PBDB Taxon

Basic info	Taxonomic history	Classification	Included Taxa
Morphology	Ecology and taphonomy	External Literature Search	Age range and collections

Hydroptilidae (microcaddisfly)

Insecta - Trichoptera - Hydroptilidae

Taxonomy

Hydroptilidae was named by Stephens (1836). It is extant. Its type is Hydroptila.

It was assigned to Trichoptera by Carpenter (1992); to Spicipalpia by Wichard et al. (2009) and Wichard (2013); to Integripalpia by Thomas et al. (2020); and to Hydroptiloidea by Thomas et al. (2023).

Subtaxa

Burminoptilinae, Electrotrichia, Hydroptilinae, Orthotrichiinae

Synonymy list

Year	Name and author
1836	Hydroptilidae Stephens p. 148
1992	Hydroptilidae Carpenter
2009	Hydroptilidae Wichard et al. p. 294
2013	Hydroptilidae Wichard
2020	Hydroptilidae Thomas et al.
2023	Hydroptilidae Thomas et al.

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Rank	Name	Author
kingdom	Animalia	()
—	Bilateria
—	Eubilateria	Ax 1987
—	Protostomia
—	Ecdysozoa
—	Panarthropoda
phylum	Arthropoda	Latreille 1829
superclass	Hexapoda	Latreille 1825
class	Insecta
—	Dicondylia
—	Paranotalia
subclass	Pterygota	()
—	Neopterygota	Crampton 1924
infraclass	Neoptera	Martynov 1923
—	Holometabola

Rank	Name	Author
—	Aparaglossata
—	Panorpida
superorder	Amphiesmenoptera	Kiriakoff 1948
—	Metamphiesmenoptera
—	Euamphiesmenoptera
—	Panendymenoptera
—	Endymenoptera
—	Stelloptera
order	Trichoptera	Kirby 1815
—	Eutrichoptera
suborder	Integripalpia
superfamily	Hydroptiloidea
family	Hydroptilidae
family	Hydroptilidae

If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.

Fm. Hydroptilidae Stephens 1836 [microcaddisfly]

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Subfm. †Burminoptilinae Wichard 2021

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G. †Burminoptila Botosaneanu 1981

†Burminoptila bemeneha Botosaneanu 1981

G. †Cretacoptila Wichard 2021

†Cretacoptila botosaneanui Wichard 2021

G. †Electrotrichia Ulmer 1912

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†Electrotrichia rasnitsyni Melnitsky et al. 2021

†Electrotrichia subtilis Ulmer 1912

Subfm. Hydroptilinae Stephens 1836

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Tr. Hydroptilini Stephens 1836

G. Agraylea Curtis 1834

Subg. †Agraylea (Nanoagraylea) Botosaneanu 1995 [micro caddis fly]

†Agraylea (Nanoagraylea) cretaria Botosaneanu 1995 [micro caddis fly]

†Agraylea (Nanoagraylea) parva Wichard and Bölling 2000

†Agraylea cumsacculo Wichard 2013

†Agraylea electroscientia Melnitsky and Ivanov 2010

†Agraylea glaesaria Wichard 2013

†Agraylea lentiginosa Botosaneanu et al. 1998

†Agraylea spathifera Ulmer 1912

Invalid names: Hydrorchestria Kolenati 1848 [synonym]

G. Allotrichia McLachlan 1880

†Allotrichia ampullata Ulmer 1912

†Allotrichia clara Wichard 2013

†Allotrichia succinica Hagen 1856

†Allotrichia superba Wichard 2013

G. Hydroptila Dalmon 1819

†Hydroptila phileos Cockerell 1921

G. Oxyethira Eaton 1873

†Oxyethira lurida Melnitsky and Ivanov 2016

Tr. Leucotrichiini Flint 1970

G. Alisotrichia Flint 1964

†Alisotrichia arizela Wells and Wichard 1989

G. Leucotrichia Mosely 1934

†Leucotrichia adela Wells and Wichard 1989

G. †Novajerseya Botosaneanu et al. 1998

†Novajerseya glesumica Botosaneanu et al. 1998

Tr. Ochrotrichiini Marshall 1979

G. Ochrotrichia Mosely 1934

Subg. Ochrotrichia (Ochrotrichia) Mosely 1934

Ochrotrichia (Ochrotrichia) aldama Mosely 1937

†Ochrotrichia (Ochrotrichia) brodzinskyi Wells and Wichard 1989

†Ochrotrichia (Ochrotrichia) chaulioda Wells and Wichard 1989

†Ochrotrichia (Ochrotrichia) denaia Wells and Wichard 1989

†Ochrotrichia (Ochrotrichia) doehleri Wichard 1981

†Ochrotrichia aliceae Wichard 2000

Invalid names: Polytrichia Sibley 1926 [replaced]

Subfm. Orthotrichiinae Nielsen 1948

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G. Orthotrichia Eaton 1873

†Orthotrichia umbra Melnitsky and Ivanov 2016

Diagnosis

No diagnoses are available

Measurements

No measurements are available

Composition:

chitinc

Environment:

terrestrialc

Locomotion:

actively mobilep

Created:

2017-04-17 09:56:18

Modified:

2017-04-17 09:56:18

Source: c = class, p = phylum

References: Kiessling 2004, Bush and Bambach 2015

Age range

Maximum range based only on fossils: base of the Turonian to the top of the Langhian or 93.90000 to 13.82000 Ma
Minimum age of oldest fossil (stem group age): 93.5 Ma

Collections (22 total)

Time interval	Ma	Country or state	Original ID and collection number
Early/Lower Cenomanian	99.6 - 93.5	Myanmar (Kachin)	Hydroptilidae indet. (104053)
Early/Lower Cenomanian	99.6 - 93.5	Myanmar	Burminoptila bemeneha (123020) Cretacoptila botosaneanui (161726)
Turonian	93.9 - 89.8	USA (New Jersey)	Agraylea sp., Agraylea cretaria, Agraylea parva (110895) Novajerseya glesumica, Agraylea lentiginosa (156002)
Ypresian	56.0 - 47.8	USA (Colorado)	Hydroptila phileos (105004)
Priabonian	38.0 - 33.9	Russian Federation	Electrotrichia subtilis (123930)
Priabonian	38.0 - 33.9	Russian Federation (Kaliningrad)	Agraylea spathifera (110735) Agraylea spathifera, Allotrichia ampullata, Electrotrichia subtilis (123927) Agraylea spathifera, Allotrichia succinica, Electrotrichia subtilis (124073) Electrotrichia subtilis (124951) Electrotrichia subtilis, Agraylea glaesaria, Agraylea cumsacculo, Allotrichia clara (124468) Electrotrichia subtilis, Agraylea glaesaria, Agraylea cumsacculo, Allotrichia clara, Allotrichia superba (123757) Electrotrichia subtilis, Allotrichia clara (120814)
Priabonian	38.0 - 33.9	Ukraine	Agraylea electroscientia (122996) Agraylea electroscientia, Oxyethira lurida, Orthotrichia umbra (122997) Electrotrichia rasnitsyni (185682)
Priabonian	38.0 - 33.9	Poland	Electrotrichia subtilis, Hydrorchestria succinica (120811) Hydrorchestria succinica (109625)
Burdigalian - Langhian	20.44 - 13.82	Dominican Republic	Ochrotrichia aliceae, Alisotrichia arizela, Ochrotrichia sp., Ochrotrichia brodzinskyi, Ochrotrichia aldama (124255) Ochrotrichia aliceae, Leucotrichia adela, Ochrotrichia chaulioda, Ochrotrichia denaia (122473) Ochrotrichia doehleri (122474)