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Berothidae (beaded lacewing)
Taxonomy
Berothidae was named by Handlirsch (1906). It is extant.
It was assigned to Planipennia by Carpenter (1992); to Neuroptera by Martins-Neto (2002); to Mantispidea by Engel and Grimaldi (2008); and to Mantispoidea by Engel et al. (2018).
It was assigned to Planipennia by Carpenter (1992); to Neuroptera by Martins-Neto (2002); to Mantispidea by Engel and Grimaldi (2008); and to Mantispoidea by Engel et al. (2018).
Subtaxa
Aggregataberotha, Ansoberotha, Araripeberotha, Banoberotha, Berotha, Berothinae, Cantabroberotha, Caririberotha, Cornoberotha, Cyrenoberothinae, Dasyberotha, Dolichoberotha, Epimesoberotha, Ethiroberotha, Ferganoberotha, Haploberotha, Iceloberotha, Jersiberotha, Khasurtoberotha, Krokhathone, Maculaberotha, Magniberotha, Mesithoninae, Mesoberotha (syn. Proberotha), Microberotha, Nascimberotha, Oloberotha, Osmyloberotha, Proberothella, Protoberotha, Rhachiberothinae, Sejunctaberotha, Sibelliberotha, Sinosmylites, Systenoberotha, Telistoberotha, Xiaoberotha
Synonyms
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Synonymy list
Year | Name and author |
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1906 | Berothidae Handlirsch p. 42 |
1955 | Proberothidae Riek p. 674 |
1991 | Mesoberothidae Carpenter p. 87 |
1992 | Berothidae Carpenter |
1992 | Mesoberothidae Carpenter |
2002 | Berothidae Martins-Neto p. 37 |
2008 | Berothidae Engel and Grimaldi |
2008 | Mesoberothidae Engel and Grimaldi |
2018 | Berothidae Engel et al. |
2018 | Mesoberothidae Engel et al. |
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If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.
Fm. Berothidae Handlirsch 1906 [beaded lacewing]
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G. †Aggregataberotha Wang et al. 2022
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†Aggregataberotha paucipunctata Chen et al. 2024
†Aggregataberotha punctata Wang et al. 2022
G. †Araripeberotha Martins-Neto and Vulcano 1990
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†Araripeberotha fairchildi Martins-Neto and Vulcano 1990
G. Berotha Walker 1860
Subfm. Berothinae Handlirsch 1906
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G. †Cantabroberotha Pérez-de la Fuente et al. 2020
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†Cantabroberotha soplaensis Pérez-de la Fuente et al. 2020
G. †Caririberotha Martins-Neto and Vulcano 1990
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†Caririberotha martinsi Martins-Neto and Vulcano 1990
G. †Cornoberotha Yang et al. 2019
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†Cornoberotha anomala Yang et al. 2019
†Cornoberotha aspoeckae Yang et al. 2019
†Cornoberotha monogona Yang et al. 2019
G. †Dolichoberotha Yang et al. 2019
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†Dolichoberotha bifurcata Yang et al. 2019
†Dolichoberotha burmana Yang et al. 2019
G. †Haploberotha Engel and Grimaldi 2008
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†Haploberotha carsteni Makarkin 2018
†Haploberotha persephone Engel and Grimaldi 2008
G. †Iceloberotha Engel and Grimaldi 2008
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†Iceloberotha kachinensis Engel and Grimaldi 2008
†Iceloberotha simulatrix Engel and Grimaldi 2008
G. †Jersiberotha Grimaldi 2000
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†Jersiberotha luzzii Grimaldi 2000
†Jersiberotha musivum Zhang et al. 2022
†Jersiberotha myanmarensis Engel and Grimaldi 2008
†Jersiberotha similis Grimaldi 2000
†Jersiberotha tauberorum Engel and Grimaldi 2008
Subfm. †Mesithoninae Panfilov 1980
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G. †Mesithone Panfilov 1980
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†Mesithone angusta Makarkin 1999
†Mesithone grandis Panfilov 1980
†Mesithone maculata Panfilov 1980
†Mesithone maculosa Makarkin 1999
†Mesithone magna Panfilov 1980
†Mesithone proberotha Makarkin 1999
G. †Mesoberotha Carpenter 1991
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†Mesoberotha superba Riek 1955
Invalid names: Proberotha Riek 1955 [replaced]
G. †Microberotha Archibald and Makarkin 2004
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†Microberotha macculloughi Archibald and Makarkin 2004
G. †Sejunctaberotha Chen et al. 2024
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†Sejunctaberotha sphaerica Chen et al. 2024
†Sejunctaberotha tenuis Chen et al. 2024
†Sejunctaberotha transversa Chen et al. 2024
G. †Sinosmylites Hong 1983
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†Sinosmylites auliensis Khramov 2015
†Sinosmylites fumosus Makarkin et al. 2011
†Sinosmylites hotgoricus Khramov 2015
†Sinosmylites karatavicus Khramov 2015
†Sinosmylites pectinatus Hong 1983
†Sinosmylites rasnitsyni Makarkin et al. 2011
G. †Xiaoberotha Shi et al. 2019
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†Xiaoberotha bipunctata Shi et al. 2019
†Xiaoberotha dilatapoda Wang et al. 2022
Invalid names: Cyrenoberothinae Macleod and Adams 1967 [empty], Mesoberothidae Carpenter 1991 [synonym], Rhachiberothinae Tjeder 1959 [empty]
Diagnosis
No diagnoses are available
Measurements
No measurements are available
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Source: c = class, p = phylum | |||||
References: Kiessling 2004, Bush and Bambach 2015 |
Age range
Maximum range based only on fossils: base of the Ladinian to the top of the Priabonian or 242.00000 to 33.90000 Ma
Minimum age of oldest fossil (stem group age): 237.0 Ma
Minimum age of oldest fossil (stem group age): 237.0 Ma
Collections (48 total)
Time interval | Ma | Country or state | Original ID and collection number |
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Ladinian | Kyrgyzstan (Osh) | Ferganoberotha minutissima (105007) | |
Norian | Australia (Queensland) | Proberotha superba, Proberothella elongata (106484) | |
Callovian - Oxfordian | China (Nei Mongol) | Sinosmylites sp., Sinosmylites rasnitsyni, Sinosmylites fumosus (117610) | |
Callovian - Oxfordian | Kazakhstan (Chimkent) | Berothone protea (106279) Krokhathone tristis (106101) Mesithone gracilis (105970) Mesithone maculata, Mesithone magna, Berothone protea (113439) Mesithone protea, Mesithone grandis, Krokhathone parva, Sinosmylites auliensis (105971) Sinosmylites karatavicus (105426) | |
Callovian - Oxfordian | China (Liaoning) | Sinosmylites pectinatus (154730) | |
Tithonian | Mongolia (Dund-Gov’ Aymag) | Sinosmylites hotgoricus (122321) | |
Early/Lower Berriasian | United Kingdom (England) | Epimesoberotha parva (122911 122913) | |
Early/Lower Barremian | Lebanon | Berothidae indet., Banoberotha enigmatica (123001) Sibelliberotha rihanensis (153333) | |
Late/Upper Barremian | China (Liaoning) | Oloberotha sinica (138436) | |
Aptian | Russian Federation (Buryatia) | Khasurtoberotha bellissima (113350) Mesithone angusta (154573) Mesithone maculosa (122548) Mesithone proberotha (119498 123465) Pseudosisyra minima (122549) | |
Late/Upper Aptian | Brazil (Ceará) | Araripeberotha fairchildi (123013) Caririberotha martinsi (125511) | |
Early/Lower Aptian | Russian Federation (Chita) | Mesithone sp. (119499) | |
Early/Lower Albian | Spain | Cantabroberotha soplaensis (109438) | |
Late/Upper Albian | Spain (Álava) | Berothidae indet., Berothidae indet. 1, Berothidae indet. 2, Berothidae indet. 3, Berothidae indet. 4 (108769) | |
Late/Upper Albian | Spain (Aragón) | Berothidae indet. (91626) | |
Early/Lower Cenomanian | Myanmar | Ansoberotha jiewenae, Cornoberotha aspoeckae, Cornoberotha monogona, Dolichoberotha bifurcata, Dolichoberotha burmana, Xiaoberotha sp. (161726) Berothidae indet., Berothidae larva indet., Jersiberotha myanmarensis, Jersiberotha tauberorum, Iceloberotha kachinensis, Iceloberotha simulatrix, Haploberotha persephone, Telistoberotha libitina, Dasyberotha eucharis, Systenoberotha magillae, Ethiroberotha elongata (111688) Cornoberotha anomala, Cornoberotha aspoeckae, Dolichoberotha bifurcata, Xiaoberotha bipunctata (178121) Haploberotha carsteni (172900) Haploberotha persephone (123116) Jersiberotha musivum (216019) Maculaberotha nervosa, Magniberotha recurrens, Protoberotha minuta, Ansoberotha jiewenae, Aggregataberotha punctata, Xiaoberotha dilatapoda, Aggregataberotha paucipunctata, Sejunctaberotha sphaerica, Sejunctaberotha tenuis, Sejunctaberotha transversa (165561) Osmyloberotha simpla (190961) Xiaoberotha bipunctata (178159) | |
Turonian | USA (New Jersey) | Jersiberotha luzzii, Jersiberotha similis, Nascimberotha picta (110895) | |
Late/Upper Campanian | Myanmar | Berothidae indet. (195455) | |
Middle Campanian | Canada (Alberta) | Berothidae indet. (104932) Chrysopidae indet. (122384) | |
Ypresian | Canada (British Columbia) | Microberotha macculloughi (123795) | |
Uintan | USA (Colorado) | Xenoberotha angustialata (184083) | |
Priabonian | Russian Federation (Kaliningrad) | Berothidae indet. larva A, Berothidae indet. larva B (140480) Berothidae indet. larva D (152019) Elektroberotha groehni (110735 147612) | |
Priabonian | Russian Federation | Berothidae indet. larva C (123419) |