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Carnivoramorpha

Mammalia

Taxonomy
Carnivoramorpha was named by Wyss and Flynn (1993). It is extant.

It was synonymized subjectively with Carnivora by Polly et al. (2006).

It was assigned to Laurasiatheria by Wesley-Hunt and Werdelin (2005); and to Ferae by Spaulding et al. (2009).

Synonymy list
YearName and author
1993Carnivoramorpha Wyss and Flynn
2005Carnivoramorpha Wesley-Hunt and Werdelin p. 838
2009Carnivoramorpha Spaulding et al. p. 5 figs. Figure 2

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RankNameAuthor
kingdomAnimalia()
—Bilateria
—EubilateriaAx 1987
—Deuterostomia
phylumChordataHaeckel 1874
subphylumVertebrata
superclassGnathostomata
—Osteichthyes()
subclassSarcopterygii()
subclassDipnotetrapodomorpha(Nelson 2006)
subclassTetrapodomorpha()
—Tetrapoda
—Reptiliomorpha
—Anthracosauria
subclassAmphibiosauriaKuhn 1967
—Cotylosauria()
—Amniota
subclassSynapsida
suborderEupelycosauria
—Therapsida()
RankNameAuthor
infraorderCynodontia()
—Mammaliamorpha
—Mammaliaformes
classMammalia
—Theriamorpha(Rowe 1993)
—Theriiformes()
—Trechnotheria
—Cladotheria
—Zatheria
subclassTribosphenida()
subclassTheria
—Eutheria()
—Placentalia
—Boreoeutheria
—Laurasiatheria
—Scrotifera
—Ferae()
—CarnivoramorphaWyss and Flynn 1993
—CarnivoramorphaWyss and Flynn 1993

If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.

Unr. Carnivoramorpha Wyss and Flynn 1993
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Unr. †Carnivoraformes Flynn et al.
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G. †Africtis Mattingly et al. 2020
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†Africtis sirtensis Mattingly et al. 2020
Or. Carnivora Bowditch 1821 [carnivoran]
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G. †Adracon Filhol 1884
Subor. Caniformia Kretzoi 1943
+
Fm. †Amphicyonidae Haeckel 1886 [bear-dog]
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G. †Amphicyanis Springhorn 1977
Subfm. †Amphicyoninae Trouessart 1885
G. †Angelarctocyon Tomiya and Tseng 2016
G. †Arctamphicyon Pilgrim 1932
G. †Bonisicyon Werdelin 2009
G. †Brachycyon Filhol 1872
G. †Cynodictis Bravard and Pomel 1850
Subfm. †Daphoeninae Hunt, Jr. 1998
G. †Guangxicyon Zhai et al. 2003
G. †Gustafsonia Tomiya and Tseng 2016
Subfm. †Haplocyoninae
G. †Harpagophagus De Bonis 1971
G. †Notoamphicyon Ameghino 1904
G. †Protemnocyon Hatcher 1901
G. †Pseudamphicyon Schlosser 1887
G. †Pseudocyonopsis Kuss 1965
G. †Storchictis de Bonis 2020
G. †Symplectocyon Springhorn 1979
Subfm. †Temnocyoninae Hunt, Jr. 1998
Subfm. †Thaumastocyoninae Hurzeller 1940
Invalid names: Aphicyoninae Trouessart 1885 [empty]
Infraor. Canoidea Simpson 1931
+
Superfm. Arctoidea Flower 1869
Fm. Canidae Fischer 1817 [canid]
G. †Lycophocyon Tomiya 2011
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†Lycophocyon hutchisoni Tomiya 2011
Invalid names: Cynoidea Flower 1869 [empty]
G. †Cryptailurus Martin and Lim 2001
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†Cryptailurus tinaynakti Barrett et al. 2020
Subor. Feliformia Kretzoi 1945
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Infraor. Aeluroida Flynn and Galiano 1982
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G. †Ictidopappus Simpson 1935
Invalid names: Viverravoidea Wortman and Matthew 1899 [empty]
Unr. Aeluroidea Flower 1869
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G. †Anictis Kretzoi 1945
G. †Haplogale
G. †Herpestides de Beaumont 1967
G. †Palaeoprionodon Filhol 1880
G. †Shandgolictis Hunt, Jr. 1998
G. †Stenoplesictis Filhol 1880
Infraor. Viverroidea Gray 1821
Invalid names: Stenoplesictinae Schlosser 1923 [invalid subgroup]
G. †Alagtsavbaatar Egi et al. 2016
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†Alagtsavbaatar indigenus Dashzeveg 1996
G. †Asiavorator Spassov and Lange-Badré 1995
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†Asiavorator gracilis Matthew and Granger 1924
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Invalid names: Asiavorator altidens Spassov and Lange-Badré 1995 [synonym], Stenoplesictis simplex Dashzeveg 1996 [synonym]
Superfm. Feloidea Hay 1930
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Fm. Felidae Gray 1821 [cat]
Fm. Prionodontidae Horsfield 1824 [asiatic linsang]
G. †Stenogale Schlosser 1888
G. †Viretictis de Bonis et al. 1999
Fm. Nandiniidae Pocock 1929
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G. Nandinia Gray 1843 [African palm civet]
Fm. †Nimravidae Cope 1880 [false sabre-tooth]
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G. †Aelurogale Filhol 1872
G. †Archaelurus Cope 1879
Subfm. †Barbourofelinae Schultz et al. 1970
G. †Maofelis Averianov et al. 2016
Subfm. †Nimravinae Trouessart 1885
+
Invalid names: Hoplophoneinae [synonym]
Tr. †Nimravini Cope 1880
G. †Pangurban Poust et al. 2022
Invalid names: Eusmilus insignis Filhol 1876 [nomen nudum]
Fm. †Palaeogalidae Martin and Lim 2001
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G. †Palaeogale von Meyer 1846
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Invalid names: Bunaelurus Cope 1873 [synonym]
Invalid names: Didymictida Flynn and Galiano 1982 [empty]
Subor. Fissipedia Blumenbach 1791
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Invalid names: Arctictidae Cope 1889 [empty], Bassarididae Cope 1889 [empty], Cynictidae Cope 1882 [empty]
Superfm. †Miacoidea Cope
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Fm. †Miacidae Cope 1880
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G. †Eogale Beard and Dawson 2009
Subfm. †Miacinae Cope 1880
G. †Miocyon Matthew 1909
G. †Pappidictops Chiu and Li 1977
G. †Simamphicyon Viret 1942
Subfm. †Viverravinae Matthew 1909
G. †Monachiopsis Kretzoi 1941
Fm. †Percrocutidae Werdelin and Solounias 1991
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G. †Dinocrocuta Schmidt-Kittler 1976
G. †Percrocuta Kretzoi 1938
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†Percrocuta australis Hendey 1974
†Percrocuta hebeiensis Chen and Wu 1976
†Percrocuta primordialis Qiu and Cao 1988
G. †Pseudailurus Gervais 1848
G. †Pristinictis Fox and Youzwyshyn 1994
+
†Pristinictis connata Fox and Youzwyshyn 1994
G. †Ravenictis Fox and Youzwyshyn 1994
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†Ravenictis krausei Fox and Youzwyshyn 1994
G. †Thos Bogachev
Invalid names: Eosictis Scott 1945 [nomen dubium], Felidi Jaekel 1911 [empty], Phocidi Jaekel 1911 [empty], Protelidi Jaekel 1911 [empty], Protoferi Jaekel 1911 [empty], Ursina Gray 1825 [empty], Ursocanidi Jaekel 1911 [empty], Viverra memphitis Linnaeus 1758 [nomen vanum]
G. †Dormaalocyon Solé et al. 2014
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†Dormaalocyon latouri Quinet 1966
G. †Gracilocyon Smith and Smith 2010
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†Gracilocyon igniculus Beard and Dawson 2009
†Gracilocyon rosei Heinrich et al. 2008
†Gracilocyon rundlei Hooker 2010
†Gracilocyon solei Smith and Smith 2010
†Gracilocyon winkleri Gingerich 1983
G. Lycarion Matthew 1909
+
†Lycarion medius Matthew 1909
G. †Miacis Cope 1872
+
†Miacis bathygnathus Scott 1888
†Miacis boqinghensis Huang et al. 1999
†Miacis deutschi Gingerich 1983
†Miacis edax Leidy 1872
†Miacis exiguus Matthew and Granger 1915
†Miacis gracilis Clark 1939
†Miacis hookwayi Stock 1934
†Miacis invictus Matthew and Granger 1925
†Miacis latidens Matthew and Granger 1915
†Miacis lushiensis Chow 1975
†Miacis parvivorus Cope 1872
†Miacis petilus Gingerich 1983
†Miacis rundlei Hooker 2010
†Miacis vorax Leidy 1872
†Miacis vulpinus Scott and Osborn 1887
Invalid names: Harpalodon Marsh 1872 [synonym], Prodaphaenus Wortman and Matthew 1899 [synonym]
G. †Neovulpavus Wortman 1901
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†Neovulpavus mccarrolli Tomiya et al. 2021
†Neovulpavus washakius Wortman 1901
G. †Palaearctonyx Matthew 1909
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†Palaearctonyx meadi Matthew 1909
G. †Paramiacis Filhol 1876
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†Paramiacis exilis Filhol 1876
†Paramiacis teilhardi Mathis 1987
G. †Procynodictis Wortman and Matthew 1899
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†Procynodictis vulpiceps Wortman and Matthew 1899
Invalid names: Plesiomiacis Simpson 1945 [synonym]
G. †Quercygale Kretzoi 1945
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†Quercygale angustidens Filhol 1872
†Quercygale hastingsae Davies 1884
†Quercygale helvetica Rutimeyer 1882
†Quercygale smithi Solé 2014
Invalid names: Humbertia De Beaumont 1965 [objective synonym]
G. †Uintacyon Leidy 1873
+
†Uintacyon acutus Thorpe 1923
†Uintacyon asodes Gazin 1952
†Uintacyon gingerichi Heinrich et al. 2008
†Uintacyon hookeri Solé 2014
†Uintacyon jugulans Matthew 1909
†Uintacyon rudis Rose 1981
G. †Vassacyon Matthew 1909
+
†Vassacyon bowni Heinrich et al. 2008
†Vassacyon prieuri Solé et al. 2016
†Vassacyon promicrodon Wortman and Matthew 1899
†Vassacyon taxidiotis Solé et al. 2013
G. †Vulpavus Marsh 1871
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Subg. †Vulpavus (Phlaodectes) Matthew 1909
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†Vulpavus australis Matthew and Granger 1915
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Invalid names: Vulpavus asius Gazin 1952 [synonym]
†Vulpavus canavus Cope 1881
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Invalid names: Miacis brevirostris Cope 1881 [synonym]
†Vulpavus completus Matthew 1909
†Vulpavus farsonensis Gunnell 1998
†Vulpavus hargeri Wortman 1901
†Vulpavus ovatus Matthew 1909
†Vulpavus palustris Marsh 1871
†Vulpavus profectus Matthew 1909
G. †Xinyuictis Zheng et al. 1975
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†Xinyuictis tenuis Zheng et al. 1975
G. †Zodiocyon Tong and Wang 2006
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†Zodiocyon zetesios Tong and Wang 2006
G. †Dawsonicyon Spaulding et al. 2010
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†Dawsonicyon isami Spaulding et al. 2010
G. †Oodectes Wortman 1901
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†Oodectes herpestoides Wortman 1901
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Invalid names: Oodectes proximus Matthew 1909 [synonym], Paeneprolimnocyon amissadomus Guthrie 1967 [synonym], Triacodon nanus Marsh 1872 [synonym]
†Oodectes jepseni Guthrie 1967
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Invalid names: Paeneprolimnocyon iudei Guthrie 1967 [synonym]
†Oodectes pugnax Wortman and Matthew 1899
Invalid names: Paeneprolimnocyon Guthrie 1967 [synonym]
G. †Tapocyon Stock 1934
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†Tapocyon dawsonae Wesley and Flynn 2003
†Tapocyon robustus Peterson 1919
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Invalid names: Tapocyon occidentalis Stock 1934 [synonym]
Fm. †Viverravidae Wortman and Matthew 1899
+
G. †Didymictis Cope 1875
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†Didymictis altidens Cope 1880
†Didymictis dellensis Dorr 1952
†Didymictis leptomylus Cope 1880
†Didymictis protenus Cope 1874
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Invalid names: Didymictis curtidens Cope 1882 [synonym]
†Didymictis proteus Polly 1997
†Didymictis vancleveae Robinson 1966
G. †Orientictis Huang and Zheng 2005
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†Orientictis spanios Huang and Zheng 2005
G. †Preonictis Tong and Wang 2006
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†Preonictis youngi Tong and Wang 2006
G. †Protictis Matthew
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Subg. †Protictis (Bryanictis) MacIntyre 1966
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†Bryanictis microlestes Simpson 1935
†Bryanictis paulus Meehan and Wilson 2002
Subg. †Protictis (Protictis) Matthew 1937
Subg. †Protictis (Protictoides) Flynn and Galiano 1982
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†Protictoides aprophatos Flynn and Galiano 1982
†Protictis agastor Gingerich and Winkler 1985
†Protictis haydenianus Cope 1882
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Invalid names: Didymictis primus Cope 1884 [synonym], Prolimnocyon macfaddeni Rigby, Jr. 1980 [synonym]
†Protictis minor Meehan and Wilson 2002
†Protictis paralus Holtzman 1978
†Protictis simpsoni Meehan and Wilson 2002
G. †Simpsonictis MacIntyre 1962
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†Simpsonictis jaynanneae Rigby, Jr. 1980
†Simpsonictis pegus Gingerich and Winkler 1985
†Simpsonictis tenuis Simpson 1935
G. †Variviverra Tong and Wang 2006
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†Variviverra vegetatus Tong and Wang 2006
G. †Viverravus Marsh 1872
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†Viverravus acutus Matthew and Granger 1915
†Viverravus gracilis Marsh 1872
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Invalid names: Didymictis dawkinsianus Cope 1881 [synonym], Harpalodon vulpinus Marsh 1872 [synonym], Triacodon fallax Marsh 1871 [synonym], Ziphacodon rugatus Marsh 1872 [synonym]
†Viverravus lawsoni Hooker 2010
†Viverravus laytoni Gingerich and Winkler 1985
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Invalid names: Viverravus bowni Gingerich 1987 [synonym]
†Viverravus lutosus Gazin 1952
†Viverravus minutus Wortman 1901
†Viverravus nitidus Marsh 1872
†Viverravus politus Matthew and Granger 1915
†Viverravus rosei Polly 1997
†Viverravus schaffi Gingerich and Winkler 1985
†Viverravus sicarius Matthew 1909
Invalid names: Ziphacodon Marsh 1872 [synonym]
G. †Viverriscus Beard and Dawson 2009
+
†Viverriscus omnivorus Beard and Dawson 2009
Invalid names: Deltatherium durini Van Valen 1978 [nomen dubium], Didymictidae Flynn and Galiano 1982 [synonym]
Diagnosis
No diagnoses are available
Measurements
No measurements are available
Composition: phosphaticsubp
Maximum body mass: 300 kgs
Minimum body mass: 10 kgs
Environment: terrestrialsubc
Locomotion: actively mobilec
Life habit: scansorialsubc
Diet: insectivoresubc
Reproduction: viviparoussubc
Created: 2005-09-06 20:43:42
Modified: 2005-09-06 22:44:32
Source: s = species, subc = subclass, c = class, subp = subphylum
References: Martin 1990, Lillegraven 1979, Carroll 1988, Ji et al. 2002, Hendy et al. 2009

Age range

Maximum range based only on fossils: base of the Puercan to the top of the Holocene or 66.00000 to 0.00000 Ma
Minimum age of oldest fossil (stem group age): 63.3 Ma

Collections (5568 total)

Oldest occurrences

Time interval Ma Country or state Original ID and collection number
Puercan66.0 - 63.3Canada (Saskatchewan) Carnivora indet., Ravenictis krausei (14768)
Early/Lower Paleocene66.0 - 61.7China (Anhui) Pappictidops orientalis (132943 146556)
Early/Lower Paleocene66.0 - 61.7China (Guangdong) Pappictidops acies, Pappictidops obtusus (140273 176607)
Danian66.0 - 61.6USA (North Dakota) Miacidae indet., Viverravinae indet., Protictis paralus (14962)
Cenozoic66.0 - 0.0Mongolia (South Gobi Aimag) Carnivora indet. (137861)
To263.3 - 61.7Canada (Alberta) Simpsonictis jaynanneae, Protictis sp. (14894)
Torrejonian63.3 - 61.7USA (Utah) Protictis haydenianus (14830)
Torrejonian63.3 - 61.7USA (New Mexico) Bryanictis paulus, Didymictis sp. b, Didymictis sp. a (14816) Didymictis haydenianus (14819 14873) Protictis haydenianus (14812 14815 14824 14832 14835 14862) Protictis haydenianus, Protictis sp. (14865) Protictis haydenianus, Protictis vanvaleni (14833 14893) Protictis sp. (14868) Protictis vanvaleni (14825)
Torrejonian63.3 - 61.7USA (Texas) Bryanictis sp. (14829) Protictis sp. (14887)
Torrejonian63.3 - 61.7USA (Wyoming) Bryanictis vanvaleni, Simpsonictis jaynanneae, Protictis haydenianus, Prolimnocyon macfaddeni (14886) Protictis haydenianus, Protictis microlestes (14896) Protictis microlestes (14899 14900 14901 14904 14905) Protictis sp., Protictis microlestes (14897 14898) Simpsonictis tenuis, Bryanictis microlestes, Protictis haydenianus, Protictis sp. (14912)
Torrejonian63.3 - 61.7USA (Montana) Didymictis haydenianus (14919 14920 14923) Didymictis microlestes (14917) Didymictis microlestes, Didymictis haydenianus (14913) Ictidopappus mustelinus, Protictis haydenianus, Bryanictis vanvaleni, Protictis tenuis, Protictis microlestes (14906)
Torrejonian63.3 - 61.7Canada (Alberta) Bryanictis microlestes (143200) Protictis sp. (14822) Simpsonictis jaynanneae (14828)
Torrejonian - Tiffanian63.3 - 56.8USA (Wyoming) Intyrictis vanvaleni (14853)
Ti161.7 - 56.8Canada (Alberta) Pristinictis connata, Simpsonictis pegus, Simpsonictis tenuis (14939)
Ti361.7 - 56.8Canada (Alberta) Raphictis gausion, Protictis paralus (14928 14992)
Ti461.7 - 56.8Canada (Saskatchewan) Viverravidae indet., Raphictis sp., Protictis paralus, Didymictis dellensis, Raphictis gausion, Protictis dellensis (14978)
Tiffanian61.7 - 56.8USA (California) Protictis agastor, Protictis paralus (154724)
Tiffanian61.7 - 56.8Canada (Alberta) Didymictis sp. (14995) Protictis dellensis (14996) Protictis sp. (15005) Raphictis sp., Raphictis gausion (14954)
Tiffanian61.7 - 56.8USA (North Dakota) Miacidae indet., Protictis paralus (14931) Protictis microlestes (15000)
Tiffanian61.7 - 56.8USA (Wyoming) Didymictis dellensis (14944 190436 190438 190462 190481) Didymictis sp. (14999) Didymictis sp. A (15034) Didymictis tenuis (14989) Protictis haydenianus (14974) Protictis microlestes (15014) Protictis paralus (15027) Protictis paralus, Protictis sp., Protictis haydenianus (15017) Protictis schaffi (15057 15065 15068) Protictis sp., Protictis microlestes (15067) Raphictis gausion, Protictis agastor, Protictis paralus (14936) Viverravinae indet., Protictis schaffi, Protictis laytoni, Protictis dellensis (15049) Viverravinae indet., Protictis sp. (15032) Viverravus laytoni (85096) Viverravus schaffi (26569 53059 190441)