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Protoceratopsidae

Reptilia - Protoceratopsidae

Taxonomy
Protoceratopsidae was named by Granger and Gregory (1923). Its type is Protoceratops. It was considered paraphyletic by Dodson (1997) and monophyletic by Sereno (2000).

It was synonymized subjectively with Leptoceratopsinae by Huene (1934) and Huene (1934); it was corrected as Protoceratopidae by Olshevsky and Ford (1994), Tereshchenko and Alifanov (2003) and Tereshchenko (2007).

It was assigned to Orthopoda by Bohlin (1953); to Ponderopoda by Huene (1948) and Huene (1959); to Ceratopsoidea by Zhao (1983), Norman (1984) and Cooper (1985); to Neoceratopia by Olshevsky and Ford (1994); to Ceratopsia by Gregory and Mook (1925), Lull (1933), Roxo (1937), Brown and Schlaikjer (1942), Kuhn (1946), Huene (1950), Romer (1956), Parsch (1963), Tatarinov (1964), Kuhn (1964), Kuhn (1966), Romer (1966), Charig (1967), Swinton (1970), Thulborn (1971), Maryanska and Osmólska (1975), Rozhdestvensky (1977), Ostrom (1980), McIntosh (1981), McGinnis (1982), Sues and Galton (1982), Russell (1984), Maryanska and Osmólska (1985), Ostrom and Wellnhofer (1986), Bakker (1986), Carroll (1988) and Nessov (1995); to Neoceratopsia by Dodson and Currie (1990), Dodson (1990), Dodson (1997), Ryan and Currie (1998), Chinnery and Weishampel (1998), Lambert et al. (2001), Tereshchenko and Alifanov (2003), Chinnery (2004) and Ott (2007); to Protoceratopoidea by Tereshchenko (2007); and to Coronosauria by Sereno (1997), Sereno (1998), Sereno (1999), Sereno (2000), Makovicky (2001), You and Dodson (2003), Butler et al. (2008) and Morschhauser et al. (2019).

Synonymy list
YearName and author
1923Protoceratopsidae Granger and Gregory p. 4
1925Protoceratopsidae Gregory and Mook p. 4
1933Protoceratopsidae Lull p. 74
1937Protoceratopsidae Roxo p. 65
1942Protoceratopsidae Brown and Schlaikjer p. 2
1946Protoceratopsidae Kuhn p. 66
1948Protoceratopsidae Huene p. 95
1950Protoceratopsidae Huene p. 351
1953Protoceratopsidae Bohlin p. 13
1956Protoceratopsidae Romer p. 637
1959Protoceratopsidae Huene p. 122
1963Protoceratopsidae Parsch p. 4
1964Protoceratopsidae Kuhn p. 52
1964Protoceratopsidae Tatarinov p. 578
1966Protoceratopsidae Kuhn p. 106
1966Protoceratopsidae Romer p. 370
1967Protoceratopsidae Charig p. 717
1970Protoceratopsidae Swinton p. 254
1971Protoceratopsidae Thulborn pp. 77-78 fig. 4
1975Protoceratopsidae Maryanska and Osmólska p. 137
1977Protoceratopsidae Rozhdestvensky p. 114
1980Protoceratopsidae Ostrom p. 23
1981Protoceratopsidae McIntosh p. 40
1982Protoceratopsidae McGinnis p. 96
1982Protoceratopsidae Sues and Galton p. 189
1983Protoceratopsidae Zhao p. 300
1984Protoceratopsidae Norman p. 160 fig. 2
1984Protoceratopsidae Russell p. 29
1985Protoceratopsidae Cooper p. 292
1985Protoceratopsidae Maryanska and Osmólska pp. 143-144 fig. 1
1986Protoceratopsidae Bakker p. 461
1986Protoceratopsidae Ostrom and Wellnhofer p. 116
1988Protoceratopsidae Carroll
1990Protoceratopsidae Dodson p. 578
1990Protoceratopsidae Dodson and Currie p. 611
1994Protoceratopidae Olshevsky and Ford
1995Protoceratopsidae Nessov p. 52
1997Protoceratopsidae Dodson p. 473–474
1997Protoceratopsidae Sereno p. 443
1998Protoceratopsidae Chinnery and Weishampel p. 578 fig. 10
1998Protoceratopsidae Ryan and Currie p. 821
1998Protoceratopsidae Sereno p. 62
1999Protoceratopsidae Sereno p. 2138 fig. 2
2000Protoceratopsidae Sereno p. 482
2001Protoceratopsidae Lambert et al. pp. 7, 18
2001Protoceratopsidae Makovicky p. 254 fig. 18.3
2003Protoceratopidae Tereshchenko and Alifanov p. 293
2003Protoceratopsidae You and Dodson
2004Protoceratopsidae Chinnery pp. 584-585 fig. 9
2007Protoceratopsidae Ott p. 214
2007Protoceratopidae Tereshchenko p. 175
2008Protoceratopsidae Butler et al. p. 8
2019Protoceratopsidae Morschhauser et al. p. 129 fig. 10

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RankNameAuthor
kingdomAnimalia()
Bilateria
EubilateriaAx 1987
Deuterostomia
phylumChordataHaeckel 1874
subphylumVertebrata
superclassGnathostomata
Osteichthyes()
subclassSarcopterygii()
subclassDipnotetrapodomorpha(Nelson 2006)
subclassTetrapodomorpha()
Tetrapoda
Reptiliomorpha
Anthracosauria
subclassAmphibiosauriaKuhn 1967
Cotylosauria()
Amniota
Sauropsida
classReptilia
subclassEureptilia()
Romeriida
RankNameAuthor
Diapsida()
Archosauromorpha(Huene 1946)
Crocopoda
ArchosauriformesGauthier 1986
Eucrocopoda
Archosauria()
informalAvemetatarsalia
Ornithodira
Dinosauromorpha
Dinosauriformes
Dinosauria()
Ornithischia()
Genasauria
Cerapoda
Marginocephalia
Ceratopsia()
infraorderNeoceratopsia
Coronosauria
familyProtoceratopsidae
familyProtoceratopsidae

If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.

Fm. †Protoceratopsidae Granger and Gregory 1923
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G. †Bagaceratops Maryanska and Osmólska 1975
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Bagaceratops rozhdestvenskyi Maryanska and Osmólska 1975
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Invalid names: Gobiceratops minutus Alifanov 2008 [synonym], Lamaceratops tereschenkoi Alifanov 2003 [synonym], Magnirostris dodsoni You and Dong 2003 [synonym], Platyceratops tatarinovi Alifanov 2003 [synonym]
Invalid names: Gobiceratops Alifanov 2008 [synonym], Lamaceratops Alifanov 2003 [synonym], Magnirostris You and Dong 2003 [synonym], Platyceratops Alifanov 2003 [synonym]
G. †Breviceratops Kurzanov 1990
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Breviceratops kozlowskii Maryanska and Osmólska 1975
G. †Protoceratops Granger and Gregory 1923
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Protoceratops andrewsi Granger and Gregory 1923
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Invalid names: Bainoceratops efremovi Tereshchenko and Alifanov 2003 [synonym]
Protoceratops hellenikorhinus Lambert et al. 2001
Invalid names: Bainoceratops Tereshchenko and Alifanov 2003 [synonym]
Invalid names: Asiaceratopsinae Nessov et al. 1989 [empty]
Diagnosis
ReferenceDiagnosis
F. v. Huene 1948Few teeth in posteiror part of premaxilla; 1-2 rows of single rooted teeth in jaws; large preorbit; two pairs of large temporal fossae; frontals and parietals in exterior roof of skull; short high mandibles; relatively long pubis.