|Basic info||Taxonomic history||Classification||Relationships|
|Morphology||Ecology and taphonomy||External Literature Search||Age range and collections|
Mammalia - Carnivora - Ursidae
If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.
|L. de Bones et al. 2017||medium-sized ursid, primitive by the anteriorly situated protocone of P4 and the short talon of M2; vertical ascending ramus of the mandible; lower p1-p3 reduced but p2 and p3 two-rooted; p4 with a well-developed posterior accessory cuspid; low trigonid of m1 with blunt cuspids, mesio-distally oriented paraconid, gentle mesial and distal slopes of the protoconid, distally situated metaconid, talonid wider than trigonid; oval m3; low upper canine, small but two-rooted P3; P4 with very tiny parastyle, protocone at the same level than paracone; M2 with a small talon.|