poecilodus Agassiz 1843
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Poecilodus aliformis M'Coy 1848
Poecilodus carbonarius Saint John and Worthen 1883
Poecilodus cestriensis Saint John and Worthen 1883
Poecilodus convolutus Newberry and Worthen 1870
Poecilodus foveolatus M'Coy 1848
Poecilodus jonesii Agassiz 1843
Poecilodus oliquus Agassiz 1843
Poecilodus paradoxus Waagen 1879
Poecilodus parallelus Agassiz 1843
Poecilodus rugosus Newberry and Worthen 1866
Poecilodus sanctiludovici Saint John and Worthen 1883
Poecilodus springeri Saint John and Worthen 1883
Poecilodus tribulis Stjohn and Worthen 1883
Poecilodus varsoviensis Saint John and Worthen 1883
Poecilodus wortheni Saint John 1883
G. †Rhamphodus Davis 1883
G. †Sandalodus Newberry and Worthen 1866
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Sandalodus angustus Newberry and Worthen 1866
Sandalodus carbonarius Newberry and Worthen 1866
Sandalodus complanatus Newberry and Worthen 1866
Sandalodus crassus Newberry and Worthen 1866
Sandalodus elllipticus Newberry 1897
Sandalodus laevissimus Newberry and Worthen 1866
Sandalodus minor Newberry and Worthen 1866
Sandalodus morrisii Davis 1883
Sandalodus parvulus Newberry and Worthen 1866
Sandalodus rhomboideus Newberry and Worthen 1866
G. †Stenopterodus St John and Worthen 1883
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Stenopterodus elongatus Saint John and Worthen 1883
Stenopterodus planus Saint John and Worthen 1883
G. †Streblodus Agassiz 1843
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Streblodus angustus Eastman 1903
Streblodus colei Davis 1883
Streblodus egertoni Davis 1883
Streblodus oblongus Agassiz 1843
G. †Taeniodus St John and Worthen 1883
G. †Trigonodus Newberry and Worthen 1866
G. †Vaticinodus St John and Worthen 1883
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Vaticinodus carbonarius Saint John and Worthen 1883
Vaticinodus discrepans Saint John and Worthen 1883
Vaticinodus lepis Saint John and Worthen 1883
Vaticinodus similis Saint John and Worthen 1883
Vaticinodus simplex Saint John and Worthen 1883
Vaticinodus vetustus Saint John and Worthen 1883
G. †Venustodus St. John and Worthen 1875
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Venustodus argutus St John and Worthen 1875
Venustodus leidyi St. John and Worthen 1875
Venustodus robustus Saint John and Worthen 1875
Venustodus tenuicristatus Saint John and Worthen 1875
Venustodus variabilis Saint John and Worthen 1875
Venustodus venustus Leidy 1857
G. †Xystrodus Plieninger 1860
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Xystrodus egertoni Davis 1883
Fm. †Psephodontidae Zangerl 1981
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G. †Kanodus Lebedev 2014
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Kanodus robustus Lebedev 2014
G. †Psephodus Morris and Roberts 1862
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Psephodus crenulatus Newberry and Worthen 1866
Psephodus dubius Woodward 1889
Psephodus latus Saint John and Worthen 1883
Psephodus levissimus Agassiz 1837
Psephodus lunulatus Saint John and Worthen 1883
Psephodus magnus Agassiz 1843
Psephodus obliquus Saint John and Worthen 1883
Psephodus placenta Newberry and Worthen 1866
Psephodus politus Newberry 1897
Psephodus regularis Saint John and Worthen 1883
Psephodus reticulatus Saint John and Worthen 1875
Psephodus salopiensis Woodward 1889
Psephodus symmetricus Saint John and Worthen 1883
Invalid names: Psephodus depressus Waagen 1880 [nomen vanum], Psephodus indicus Waagen 1880 [nomen vanum]
G. †Solenodus Trautschold 1874
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Solenodus crenulatus Trautschold 1874
Or. †Copodontiformes Obruchev 1953
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Fm. †Copodontidae Davis 1883
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G. †Copodus Morris and Roberts 1862
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Copodus auriculatus Davis 1883
Copodus cornutus Davis 1843
Copodus furcatus Davis 1862
Copodus lingua Davis 1883
Copodus minimus Davis 1883
Copodus oblongus Davis 1883
Copodus pusillus Saint John and Worthen 1883
Copodus spatulatus Davis 1862
Copodus spatulatus Davis 1862
Copodus vanhornii Saint John and Worthen 1883
G. †Melanodus Darras et al. 2008
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Melanodus loonesi Darras et al. 2008
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Fm. †Deltoptychiidae Zangerl 1981
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G. †Deltoptychius Morris and Roberts 1862
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Deltoptychius acutus Agassiz 1843
Deltoptychius expansus Saint John and Worthen 1883
Deltoptychius nitidus Leidy 1857
Deltoptychius primus Saint John and Worthen 1883
Deltoptychius varsoviensis Saint John and Worthen 1883
Deltoptychius wachsmuthi Saint John and Worthen 1883
Fm. †Menaspidae Woodward 1891
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G. †Menaspis Ewald 1848
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Menaspis armata Ewald 1848
Fm. †Traquairiidae Lund and Grogan 1997
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G. †Traquairius Lund and Grogan 1997
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Traquairius agkistrocephalus Lund and Grogan 1997
Traquairius nudus Lund and Grogan 1997
Traquairius spinosus Lund and Grogan 1997
Or. †Psammodontiformes Obruchev 1953
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+
G. †Apodometes Gistel 1848
G. †Archaeobatis Newberry 1878
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Archaeobatis gigas Newberry 1878
G. †Astrabodus Davis 1884
G. †Dimyleus Davis 1883
G. †Homalodus Davis 1883
G. †Mazodus Newberry 1899
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Mazodus kepleri Newberry 1889
G. †Mylacodus Davis 1883
G. †Mylax Morris and Roberts 1883
G. †Pinacodus Davis 1883
G. †Pleurogomphus Agassiz 1862
G. †Psammodus Agassiz 1834
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Psammodus angularis Newberry and Worthen 1866
Psammodus caelatus Saint John and Worthen 1883
Psammodus cornutus Agassiz 1843
Psammodus crassidens Saint John and Worthen 1883
Psammodus expansus Davis 1884
Psammodus glyptus Saint John and Worthen 1883
Psammodus grandis Saint John and Worthen 1883
Psammodus longidens Agassiz 1837
Psammodus lovianus Saint John and Worthen 1883
Psammodus plenus Saint John and Worthen 1883
Psammodus porosus Agassiz 1837
Psammodus reticulatus Newberry and Worthen 1866
Psammodus rugosus Agassiz 1836
Psammodus salopiensis Woodward 1889
Psammodus springeri Saint John and Worthen 1883
Psammodus tumidus Saint John and Worthen 1883
Psammodus turgidus Saint John and Worthen 1883
Psammodus turgidus Agassiz 1838
G. †Rhizodus
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Rhizodus gracilis M'Coy 1848
Rhizodus hibberti Agassiz 1843
Rhizodus reticulatus Newberry and Worthen 1870
Invalid names: Characodus Morris and Roberts 1862 [synonym]
G. †Rhymodus Davis 1883
Or. †Squalorajiformes Patterson 1965
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Fm. †Squalorajidae Woodward 1886
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G. †Squaloraja Riley 1833
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Squaloraja polyspondyla Agassiz 1836
Squaloraja tenuispina Woodward 1886
Invalid names: Spinacorhinus Agassiz 1836 [synonym]
G. †Macrodontacanthus Romer 1942
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G. †Menaspacanthus Patterson 1965
Or. †Orodontiformes Zangerl 1981
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Fm. †Leiodontidae Ginter et al. 2010
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G. †Cristatodens Ginter and Sun 2007
G. †Leiodus Saint John and Worthen 1875
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Leiodus calcaratus Saint John and Worthen 1875
Leiodus grossipunctatus Saint John and Worthen 1875
Fm. †Orodontidae de Koninck 1878
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G. †Adiapneustes Gistel 1848
G. †Eoorodus Branson 1914
G. †Hercynolepis Gross 1973
G. †Lophodus
G. †Octinapsis Trautschold 1874
G. †Orodus Agassiz 1836
+
Orodus alleni Saint John and Worthen 1875
Orodus carinatus Saint John and Worthen 1875
Orodus catenatus Davis 1883
Orodus cinctus Agassiz 1836
Orodus colletti Newberry 1879
Orodus daedalus Saint John and Worthen 1875
Orodus decussatus Saint John and Worthen 1875
Orodus elegantulus Newberry and Worthen 1866
Orodus elongatus Davis 1883
Orodus fastigiatus Saint John and Worthen 1875
Orodus gibbosus Newberry and Worthen 1866
Orodus gibbus Davis 1883
Orodus greggi Zangerl 1981
Orodus intermedius Eastman 1903
Orodus ipeunaensis Chahud et al. 2010
Orodus major Saint John and Worthen 1875
Orodus mammillaris Newberry and Worthen 1866
Orodus micropterygius Zangerl 1981
Orodus minutus Newberry and Worthen 1866
Orodus moniliformis Davis 1883
Orodus neglectus Saint John and Worthen 1875
Orodus ornatus Newberry and Worthen 1866
Orodus parallelus Saint John and Worthen 1875
Orodus parvulus Saint John and Worthen 1875
Orodus plicatus Newberry and Worthen 1866
Orodus ramosus Agassiz 1836
Orodus tenuis Woodward 1889
Orodus tuberculatus Newberry and Worthen 1866
Orodus turgidus Saint John and Worthen 1875
Orodus variabilis Newberry 1875
Orodus varicostatus Saint John and Worthen 1875
Or. †Petalodontiformes Zangerl 1981
+
Fm. †Belantseidae Lund 1989
+
G. †Belantsea Lund 1989
+
Belantsea montana Lund 1989
Belantsea occidentalis St. John and Worthen 1875
G. †Ctenoptychius Agassiz 1838
+
Ctenoptychius acuminatus Saint John and Worthen 1875
Ctenoptychius apicalis Agassiz 1838
Ctenoptychius cristatus Dawson 1868
Ctenoptychius lobatus Etheridge 1875
Ctenoptychius pertenius Saint John and Worthen 1875
Ctenoptychius stevensoni St. John and Worthen 1875
G. †Netsepoye Lund 1989
+
Netsepoye hawesi Lund 1989
Fm. †Janassidae Jaekel 1899
+
G. †Cholodus John and Worthen 1875
G. †Ctenopetalus Agassiz 1837
+
Ctenopetalus bellulus Saint John and Worthen 1875
Ctenopetalus limatulus Saint John and Worthen 1875
Ctenopetalus medius Saint John and Worthen 1875
Ctenopetalus vinosus Saint John and Worthen 1875
G. †Cymatodus Newberry and Worthen 1870
+
Cymatodus oblongus Newberry and Worthen 1870
+
Dactylodus excavatus Saint John and Worthen 1866
Dactylodus princeps Newberry and Worthen 1866
G. †Fissodus St. John and Worthen 1875
+
Fissodus bifidus Saint John and Worthen 1875
Fissodus dentatus Eastman 1903
Fissodus inaequalis Stjohn and Worthen 1875
G. †Gamphacanthus Miller 1892
+
Gamphacanthus politus Newberry 1889
Gamphacanthus uddeni Lindahl 1897
G. †Gomphacanthus Davis 1884
G. †Janassa
+
Janassa angularis Branson 1916
Janassa bituminosa Schlotheim 1820
+
Invalid names: Acrodus larva Agassiz 1839 [synonym], Byzenos latipinnatus Münster 1843 [synonym], Dictea striata Münster 1840 [synonym], Janassa angulata Münster 1839 [synonym], Janassa dictea Münster 1842 [synonym], Janassa humboldii Münster 1839 [synonym]
Janassa brevis Newberry 1856
Janassa clarki Lund 1989
Janassa clavatus M'Coy 1848
Janassa gurleiana Cope 1877
Janassa imbricatus M'Coy 1848
Janassa kochi Nielsen 1932
Janassa korni Weigelt 1930
Janassa linguaeformis Atthey 1868
Janassa maxima Eastman 1903
Janassa ordiana Cope 1881
Janassa strigilina Cope 1881
Janassa unguicula Eastman 1903
Invalid names: Byzenos Münster 1843 [synonym], Climaxodus McCoy 1848 [synonym], Dictea Münster 1840 [synonym]
G. †Petalodopsis Barkas 1874
+
Petalodopsis mirabilis Barkas 1874
G. †Phoderacanthus Davis 1883
G. †Platacanthus M'Coy 1848
G. †Pnigeacanthus St John and Worthen 1883
G. †Pristicladodus M'Coy 1848
+
Pristicladodus goughi M'Coy 1848
G. †Stichacanthus Koninck 1878
+
Stichacanthus coemansi Koninck 1878
G. †Thoracodus Cope 1883
+
Thoracodus emydinus Cope 1883
Fm. †Petalodontidae Newberry and Worthen 1866
+
G. †Antliodus Newberry and Worthen 1866
+
Antliodus arcuatus Newberry 1889
Antliodus cucullus Newberry and Worthen 1866
Antliodus gracilis Saint John and Worthen 1875
Antliodus mucronatus Newberry and Worthen 1866
Antliodus parvulus Newberry and Worthen 1866
Antliodus perovalis Saint John and Worthen 1875
Antliodus politus Newberry and Worthen 1866
Antliodus robustus Newberry and Worthen 1866
Antliodus sarcululus Newberry and Worthen 1870
Antliodus similis Newberry and Worthen 1866
Antliodus simplex Newberry and Worthen 1866
Antliodus sulcatus Newberry and Worthen 1866
G. †Chomatodus Agassiz 1838
+
Chomatodus affinis Newberry and Worthen 1866
Chomatodus angustus Newberry 1879
Chomatodus chesterensis Saint John and Worthen 1875
Chomatodus cinctus Agassiz 1838
Chomatodus comptus Saint John and Worthen 1875
Chomatodus costatus Newberry and Worthen 1866
Chomatodus cultellus Newberry and Worthen 1866
Chomatodus elegans Newberry and Worthen 1866
Chomatodus inconstans Saint John and Worthen 1875
Chomatodus incrassatus Saint John and Worthen 1875
Chomatodus insignis Leidy 1857
Chomatodus linearis Agassiz 1836
Chomatodus loriformis Newberry and Worthen 1866
Chomatodus molaris Newberry and Worthen 1866
Chomatodus oliquus M'Coy 1848
Chomatodus parallelus Saint John and Worthen 1875
Chomatodus pusillus Newberry and Worthen 1866
Chomatodus selliformis Newberry 1879
Chomatodus truncatus Agassiz 1843
Chomatodus varsouviensis Saint John and Worthen 1875
G. †Glossodus M'Coy 1848
+
Glossodus heckelii Costa 1864
Glossodus lingusbovis Mcoy 1848
Glossodus marginatus M'Coy 1848
G. †Glyphanodus Davis 1883
G. †Harpacodus St. John and Worthen 1875
+
Harpacodus clavatus Davis 1883
Harpacodus compactus St. John and Worthen 1875
Harpacodus dentatus Davis 1883
Harpacodus occidentalis St. John and Worthen 1875
G. †Itapyrodus Silva Santos 1990
+
Itapyrodus punctatus Silva Santos 1990
G. †Lisgodus Saint John and Worthen 1875
+
Lisgodus affinis Newberry 1879
Lisgodus curtus Saint John and Worthen 1875
Lisgodus selluliformis Saint John and Worthen 1875
Lisgodus serratus Saint John and Worthen 1875
G. †Neopetalodus Murata and Uyeno 1979
G. †Peltodus Newberry and Worthen 1870
+
Peltodus plicomphalus Saint John and Worthen 1875
Peltodus pulvinulus Newberry and Worthen 1870
Peltodus quadratus Saint John and Worthen 1875
Peltodus transversus Saint John and Worthen 1875
Peltodus unguiformis St John and Worthen 1870
G. †Petalodus
+
Petalodus acuminatus Agassiz 1838
+
Invalid names: Petalodus destructor Newberry and Worthen 1870 [synonym], Sicarius extinctus Leidy 1856 [synonym]
Petalodus arcuatus St. John 1870
Petalodus curtus Newberry and Worthen 1870
Petalodus davisii Woodward 1889
Petalodus flabellula Woodward 1889
Petalodus grandis Davis 1883
Petalodus hastingsiae Owen 1840
Petalodus hybridus Saint John and Worthen 1875
Petalodus jewetti Miller 1957
Petalodus knappi Newberry 1879
Petalodus linguifer Newberry and Worthen 1866
Petalodus ohioensis Safford 1853
Petalodus proximus Saint John and Worthen 1875
Petalodus sagittatus Agassiz 1843
Invalid names: Sicarius Leidy 1856 [synonym]
G. †Polyrhizodus M'Coy 1848
+
Polyrhizodus carbonarius Saint John and Worthen 1875
Polyrhizodus colei Davis 1883
Polyrhizodus concavus Trautschold 1874
Polyrhizodus dentatus Newberry and Worthen 1866
Polyrhizodus digitatus Leidy 1857
+
Invalid names: Polyrhizodus amplus St. John and Worthen 1875 [synonym], Polyrhizodus excavatus Saint John and Worthen 1866 [synonym], Polyrhizodus littoni Newberry and Worthen 1870 [synonym], Polyrhizodus lobatus Newberry and Worthen 1866 [synonym], Polyrhizodus princeps Newberry and Worthen 1866 [synonym], Polyrhizodus williamsi St. John and Worthen 1875 [synonym]
Polyrhizodus elongatus Davis 1883
Polyrhizodus inflexus Newberry and Worthen 1866
Polyrhizodus latus Newberry 1897
Polyrhizodus magnus M'Coy 1848
Polyrhizodus minimus Saint John and Worthen 1875
Polyrhizodus modestus Newberry 1875
Polyrhizodus nanus Saint John and Worthen 1875
Polyrhizodus piasaensis Saint John and Worthen 1875
Polyrhizodus ponticulus Newberry and Worthen 1866
Polyrhizodus porosus Newberry and Worthen 1866
Polyrhizodus pusillus M'Coy 1848
Polyrhizodus rectus Newberry 1897
Polyrhizodus truncatus Newberry and Worthen 1870
G. †Strigilina Cope 1877
+
G. †Tanaodus St. John and Worthen 1875
+
Tanaodus angularis Newberry and Worthen 1866
Tanaodus bellcinctus Saint John and Worthen 1875
Tanaodus depressus Saint John and Worthen 1875
Tanaodus gracillimus Newberry and Worthen 1866
Tanaodus grossiplicatus Saint John and Worthen 1875
Tanaodus multiplicatus Newberry and Worthen 1866
Tanaodus obscurus Leidy 1857
Tanaodus polymorphus Saint John and Worthen 1875
Tanaodus praenuntius Saint John and Worthen 1875
Tanaodus pumilis Saint John and Worthen 1875
Tanaodus sculptus Saint John and Worthen 1875
Tanaodus sublunatus Saint John and Worthen 1875
Tanaodus wisei Traquair 1901
Fm. †Pristodontidae Woodward 1889
+
G. †Calopodus John and Worthen 1875
+
Calopodus apicalis Saint John and Worthen 1875
G. †Davodus Hansen 1985
G. †Megactenopetalus David 1944
+
+
Invalid names: Petalodus shingkuoi Young 1950 [synonym]
G. †Peripristis St. John 1870
+
Peripristis semicircularis Newberry and Worthen 1866
G. †Petalorhynchus Newberry and Worthen 1866
+
Petalorhynchus distortus Saint John and Worthen 1875
Petalorhynchus indicus Waagen 1879
Petalorhynchus pseudosagittatus Saint John and Worthen 1875
Petalorhynchus spatulatus Saint John and Worthen 1875
Petalorhynchus striatus Newberry and Worthen 1866
G. †Pristodus Davis 1883
+
Pristodus benniei Etheridge 1875
Pristodus concinnus Davis 1883
Pristodus falcatus Davis 1883
G. †Siksika Lund 1989
+
Siksika ottae Lund 1989
Invalid names: Peripristidae Eastman 1902 [synonym]
Invalid names: Bradyodonti Arambourg and Bertin 1958 [synonym], Chimaeri Jaekel 1911 [empty], Debeeriformes [empty], Euchondrocephali Lund and Grogan 1997 [synonym], Iniopterygiformes Zangerl and Case 1973 [empty], Paraselachiamorpha Lund 1977 [empty], Paraselachimorpha Lund 1977 [invalid subgroup], Trachyacanthi Jaekel 1911 [empty]
Diagnosis
ReferenceDiagnosis
R. Owen 1859Jaws bony, traversed and encased by dental plates; endo-skeleton cartilaginous; exo-skeleton as placoid granules; most of the fins with a strong spine for the first ray; ventrals abdominal; gills laminated, attached by their margins; a single external gill aperture.