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Compsemys victa

Reptilia - Testudines - Compsemydidae

Taxonomy
Compsemys victus was named by Leidy (1856). Its type specimen is USNM 960, a partial shell (a neural and two costal fragments), and it is a 3D body fossil. Its type locality is Long Lake, which is in a Maastrichtian terrestrial horizon in the Hell Creek Formation of North Dakota.

It was misspelled as Compsemys victus by Leidy (1860) and Hay (1902); it was corrected as Compsemys victa by Leidy (1856), Leidy (1857), Estes (1964), Estes and Berberian (1970), Gaffney (1972), Estes (1976), Bryant (1989), Holroyd and Hutchison (2002), Hutchison and Holroyd (2003), Burger (2007), Lyson and Joyce (2011), Hutchison et al. (2013) and López-Conde et al. (2018).

Synonyms
Synonymy list
YearName and author
1856Compsemys victa Leidy p. 312
1856Compsemys victus Leidy p. 312
1856Emys obscurus Leidy p. 312
1857Compsemys victa Leidy p. 90
1857Emys obscurus Leidy p. 90
1860Compsemys victus Leidy p. 152
1860Emys obscurus Leidy p. 153
1902Compsemys obscurus Hay
1902Compsemys victus Hay
1908Glyptops depressus Hay figs. Text-figs. 33, 34
1908Compsemys obscura Hay p. 235 figs. Plate 34, fig. 4
1910Compsemys parva Hay
1910Compsemys vafer Hay
1910Compsemys parva Hay pp. 308-310 figs. fig. 1, pl. 10 figs. 1-3
1910Compsemys vafer Hay pp. 311-313 figs. 2-5
1919Compsemys torrejonensis Gilmore p. 21 figs. Pl. 4:1-2
1919Compsemys puercensis Gilmore p. 198 figs. Pl. 3:1-2
1964Compsemys victa Estes p. 98
1970Compsemys victa Estes and Berberian p. 5
1972Compsemys victa Gaffney pp. 249, 289 fig. 37
1976Compsemys victa Estes p. 507
1989Compsemys victa Bryant p. 41
2002Compsemys victa Holroyd and Hutchison p. 181
2003Compsemys victa Hutchison and Holroyd
2007Compsemys victa Burger p. 144
2011Compsemys victa Lyson and Joyce p. 790 figs. 1-2
2013Compsemys victa Hutchison et al.
2018Compsemys victa López-Conde et al.

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RankNameAuthor
kingdomAnimalia()
Triploblastica
Nephrozoa
Deuterostomia
phylumChordataHaeckel 1847
subphylumVertebrata
superclassGnathostomata
Osteichthyes()
Sarcopterygii
subclassDipnotetrapodomorpha(Nelson 2006)
subclassTetrapodomorpha()
Tetrapoda()
Reptiliomorpha
RankNameAuthor
Anthracosauria
Batrachosauria()
Cotylosauria()
Amniota
Sauropsida
classReptilia
Testudinata(Oppel 1811)
orderTestudinesBatsch 1788
Paracryptodira
familyCompsemydidae
genusCompsemys
speciesvicta()

If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.

Diagnosis
ReferenceDiagnosis
E. S. Gaffney 1972[as for genus Compsemys]
T. R. Lyson and W. G. Joyce 2011Placed along the phylogenetic stem of Paracryptodira by nasals that prevent frontals from entering narial opening; no medial contact of the prefrontals; foramen posterius canalis carotici interni located halfway along contact between pterygoid and basisphenoid; posterior plastral thickening medial to the bridge. Autapomorphies include: no cheek emargination; rectangular quadratojugal; quadratojugal extends ventrally to a level even with mandibular condyles of quadrate, covering up these condyles; postorbital contributes to rim of cavum tympani; cavum tympani diameter less than diameter of orbit; mesoplastra unusually large and with broad midline contact; first peripherals meet medially in front of nuchal, preventing the nuchal from being exposed along the anterior margin of carapace; deep xiphiplastral notch present, varying from Ushaped to V-shaped; sinuous midline sulcus on plastron.