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Gopherus laticuneus

Reptilia - Testudines - Testudinidae

Taxonomy
Testudo laticuneus was named by Cope (1873). Its type specimen is AMNH 1160, a shell (essentially complete shell with some of the internal skeleton), and it is a 3D body fossil. Its type locality is Horsetail Creek, which is in a Chadronian terrestrial horizon in the White River Formation of Colorado.

It was corrected as Testudo laticunea by Hay (1902), Hay (1930) and Gilmore (1946); it was recombined as Scaptochelys laticunea by Bramble (1982); it was recombined as Gopherus (Oligopherus) laticuneus by Hutchison (1996); it was recombined as Gopherus laticuneus by Wall and Maddox (1998) and Reynoso and Montellano-Ballesteros (2004); it was misspelled as Gopherus laticunea by Williams (1950); it was recombined as Oligopherus laticuneus by Vlachos and Rabi (2018) and Vlachos (2018).

Synonyms
Synonymy list
YearName and author
1873Testudo laticuneus Cope p. 6
1902Testudo laticunea Hay p. 451
1908Testudo thomsoni Hay
1909Testudo undabuna Loomis
1913Testudo praeextans Lambe
1930Testudo laticunea Hay p. 104
1930Testudo praeextans Hay p. 104
1930Testudo thomsoni Hay p. 105
1930Testudo undabuna Hay p. 105
1946Testudo praeextans Gilmore p. 293
1946Testudo laticunea Gilmore p. 307
1950Gopherus laticunea Williams p. 25
1982Scaptochelys laticunea Bramble p. 854
1982Stylemys undabuna Bramble p. 854
1996Gopherus (Oligopherus) laticuneus Hutchison p. 348
1998Gopherus laticuneus Wall and Maddox p. 11
2004Gopherus laticuneus Reynoso and Montellano-Ballesteros p. 824
2018Oligopherus laticuneus Vlachos
2018Oligopherus laticuneus Vlachos and Rabi p. 663

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RankNameAuthor
kingdomAnimalia()
Bilateria
EubilateriaAx 1987
Life
Deuterostomia
phylumChordataHaeckel 1874
subphylumVertebrata
superclassGnathostomata
Osteichthyes()
subclassSarcopterygii()
subclassDipnotetrapodomorpha(Nelson 2006)
subclassTetrapodomorpha()
Tetrapoda
Reptiliomorpha
RankNameAuthor
Anthracosauria
subclassAmphibiosauriaKuhn 1967
Cotylosauria()
Amniota
Sauropsida
classReptilia
Testudinata(Oppel 1811)
orderTestudinesBatsch 1788
suborderCryptodira
Pantestudinoidea
superfamilyTestudinoidea
familyTestudinidaeBatsch 1788
genusGopherusRafinesque 1832
specieslaticuneus()

If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.

Gopherus laticuneus Cope 1873
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Diagnosis
ReferenceDiagnosis
E. Vlachos 2018Oligopherus laticuneus can be diagnosed as a member of Pan-Testudinidae based on the presence of a rectangular neural III. Oligopherus laticuneus differs from other members of Pan-Testudinidae by the following combination of characters: low-domed shell, shell bones moderately thin, epiplastron flattened with a very shallow excavation, gular protrusion present, cervical scute wide as long and often wedge-shaped, pleural I not covering the nuchal, and presence of a pygal scute.