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Hadrianus majusculus

Reptilia - Testudines

Taxonomy
Hadrianus majusculus was named by Hay (1904). Its type specimen is YPM 927 and is a 3D body fossil. Its type locality is Badlands near Gallina, which is in a Wasatchian terrestrial horizon in the San Jose Formation of New Mexico.

It was recombined as Testudo (Hadrianus) majusculus by Williams (1953); it was recombined as Geochelone (Manouria) majusculus by Auffenberg (1971).

Synonymy list
YearName and author
1904Hadrianus majusculus Hay
1930Hadrianus majusculus Hay p. 100
1953Testudo (Hadrianus) majusculus Williams p. 548
1971Geochelone (Manouria) majusculus Auffenberg p. 115
2015Hadrianus majusculus Lichtig and Lucas
2018Hadrianus majusculus Vlachos

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RankNameAuthor
kingdomAnimalia()
Bilateria
EubilateriaAx 1987
Deuterostomia
phylumChordataHaeckel 1874
subphylumVertebrata
superclassGnathostomata
Osteichthyes()
subclassSarcopterygii()
subclassDipnotetrapodomorpha(Nelson 2006)
subclassTetrapodomorpha()
Tetrapoda
Reptiliomorpha
Anthracosauria
RankNameAuthor
subclassAmphibiosauriaKuhn 1967
Cotylosauria()
Amniota
Sauropsida
classReptilia
Testudinata(Oppel 1811)
orderTestudinesBatsch 1788
suborderCryptodira
Pantestudinoidea
superfamilyTestudinoidea
Pantestudinidae
genusHadrianus
speciesmajusculus

If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.

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Diagnosis
ReferenceDiagnosis
A. J. Lichtig and S. G. Lucas 2015A turtle of the genus Hadrianus with very high peripherals and pectoral scutes more than half as long along the midline as the abdominals. Further, the striated surface sculpture of this species is unique among the turtles of the San Jose Formation.
E. Vlachos 2018Hadrianus majusculus can be diagnosed as a member of Hadrianus based on the absence of an epiplastral lip and the gulars overlapping the anterior part of the entoplastron. Hadrianus majusculus differs from other Hadrianus in the presence of higher peripherals, a small entoplastron that is longer than wide, a posteriorly concave humero-pectoral sulcus on both sides, and a laterally concave and widely omega-shaped femoro-anal sulcus.