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Protoceratopsidae
Taxonomy
Protoceratopsidae was named by Granger and Gregory (1923). Its type is Protoceratops. It was considered paraphyletic by Dodson (1997) and monophyletic by Sereno (2000).
It was synonymized subjectively with Leptoceratopsinae by Huene (1934) and Huene (1934); it was corrected as Protoceratopidae by Olshevsky and Ford (1994), Tereshchenko and Alifanov (2003) and Tereshchenko (2007).
It was assigned to Orthopoda by Bohlin (1953); to Ponderopoda by Huene (1948) and Huene (1959); to Ceratopsoidea by Zhao (1983), Norman (1984) and Cooper (1985); to Neoceratopia by Olshevsky and Ford (1994); to Ceratopsia by Gregory and Mook (1925), Lull (1933), Roxo (1937), Brown and Schlaikjer (1942), Kuhn (1946), Huene (1950), Romer (1956), Parsch (1963), Tatarinov (1964), Kuhn (1964), Kuhn (1966), Romer (1966), Charig (1967), Swinton (1970), Thulborn (1971), Maryanska and Osmólska (1975), Rozhdestvensky (1977), Ostrom (1980), McIntosh (1981), McGinnis (1982), Sues and Galton (1982), Russell (1984), Maryanska and Osmólska (1985), Ostrom and Wellnhofer (1986), Bakker (1986), Carroll (1988) and Nessov (1995); to Neoceratopsia by Dodson and Currie (1990), Dodson (1990), Dodson (1997), Ryan and Currie (1998), Chinnery and Weishampel (1998), Lambert et al. (2001), Tereshchenko and Alifanov (2003), Chinnery (2004) and Ott (2007); to Protoceratopoidea by Tereshchenko (2007); and to Coronosauria by Sereno (1997), Sereno (1998), Sereno (1999), Sereno (2000), Makovicky (2001), You and Dodson (2003), Butler et al. (2008) and Morschhauser et al. (2019).
It was synonymized subjectively with Leptoceratopsinae by Huene (1934) and Huene (1934); it was corrected as Protoceratopidae by Olshevsky and Ford (1994), Tereshchenko and Alifanov (2003) and Tereshchenko (2007).
It was assigned to Orthopoda by Bohlin (1953); to Ponderopoda by Huene (1948) and Huene (1959); to Ceratopsoidea by Zhao (1983), Norman (1984) and Cooper (1985); to Neoceratopia by Olshevsky and Ford (1994); to Ceratopsia by Gregory and Mook (1925), Lull (1933), Roxo (1937), Brown and Schlaikjer (1942), Kuhn (1946), Huene (1950), Romer (1956), Parsch (1963), Tatarinov (1964), Kuhn (1964), Kuhn (1966), Romer (1966), Charig (1967), Swinton (1970), Thulborn (1971), Maryanska and Osmólska (1975), Rozhdestvensky (1977), Ostrom (1980), McIntosh (1981), McGinnis (1982), Sues and Galton (1982), Russell (1984), Maryanska and Osmólska (1985), Ostrom and Wellnhofer (1986), Bakker (1986), Carroll (1988) and Nessov (1995); to Neoceratopsia by Dodson and Currie (1990), Dodson (1990), Dodson (1997), Ryan and Currie (1998), Chinnery and Weishampel (1998), Lambert et al. (2001), Tereshchenko and Alifanov (2003), Chinnery (2004) and Ott (2007); to Protoceratopoidea by Tereshchenko (2007); and to Coronosauria by Sereno (1997), Sereno (1998), Sereno (1999), Sereno (2000), Makovicky (2001), You and Dodson (2003), Butler et al. (2008) and Morschhauser et al. (2019).
Synonymy list
Year | Name and author |
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1923 | Protoceratopsidae Granger and Gregory p. 4 |
1925 | Protoceratopsidae Gregory and Mook p. 4 |
1933 | Protoceratopsidae Lull p. 74 |
1937 | Protoceratopsidae Roxo p. 65 |
1942 | Protoceratopsidae Brown and Schlaikjer p. 2 |
1946 | Protoceratopsidae Kuhn p. 66 |
1948 | Protoceratopsidae Huene p. 95 |
1950 | Protoceratopsidae Huene p. 351 |
1953 | Protoceratopsidae Bohlin p. 13 |
1956 | Protoceratopsidae Romer p. 637 |
1959 | Protoceratopsidae Huene p. 122 |
1963 | Protoceratopsidae Parsch p. 4 |
1964 | Protoceratopsidae Kuhn p. 52 |
1964 | Protoceratopsidae Tatarinov p. 578 |
1966 | Protoceratopsidae Kuhn p. 106 |
1966 | Protoceratopsidae Romer p. 370 |
1967 | Protoceratopsidae Charig p. 717 |
1970 | Protoceratopsidae Swinton p. 254 |
1971 | Protoceratopsidae Thulborn pp. 77-78 fig. 4 |
1975 | Protoceratopsidae Maryanska and Osmólska p. 137 |
1977 | Protoceratopsidae Rozhdestvensky p. 114 |
1980 | Protoceratopsidae Ostrom p. 23 |
1981 | Protoceratopsidae McIntosh p. 40 |
1982 | Protoceratopsidae McGinnis p. 96 |
1982 | Protoceratopsidae Sues and Galton p. 189 |
1983 | Protoceratopsidae Zhao p. 300 |
1984 | Protoceratopsidae Norman p. 160 fig. 2 |
1984 | Protoceratopsidae Russell p. 29 |
1985 | Protoceratopsidae Cooper p. 292 |
1985 | Protoceratopsidae Maryanska and Osmólska pp. 143-144 fig. 1 |
1986 | Protoceratopsidae Bakker p. 461 |
1986 | Protoceratopsidae Ostrom and Wellnhofer p. 116 |
1988 | Protoceratopsidae Carroll |
1990 | Protoceratopsidae Dodson p. 578 |
1990 | Protoceratopsidae Dodson and Currie p. 611 |
1994 | Protoceratopidae Olshevsky and Ford |
1995 | Protoceratopsidae Nessov p. 52 |
1997 | Protoceratopsidae Dodson p. 473–474 |
1997 | Protoceratopsidae Sereno p. 443 |
1998 | Protoceratopsidae Chinnery and Weishampel p. 578 fig. 10 |
1998 | Protoceratopsidae Ryan and Currie p. 821 |
1998 | Protoceratopsidae Sereno p. 62 |
1999 | Protoceratopsidae Sereno p. 2138 fig. 2 |
2000 | Protoceratopsidae Sereno p. 482 |
2001 | Protoceratopsidae Lambert et al. pp. 7, 18 |
2001 | Protoceratopsidae Makovicky p. 254 fig. 18.3 |
2003 | Protoceratopidae Tereshchenko and Alifanov p. 293 |
2003 | Protoceratopsidae You and Dodson |
2004 | Protoceratopsidae Chinnery pp. 584-585 fig. 9 |
2007 | Protoceratopsidae Ott p. 214 |
2007 | Protoceratopidae Tereshchenko p. 175 |
2008 | Protoceratopsidae Butler et al. p. 8 |
2019 | Protoceratopsidae Morschhauser et al. p. 129 fig. 10 |
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If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.
Fm. †Protoceratopsidae Granger and Gregory 1923
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G. †Bagaceratops Maryanska and Osmólska 1975
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†Bagaceratops rozhdestvenskyi Maryanska and Osmólska 1975
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Invalid names: Gobiceratops minutus Alifanov 2008 [synonym], Lamaceratops tereschenkoi Alifanov 2003 [synonym], Magnirostris dodsoni You and Dong 2003 [synonym], Platyceratops tatarinovi Alifanov 2003 [synonym]
Invalid names: Gobiceratops Alifanov 2008 [synonym], Lamaceratops Alifanov 2003 [synonym], Magnirostris You and Dong 2003 [synonym], Platyceratops Alifanov 2003 [synonym]
G. †Protoceratops Granger and Gregory 1923
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†Protoceratops andrewsi Granger and Gregory 1923
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Invalid names: Bainoceratops efremovi Tereshchenko and Alifanov 2003 [synonym]
†Protoceratops hellenikorhinus Lambert et al. 2001
Invalid names: Bainoceratops Tereshchenko and Alifanov 2003 [synonym]
Invalid names: Asiaceratopsinae Nessov et al. 1989 [empty]
Diagnosis
Reference | Diagnosis | |
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F. v. Huene 1948 | Few teeth in posteiror part of premaxilla; 1-2 rows of single rooted teeth in jaws; large preorbit; two pairs of large temporal fossae; frontals and parietals in exterior roof of skull; short high mandibles; relatively long pubis. |