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Onychocellidae
Taxonomy
Onychocellidae was named by Jullien (1882). It is extant. Its type is Onychocella.
It was assigned to Coilostega by Bassler (1953); to Membraniporidae by Brown (1958); to Flustrina by Gordon and Taylor (1999); to Anasca by Cheetham (1962) and Pouyet (2000); to Cheilostomata by Bock (2006); and to Microporoidea by Zágoršek and Kroh (2003), Schmidt (2003), Berning (2006), Di Martino and Taylor (2013), Di Martino and Taylor (2014), Gordon and Taylor (2015), Taylor and Martha (2017) and Koromyslova and Seltser (2020).
It was assigned to Coilostega by Bassler (1953); to Membraniporidae by Brown (1958); to Flustrina by Gordon and Taylor (1999); to Anasca by Cheetham (1962) and Pouyet (2000); to Cheilostomata by Bock (2006); and to Microporoidea by Zágoršek and Kroh (2003), Schmidt (2003), Berning (2006), Di Martino and Taylor (2013), Di Martino and Taylor (2014), Gordon and Taylor (2015), Taylor and Martha (2017) and Koromyslova and Seltser (2020).
Subtaxa
Aechmella, Cheethamia, Chondriovelum, Collura, Distefanella, Ehrhardina, Euritina, Floridina, Floridinella, Hoplitaechmella, Hoplocheilina, Inversaria, Nudicella, Ogiva, Ogivalia, Onychocella (syn. Reptocelleporaria) (type genus), Paraechmellina, Reptolunulites, Rhagasostoma, Rhebasia, Semieschara, Smittipora (syn. Rectonychocella, Diplopholeous, Velumella), Thyracella
Synonymy list
| Year | Name and author |
|---|---|
| 1882 | Onychocellidae Jullien |
| 1953 | Onychocellidae Bassler p. G168 |
| 1958 | Onychocellidae Brown |
| 1962 | Onychocellidae Cheetham p. 324 |
| 1999 | Onychocellidae Gordon and Taylor |
| 2000 | Onychocellidae Pouyet |
| 2003 | Onychocellidae Schmidt |
| 2003 | Onychocellidae Zágoršek and Kroh p. 402 |
| 2006 | Onychocellidae Berning p. 40 |
| 2006 | Onychocellidae Bock |
| 2013 | Onychocellidae Di Martino and Taylor p. 87 |
| 2014 | Onychocellidae Di Martino and Taylor |
| 2015 | Onychocellidae Gordon and Taylor |
| 2017 | Onychocellidae Taylor and Martha p. 26 |
| 2020 | Onychocellidae Koromyslova and Seltser p. 701 |
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If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.
Fm. Onychocellidae Jullien 1882
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G. †Aechmella Canu and Bassler 1917
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†Aechmella ambigua MacGillivray 1895
†Aechmella depressa MacGillivray 1895
†Aechmella poluzzii Moissette 1988
†Aechmella rangiauriensis Taylor and Gordon 2007
G. †Cheethamia
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†Cheethamia aktolagayensis Koromyslova et al. 2018
†Cheethamia dividia Håkansson et al. 2024
†Cheethamia volgaensis Koromyslova and Seltser 2020
G. Chondriovelum Hayward and Thorpe 1988
G. †Collura
G. †Distefanella Cipolla 1922
G. †Ehrhardina Martha and Taylor 2016
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†Ehrhardina pikeae Martha and Taylor 2016
†Ehrhardina voigti Martha and Taylor 2016
G. Floridina Jullien 1881
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†Floridina asymmetrica Canu and Bassler 1920
†Floridina elegans Gordon and Taylor 2015
†Floridina pentagonus Guha and Gopikrishna 2005
G. †Hoplitaechmella
G. †Nudicella Schmidt and Bone 2004
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†Nudicella dissidens Håkansson et al. 2024
†Nudicella fossilis Gordon and Taylor 1999
G. †Ogiva Jullien 1881
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†Ogiva concamerata Waters 1881
†Ogiva ellinorvoigtae Martha et al. 2015
†Ogiva elongata Canu and Bassler 1935
†Ogiva incompta Gordon and Taylor 1999
†Ogiva promonturiorum Voigt 1924
G. Ogivalia
G. Onychocella Jullien 1882
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Onychocella angulosa Reuss 1848
†Onychocella cyclostoma Goldfuss 1826
†Onychocella depressa von Hagenow 1851
†Onychocella lamellosa Gordon and Taylor 1999
†Onychocella michaudiana d'Orbigny 1850
†Onychocella pseudoirregularis Voigt 1924
†Onychocella reussi Prantl 1938
†Onychocella spinifera Voigt 1981
†Onychocella subtriangulata Gordon and Taylor 2015
†Onychocella torquata Guha and Gopikrishna 2005
Invalid names: Reptocelleporaria D'Orbigny [synonym]
G. †Reptolunulites d'Orbigny 1852
G. †Rhagasostoma
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†Rhagasostoma angliae Brydone 1936
†Rhagasostoma aralense Koromyslova et al. 2018
†Rhagasostoma brydonei Koromyslova et al. 2018
†Rhagasostoma gibbosulum Brydone 1936
†Rhagasostoma gibbosum Marsson 1887
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Invalid names: Rhagasostoma gibbosum weybournensis Brydone 1930 [synonym], Rhagasostoma subgibbosum Brydone 1930 [synonym]
†Rhagasostoma inelegans Lonsdale 1850
†Rhagasostoma infelix Håkansson et al. 2024
†Rhagasostoma mimosa Brydone 1930
†Rhagasostoma minuens Brydone 1936
†Rhagasostoma operculatum Koromyslova et al. 2018
†Rhagasostoma saltans Brydone 1930
†Rhagasostoma tchvanovi Favorskaya 1992
G. †Rhebasia
G. †Semieschara
G. Smittipora Jullien 1882
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Smittipora cordiformis Harmer 1926
Invalid names: Diplopholeous [synonym], Rectonychocella [synonym], Velumella [synonym]
G. †Thyracella
Diagnosis
No diagnoses are available
Measurements
No measurements are available
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| Source: f = family, o = order, c = class, uc = unranked clade | |||||
| References: Smith et al. 2006, Nesnidal et al. 2013 | |||||
Age range
Maximum range based only on fossils: base of the Late/Upper Albian to the top of the Holocene or 105.30000 to 0.00000 Ma
Minimum age of oldest fossil (stem group age): 100.5 Ma
Minimum age of oldest fossil (stem group age): 100.5 Ma
Collections (168 total)
| Time interval | Ma | Country or state | Original ID and collection number |
|---|---|---|---|
| Albian | United Kingdom (England) | Inversaria orbicularis (1816) | |
| Late/Upper Albian | USA (Texas) | Thyracella sp. (65727) | |
| Cenomanian | United Kingdom (England) | Ehrhardina voigti (220517 220518) | |
| Cenomanian | Germany (Nordrhein-Westfalen) | Ehrhardina pikeae (94265) | |
| Early/Lower Cenomanian | United Kingdom (England) | Ehrhardina voigti (220514) Euritina denticulata (220520) Euritina sp., Ehrhardina voigti (220519) | |
| Middle Cenomanian | United Kingdom (England) | Aechmella sp., Onychocella sp. (220516) | |
| Early/Lower Turonian | Czech Republic | Onychocella novaki, Onychocella reussi (90517 90518) | |
| Late/Upper Turonian | United Kingdom (England) | Rhagasostoma brydonei (216925) | |
| Coniacian | Algeria | Membranipora ambigua (71826) | |
| Coniacian | France | Onychocella acis, Onychocella aegon, Onychocella cyntia (3785) | |
| Coniacian | United Kingdom (England) | Rhagasostoma brydonei, Rhagasostoma minuens (216926) | |
| Coniacian - Santonian | France | Onychocella sp. (3736) | |
| Santonian | Germany | Ogiva ellinorvoigtae, Ogiva promonturiorum (175916) Ogiva promonturiorum (175915 175917) | |
| Santonian | United Kingdom (England) | Flustra inelegans (216920) | |
| Santonian | France | Onychocella aegle, Onychocella santonensis, Semieschara norvicia, Smittipora sp. (3786) | |
| Santonian - Maastrichtian | Russian Federation (Ulyanovsk) | Eschara volgensis (216948 216949) | |
| Late/Upper Santonian | Austria | Onychocella depressa, Onychocella michaudiana, Onychocella pseudoirregularis, Onychocella reussi, Onychocella cyclostoma (61374) | |
| Late/Upper Santonian | United Kingdom (England) | Rhagasostoma inelegans (216921) Rhagasostoma minuens (216927) | |
| Middle Santonian | Germany (Niedersachsen) | Ogiva ellinorvoigtae, Ogiva promonturiorum (175914) | |
| Campanian | Turkmenistan | Rhagasostoma operculatum (216938) | |
| Campanian | France | Onychocella claudia, Onychocella girondina, Semieschara norvicia (3787) Rhagasostoma minuens (216930) | |
| Campanian - Maastrichtian | Mexico (Baja California) | Onychocella schopforum (87374) | |
| Early/Lower Campanian | USA (Arkansas) | Aechmella ozanensis (90501) | |
| Early/Lower Campanian | Germany (Niedersachsen) | Rhagasostoma inelegans (216922) | |
| Early/Lower Campanian | Germany (Schleswig-Holstein) | Rhagasostoma minuens (216931) | |
| Early/Lower Campanian | United Kingdom (England) | Rhagasostoma inelegans (216899) Rhagasostoma minuens (216928 216929) | |
| Early/Lower Campanian | Germany (Sachsen-Anhalt) | Ogiva promonturiorum (112629) | |
| Early/Lower Campanian | Uzbekistan (Karakalpakstan) | Rhagasostoma aralense (216936) | |
| Late/Upper Campanian | Belgium (Hainaut) | Rhagasostoma rowei (216947) | |
| Late/Upper Campanian | Turkmenistan | Rhagasostoma rowei (216951) | |
| Late/Upper Campanian | Uzbekistan (Karakalpakstan) | Onychocella tchvanovi, Rhagasostoma rowei (216942) Rhagasostoma aralense, Onychocella tchvanovi (216937) | |
| Late/Upper Campanian | Kazakhstan | Rhagasostoma gibbosum, Rhagasostoma rowei (216940) | |
| Late/Upper Campanian | United Kingdom (England) | Rhagasostoma angliae (216932 216933) Rhagasostoma gibbosulum (216941) | |
| Late/Upper Campanian | Belarus | Rhagasostoma gibbosulum, Rhagasostoma inelegans, Rhagasostoma gibbosum, Rhagasostoma rowei (216924) | |
| Late/Upper Campanian | Germany | Rhagasostoma angliae (216935) | |
| Late/Upper Campanian | France | Rhagasostoma angliae (216934) | |
| Maastrichtian | USA (California) | Aechmella sp., Reptolunulites sp. (87393) | |
| Early/Lower Maastrichtian | Germany (Schleswig-Holstein) | Rhagasostoma gibbosulum, Rhagasostoma gibbosum (216939) | |
| Early/Lower Maastrichtian | USA (Arkansas) | Cheethamia howei (90500) | |
| Early/Lower Maastrichtian | United Kingdom (England) | Rhagasostoma minuens, Rhagasostoma angliae, Eschara rowei, Rhagasostoma gibbosum (159329) | |
| Early/Lower Maastrichtian | Australia (Western Australia) | Cheethamia dividia (235529 235530) Floridinella sp., Paraechmellina simulata (235527) Floridinella sp., Paraechmellina simulata, Cheethamia dividia, Rhagasostoma infelix (235528) Nudicella dissidens, Cheethamia dividia (235532) Rhagasostoma infelix (235531) | |
| Early/Lower Maastrichtian | Russian Federation (Saratov) | Hoplitaechmella sp. 1, Hoplitaechmella sp. 2, Cheethamia aktolagayensis, Cheethamia volgaensis (215304) Rhagasostoma saltans (215305) | |
| Late/Upper Maastrichtian | Turkmenistan | Rhagasostoma rowei (216950 216952) | |
| Late/Upper Maastrichtian | Netherlands (Limburg) | Onychocella spinifera (123187) | |
| Danian | Austria (Niederösterreich) | Onychocella disparilis (87001 87002) | |
| Danian | Denmark | Aechmella pindborgi, Aechmella latistoma, Floridina gothica, Onychocella columella, Smittipora prismatica, Thyracella sp. (2964) Floridina gothica (226854) Onychocella irregularis, Onychocella faxensis, Rhagasostoma elegans, Rhagasostoma nysti (89866) | |
| Danian | USA (Alabama) | Euritina tecta (83261 83297) | |
| Danian | Sweden | Aechmella pindborgi, Aechmella tenuis, Floridina gothica, Floridina sp., Floridina voigti, Onychocella ravni, Onychocella columella, Onychocella poulseni, Smittipora prismatica, Smittipora sp. (69603) | |
| Paleocene | USA (New Jersey) | Euritina torta (84051) | |
| Thanetian - Ypresian | New Zealand (Chatham Island) | Hoplitaechmella sp., Onychocella sp., Inversaria gondwanae, Chondriovelum fossilis, Ogiva incompta, Onychocella lamellosa (7221) | |
| Early/Lower Eocene | New Zealand (Chatham Island) | Floridina sp., Floridina elegans, Onychocella subtriangulata (172944) | |
| Lutetian - Bartonian | India (Gujarat) | Onychocella torquata (177175) | |
| Middle Eocene | United Kingdom (England) | Onychocella subpyriformis (3263) | |
| Middle Eocene | India | Onychocella torquata (178700) | |
| Bartonian | USA (Florida) | Floridina antiqua (7332) Floridina antiqua, Smittipora sagittellaria, Smittipora elliptica, Smittipora plicata (7330) Floridina bifoliata, Smittipora plicata, Smittipora elliptica (7321) Smittipora elliptica (7329) Smittipora levigata, Smittipora fusiformis, Smittipora tenuis (7324) Smittipora levigata, Smittipora tenuis, Smittipora fusiformis (7322) Smittipora tenuis (7327) Smittipora tenuis, Smittipora levigata (7325) | |
| Bartonian | USA (Alabama) | Floridina antiqua, Smittipora tenuis (7331) | |
| Priabonian | USA (Mississippi) | Floridina granulosa, Floridina bifoliata, Onychocella celsa, Rectonychocella tenuis (84174) | |
| Priabonian | USA (Florida) | Smittipora tenuis, Smittipora levigata (7318) | |
| Priabonian | New Zealand (North Otago) | Ogivalia sp. (172815) | |
| Late/Upper Eocene | USA (Georgia) | Floridina asymmetrica (90479) | |
| Rupelian | USA (Alabama) | Floridina antiqua, Rhagasostoma levigata (90563) Floridinella vicksburgica (90528 90549 90550 90561 90562) Rectonychocella tenuis (90548) | |
| Rupelian | USA (Mississippi) | Floridina antiqua (90565) | |
| Oligocene - Miocene | Australia (Victoria) | Aechmella ambigua, Ogiva concamerata (173510) Aechmella depressa (173521) Ogiva concamerata (173509 173512 173526 173527) Onychocella sp., Ogiva concamerata (173508) | |
| Miocene - Oligocene | Australia (Victoria) | Aechmella depressa (173516) Aechmella depressa, Aechmella ambigua (173517 173519 173522) Aechmella depressa, Aechmella ambigua, Ogiva concamerata (173520) Ogiva concamerata (173502 173504 173506 173511 173528) Ogiva concamerata, Ogiva elongata (173525) Ogiva elongata (173518) | |
| Aquitanian | India (Gujarat) | Onychocella torquata, Floridina pentagonus (177173) | |
| Aquitanian | Colombia | Smittipora sp. (211331) | |
| Early/Lower Miocene | India | Onychocella torquata, Floridina pentagonus (178702) | |
| Burdigalian | India (Gujarat) | Ogivalia vinjhanensis (46308) | |
| Burdigalian | Egypt | Onychocella angulosa (47007) | |
| Burdigalian | Germany | Onychocella sp. (188985) | |
| Burdigalian - Langhian | Indonesia | Smittipora sp. (173652) | |
| Langhian | Australia (Victoria) | Omoiosia elongata, Rectonychocella dimorphocella, Floridinella australiensis (174139) | |
| Middle Miocene | Czech Republic | Onychocella angulosa (178887 178888 178890 178892 178893 178894 178898) | |
| Middle Miocene | Australia (Victoria) | Velumella depressa (166365) | |
| Middle Miocene | Romania | Onychocella angulosa (178090) | |
| Serravallian | Indonesia | Smittipora cordiformis (173658) | |
| Serravallian | Hungary | Onychocella angulosa (34739) | |
| Tortonian | Spain | Onychocella angulosa (177482 177483 177485 177486 178465) Onychocella angulosa, Aechmella poluzzii (177481) | |
| Tortonian | Panama | Smittipora levinseni (61342) | |
| Tortonian | Indonesia | Smittipora cordiformis (173657) | |
| Messinian | Morocco | Onychocella angulosa (34723 34724 70928) | |
| Messinian | Spain | Onychocella angulosa, Aechmella poluzzii (177487) Onychocella simile (56687) | |
| Messinian | Algeria | Onychocella angulosa (70785) | |
| Zanclean | Portugal | Onychocella angulosa (152330) | |
| Pliocene | Spain | Onychocella angulosa (177490 177491) Onychocella angulosa, Aechmella poluzzii (177489) | |
| Piacenzian | Panama (Bocas) | Smittipora levinseni (79983) | |
| Piacenzian | Panama (Bocas del Toro) | Smittipora levinseni (79998) | |
| Gelasian | Greece | Onychocella angulosa (71243) | |
| Early/Lower Pleistocene | Italy | Onychocella marioni (146293) | |
| Calabrian | Costa Rica (Limon) | Floridina minima (79835) Floridinella parvula, Onychocella angulosa (79842) Smittipora acutirostris, Onychocella angulosa (79845) | |
| Holocene | USA (Florida) | Smittipora levinseni (97606 97607 97608 97609 97610) |