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Gastropoda - Cypraeidae

Bernaya was named by Jousseaume (1884) [Sepkoski's age data: J Tith T Ol-l Sepkoski's reference number: 121]. Its type is Cypraea media.

It was reranked as Cypraea (Bernayia) by Oppenheim (1906) and Vredenburg (1925); it was reranked as Cypraea (Bernaya) by Islamoglu and Taner (2002); it was synonymized subjectively with Cypraeorbis by Darragh (2011).

It was assigned to Cypraeorbini by Gliozzi and Malatesta (1983); to Cypraeidae by Cossmann (1891), Abbass (1967), Groves (1990) and Squires and Demetrion (1992); to Cypraea by Oppenheim (1906), Vredenburg (1925) and Islamoglu and Taner (2002); to Neotaenioglossa by Sepkoski (2002); to Bernayini by Groves (2004) and Groves (2011); and to Bernayinae by Schilder (1927), Perrilliat et al. (2003), Bouchet and Rosenberg (2016) and Groves and Squires (2023).

Synonymy list
YearName and author
1884Bernaya Jousseaume
1891Bernayia Cossmann
1906Cypraea (Bernayia) Oppenheim p. 301
1925Cypraea (Bernayia) Vredenburg p. 303
1927Bernaya Schilder
1967Bernaya Abbass p. 64
1983Bernaya Gliozzi and Malatesta p. 103
1990Bernaya Groves p. 277
1992Bernaya Squires and Demetrion p. 31
2002Cypraea (Bernaya) Islamoglu and Taner p. 49
2002Bernaya Sepkoski
2003Bernaya Perrilliat et al. p. 42
2004Bernaya Groves p. 46
2011Bernaya Groves p. 47
2016Bernaya Bouchet and Rosenberg
2023Bernaya Groves and Squires p. 12

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EubilateriaAx 1987
subclassCaenogastropoda(Cox 1959)
Sorbeoconcha(Ponder and Lindberg 1997)
Hypsogastropoda(Ponder and Lindberg 1997)
superorderLatrogastropodaRiedel 2000
superfamilyCypraeoidea(Rafinesque 1815)
familyCypraeidaeRafinesque 1815
subfamilyBernayinaeSchilder 1927
genusBernayaJousseaume 1884

If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.

G. †Bernaya Jousseaume 1884
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Subg. †Bernaya (Bernaya) Jousseaume 1884
Bernaya (Bernaya) azevedoi Oliveira 1957
Bernaya (Bernaya) beardi Groves 2004
Bernaya (Bernaya) depereti Roman and Mazeran 1920
Bernaya (Bernaya) ellipsoides d'Archiac and Haime 1853
Bernaya (Bernaya) jeanae Groves 2004
Bernaya (Bernaya) obesa Deshayes 1866
Bernaya grovesi Squires and Demetrion 1992
Bernaya kaylinae Groves and Squires 2023
Bernaya media Deshayes 1835
Bernaya quarunensis Abbass 1967
Bernaya squiresi Groves 2011
L. T. Groves 1990Shell medium to large size, anterior end somewhat carinate, dorsum smooth, spire of medium height and partially covered, aperture wide, sides rounded, anterior and posterior canals deep, fossula smooth, concave, wide.
L. T. Groves 2004Shell medium to large in size, anterior end somewhat carinate, dorsum smooth, spire of medium height and partially covered by successive whorls, aperture wide, sides rounded, anterior and posterior canals deep, fossula smooth, concave, wide.
L. T. Groves 2011Shell medium to large size; anterior endsomewhat carinate; dorsum smooth; spire of mediumheight and partially covered; aperture wide, sidesrounded; anterior and posterior canals deep; fossulasmooth, concave, wide.
L. T. Groves and R. L. Squires 2023Shell medium to large in size. Oval, sides inflated, rounded. Dorsum smooth. Spire of small to medium height and partially covered by successive whorls. Aperture wide. Anterior and posterior canals deep. Anterior end smooth, concave, and broad (Jousseaume 1884: p. 88).
No measurements are available
Composition: aragonitec
Locomotion: actively mobilef
Life habit: epifaunalf
Diet: omnivoref
Diet 2: grazerf
Vision: limitedc
Created: 2009-09-10 07:50:11
Modified: 2009-09-10 09:50:11
Source: f = family, c = class
References: Kiessling 2004, Hendy et al. 2009

Age range

Maximum range based only on fossils: base of the Tithonian to the top of the Bartonian or 152.10000 to 38.00000 Ma
Minimum age of oldest fossil (stem group age): 145.0 Ma

Collections (46 total)

Time interval Ma Country or state Original ID and collection number
Tithonian152.1 - 145.0Italy (Palermo) Cypraea gemmellaroi (179803)
Late/Upper Barremian130.0 - 125.45France (Barcelonne) Pseudocassis helveticus (179817)
Cenomanian - Turonian100.5 - 89.8USA (California) Cypraea argonautica (176025)
Turonian93.9 - 89.8USA (Oregon) Cypraea argonautica (86814)
Turonian93.9 - 89.8USA (California) B. argonautica (176024) Cypraea berryessae (176026)
Late/Upper Turonian93.5 - 89.3France (Provence-Alpes-Cote d'Azur) B. depereti (87462)
Coniacian89.8 - 86.3India (Tamil Nadu) Cypraea kayei (180663)
Coniacian89.8 - 86.3USA (California) Cypraea kayei (44233)
Late/Upper Santonian - Early/Lower Campanian85.8 - 70.6Canada (British Columbia) B. beardi (176395)
Campanian83.6 - 72.1India (Tamil Nadu) Cypraea cunliffei, Erato veraghoorensis (179034)
Campanian83.6 - 72.1Brazil (Pernambuco) Cypraea azevedoi (179815 180661)
Early/Lower Campanian83.5 - 70.6USA (California) B. jeanae (176365) B. popenoei (176369)
Early/Lower Campanian - Middle Campanian83.5 - 70.6USA (Mississippi) B. n. sp. (7456)
Late/Upper Campanian83.5 - 70.6USA (California) B. crawfordcatei (176396) B. crawfordcatei, B. gualalaensis, B. rineyi (60806) B. gualalaensis (99313 176400 176402) B. rineyi (99312)
Late/Upper Campanian83.5 - 70.6USA (Delaware) B. mortoni (6146)
Middle Campanian83.5 - 70.6USA (California) B. louellasaulae, B. popenoei (176363)
Middle Campanian - Late/Upper Campanian83.5 - 70.6USA (New Jersey) Palaeocypraea burlingtonensis (180662)
Late/Upper Campanian - Early/Lower Maastrichtian83.5 - 66.0USA (California) B. gualalaensis (176397 176399 176401) Cypraea gualalaensis (176398)
Maastrichtian72.1 - 66.0USA (New Jersey) B. burlingtonensis (78124)
Early/Lower Maastrichtian70.6 - 66.0Pakistan (Balochistan) Ovula baluchistanensis (180660)
Danian66.0 - 61.6Denmark B. globuliformis (176994) Protocypraea globuliformis (137438)
Paleocene66.0 - 56.0USA (California) B. sp. (194098)
Paleocene66.0 - 56.0Germany B. sp. (3115)
Selandian - Thanetian61.6 - 56.0Pakistan (North-West Frontier) Protocypraea kohatica (52132)
Ypresian56.0 - 47.8Mexico (Baja California Sur) B. grovesi (5496)
Ypresian56.0 - 47.8USA (Washington) B. grovesi (176361)
Ypresian - Lutetian56.0 - 41.3USA (California) B. kaylinae (8014) Eocypraea castacensis (183752)
Early/Lower Eocene55.8 - 48.6China (Tibet) Ovula ellipsoides (76993)
Lutetian47.8 - 41.3Mexico (Chiapas) B. obesa, B. media (79943)
Lutetian47.8 - 41.3USA (California) B. kaylinae (8012 173475)
Bartonian41.3 - 38.0France (Île-de-France) B. obesa (179130)
Bartonian41.3 - 38.0Egypt B. quarunensis (2960)