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Gopherus edae

Reptilia - Testudines - Testudinidae

Taxonomy
Testudo edae was named by Hay (1906). Its type specimen is CM 1535 (holotype), a shell, and it is a 3D body fossil. Its type locality is Running Water Creek (few miles west of Agate), which is in a Harrisonian terrestrial horizon in the Harrison Formation of Nebraska.

It was considered a nomen vanum by Reynoso and Montellano-Ballesteros (2004); it was recombined as Gopherus edae by Bramble (1982) and Vlachos (2018).

Synonyms
Synonymy list
YearName and author
1906Testudo edae Hay
1906Testudo hollandi Hay
1907Testudo edae Hay
1908Testudo vaga Hay
1908Testudo pansa Hay p. 420
1950Gopherus pansa Williams p. 26
1982Gopherus edae Bramble p. 854
1982Gopherus pansa Bramble p. 854
1982Gopherus vaga Bramble p. 854
2004Gopherus pansus Reynoso and Montellano-Ballesteros p. 824
2004Gopherus vagus Reynoso and Montellano-Ballesteros p. 824
2018Gopherus edae Vlachos

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RankNameAuthor
kingdomAnimalia()
Triploblastica
Nephrozoa
Deuterostomia
phylumChordataHaeckel 1847
subphylumVertebrata
superclassGnathostomata
Osteichthyes()
Sarcopterygii
subclassDipnotetrapodomorpha(Nelson 2006)
subclassTetrapodomorpha()
Tetrapoda()
Reptiliomorpha
Anthracosauria
RankNameAuthor
Batrachosauria()
Cotylosauria()
Amniota
Sauropsida
classReptilia
Testudinata(Oppel 1811)
orderTestudinesBatsch 1788
suborderCryptodira
Pantestudinoidea
superfamilyTestudinoidea
Pantestudinidae
familyTestudinidaeBatsch 1788
genusGopherusRafinesque 1832
speciesedae()

If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.

Diagnosis
ReferenceDiagnosis
E. Vlachos 2018Gopherus edae can be diagnosed as a member of Gopherus based on the characters mentioned above. Gopherus edae differs from Gopherus brevisternus in the presence of an octagonal neural IV and wider vertebrals, from Gopherus mohavensis in the narrower cervical, neural VI with short anterolateral sides, shorter pectorals medially, and contact between the inguinals and femorals, and from Gopherus hexagonatus in the fully differentiated neural series.