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Madrynornis mirandus

Reptilia - Avetheropoda - Spheniscidae

Taxonomy
Madrynornis mirandus was named by Acosta Hospitaleche et al. (2007). Its type specimen is MEF-PV 100, a skeleton (nearly complete and articulated skeleton with well-preserved feather impressions), and it is a 3D body fossil. Its type locality is Playa Villarino, which is in a Tortonian marine horizon in the Puerto Madryn Formation of Argentina.

Synonymy list
YearName and author
2007Madrynornis mirandus Acosta Hospitaleche et al. p. 301 figs. Figs. 2-7
2008Madrynornis mirandus Acosta Hospitaleche and Tambussi p. 123
2018Madrynornis mirandus Degrange et al. p. 1

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RankNameAuthor
kingdomAnimalia()
Triploblastica
Nephrozoa
Deuterostomia
phylumChordataHaeckel 1847
subphylumVertebrata
superclassGnathostomata
Osteichthyes()
Sarcopterygii
subclassDipnotetrapodomorpha(Nelson 2006)
subclassTetrapodomorpha()
Tetrapoda()
Reptiliomorpha
Anthracosauria
Batrachosauria()
Cotylosauria()
Amniota
Sauropsida
classReptilia
subclassEureptilia()
Romeriida
Diapsida()
Eosuchia()
Neodiapsida
SauriaGauthier 1984
RankNameAuthor
Archosauromorpha(Huene 1946)
Crocopoda
ArchosauriformesGauthier 1986
Eucrocopoda
Archosauria()
informalAvemetatarsalia
Ornithodira
Dinosauromorpha
Dinosauriformes
Dinosauria()
Saurischia()
Theropoda()
Neotheropoda
AverostraPaul 2002
Tetanurae
orderAvetheropoda
suborderCoelurosauria
Maniraptora
Paraves
classAves
orderSphenisciformesSharpe 1891
familySpheniscidaeBonaparte 1831
genusMadrynornis
speciesmirandus

If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.

Diagnosis
ReferenceDiagnosis
C. Acosta Hospitaleche and C. Tambussi 2008Crista occipitalis transversa expanded
into posterolaterally directed wings (not developed in Pygo- scelis), fossa temporalis more triangular and deeper than in Paraptenodytes, processus postorbitalis slender and longer than in Spheniscus, arcus jugalis slightly curved compared to that in Eudyptes and Pygoscelis, interorbital region nar- rower than in Spheniscus and Eudyptes. Sulcus glandulae nasalis without an external edge as in Spheniscus and Par- aptenodytes (edge present in Pygoscelis); lamina parasphe- noidalis broader than in all living species. Ramus mandibiularis straight with the fossa retroarticularis longer than in Paraptenodytes and Spheniscus, and extending beyond the fossa articularis quadratica.