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Physeterula dubusi

Mammalia - Artiodactyla - Physeteridae

Taxonomy
Physeterula dubusii was named by Van Beneden (1877) [no type specimen listed in Van Beneden (1877); information on the holotype from Lambert 2008]. Its type specimen is IRSNB M.527, a mandible (a fragmentary skull including rostral fragments of the premaxillae, parts of the supracranial basin and of the basicranium, with the associated mandible and 18 ), and it is a 3D body fossil. Its type locality is Deurne (Fort 1), Deurne Sand, which is in a Tortonian lagoonal sandstone in the Diest Formation of Belgium. It is the type species of Physeterula.

It was recombined as Kogia dubusii by Cope (1890); it was corrected as Physeterula dubusi by Lydekker (1887), Trouessart (1904), Lambert (2008) and Velez-Juarbe et al. (2015).

Synonyms
Synonymy list
YearName and author
1859Delphinus acutidens von Meyer
1861Delphinus acutidens Meyer
1874Physeterula dubusii Vanden Broeck p. 146
1876Orcopsis acutidens Van Beneden p. 489
1877Physeterula dubusii Van Beneden p. 852
1884Orcopsis acutidens Van Beneden p. 9
1887Physeterula dubusi Lydekker p. 54
1890Kogia dubusii Cope p. 608
1891Physeterula dubusii Newton p. 67
1898Beluga acutidens Studer p. 132
1898Physeterula dubusii Trouessart p. 1057
1904Physeterula dubusi Trouessart p. 774
2002Physeterula dubusii Kazár p. 163
2008Physeterula dubusi Lambert p. 293
2008Physeterula dubusii Uhen et al. p. 577
2015Physeterula dubusi Velez-Juarbe et al. p. 15 figs. Fig. 10

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RankNameAuthor
kingdomAnimalia()
Triploblastica
Nephrozoa
Deuterostomia
phylumChordataHaeckel 1847
subphylumVertebrata
superclassGnathostomata
Osteichthyes()
Sarcopterygii
subclassDipnotetrapodomorpha(Nelson 2006)
subclassTetrapodomorpha()
Tetrapoda()
Reptiliomorpha
Anthracosauria
Batrachosauria()
Cotylosauria()
Amniota
Synapsida()
RankNameAuthor
Therapsida()
infraorderCynodontia()
Epicynodontia
infraorderEucynodontia
Probainognathia
Mammaliamorpha
Mammaliaformes
classMammalia
orderArtiodactyla()
Cetacea()
Pelagiceti
Neoceti
Odontoceti()
superfamilyPhyseteroidea
familyPhyseteridae
genusPhyseterula
speciesdubusi()

If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.

Diagnosis
ReferenceDiagnosis
O. Lambert 2008Physeterula dubusi is a large physeteroid with a skull size close to Idiorophus patagonicus and Zygophyseter varolai. It differs from the other physeterids Aulophyseter and Physeter in the retention of functional maxillary teeth, the shorter anterior extension of the supracranial basin, the pre orbital process being at the same vertical level as the dorsolateral margin of the rostrum base, the frontal-maxilla suture less erected in lateral view, and lTIOre numerous dorsal infraorbital foramina. It further differs from Physeter in the maxilla being absent from the dorsal view of the rostrum for more than half its length, the elevated temporal fossa, the mandibular symphysis shorter than half the length of the mandible, and the fact that some of the caudal vertebrae are longer than wide. Finally it differs from Aulophyseter in the robust and shortened postorbital process of the frontal.