PBDB Taxon

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Ceratopogonidae (biting midge)

Insecta - Diptera - Ceratopogonidae

Taxonomy

Ceratopogonidae was named by Newman (1834). It is extant. Its type is Ceratopogon.

It was assigned to Diptera by Grogan and Szadziewski (1988); to Nematocera by Carpenter (1992); and to Chironomoidea by Yeates et al. (2007).

Subtaxa

Adelohelea, Alautunmyia, Archiculicoides, Atriculicoidinae, Bezziinae, Ceratopogon crassipalpis, Ceratopogon madagascariensis, Ceratopogon tenuipes, Ceratopogon viscatus, Ceratopogoninae, Forcipomyiinae, Gerontodacus, Heleageron, Lebanoculicoidinae, Leptoconopinae (syn. Austroconopinae), Protoculicoides, Simulia pasithea, Simulium terribilis, Sinopogonitinae

Synonyms

Heleidae is a family. It is extant.

It was synonymized subjectively with Ceratopogonidae by Clapham (2023).
Simulium terribilis was named by Förster (1891). It is a compression fossil. Its type locality is Brunstatt, horizon d3, which is in an Oligocene lacustrine - large marl in France.

It was considered a nomen dubium by Crosskey and Howard (2004).
Sinopogonites was named by Hong (2002).

It was considered a nomen nudum by Hong (2002) and Hong (2002).

It was assigned to Ceratopogonidae by Borkent (2013).
Sinopogonites eocenicus was named by Hong (2002). It is an inclusion in amber. Its type locality is Xilutian coal mine, Fushun amber (post-1980 collection), which is in a Ypresian terrestrial amber in China.

It was considered a nomen nudum by Hong (2002) and Hong (2002).
Ceratopogon viscatus was named by Meunier (1912). It is an inclusion in amber. Its type locality is Madagascar copal (Meunier collection), which is in a Holocene terrestrial amber in Madagascar.

It was considered a nomen dubium by Borkent and Dominiak (2020).
Ceratopogon madagascariensis was named by Meunier (1912). It is an inclusion in amber. Its type locality is Madagascar copal (Meunier collection), which is in a Holocene terrestrial amber in Madagascar.

It was considered a nomen dubium by Borkent and Dominiak (2020).
Ceratopogon crassipalpis was named by Meunier (1912). It is an inclusion in amber. Its type locality is Madagascar copal (Meunier collection), which is in a Holocene terrestrial amber in Madagascar.

It was considered a nomen dubium by Borkent and Dominiak (2020).
Ceratopogon tenuipes was named by Meunier (1912). It is an inclusion in amber. Its type locality is Madagascar copal (Meunier collection), which is in a Holocene terrestrial amber in Madagascar.

It was considered a nomen dubium by Borkent and Dominiak (2020).
Simulia pasithea was named by von Heyden (1870). It is a compression fossil. Its type locality is Rott (Krantz Bonn coll), which is in a MP 30 lacustrine - large shale in the Rott Formation of Germany.

It was considered a nomen dubium by Crosskey and Howard (2004).
Sinopogonitinae was named by Hong (2002).

It was considered an invalid subgroup of Ceratopogonidae by Hong (2002).

Synonymy list

Year	Name and author
1834	Ceratopogonidae Newman p. 388
1870	Simulia pasithea von Heyden p. 238 figs. Pl 44, fig 1
1891	Simulium terribilis Förster p. 467 figs. Pl 14, fig 11
1912	Ceratopogon crassipalpis Meunier p. 191
1912	Ceratopogon madagascariensis Meunier p. 191
1912	Ceratopogon viscatus Meunier p. 191
1912	Ceratopogon crassipalpis Meunier p. 365 fig. 11
1912	Ceratopogon madagascariensis Meunier p. 365 fig. 9
1912	Ceratopogon tenuipes Meunier p. 366 fig. 13
1988	Ceratopogonidae Grogan and Szadziewski
1992	Ceratopogonidae Carpenter
2002	Sinopogonitinae Hong p. 109
2002	Sinopogonites Hong p. 167
2002	Sinopogonites eocenicus Hong p. 168
2007	Ceratopogonidae Yeates et al.
2013	Sinopogonites Borkent p. 172
2013	Sinopogonites eocenicus Borkent p. 172

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Rank	Name	Author
kingdom	Animalia	()
—	Bilateria
—	Eubilateria	Ax 1987
—	Protostomia
—	Ecdysozoa
—	Panarthropoda
phylum	Arthropoda	Latreille 1829

Rank	Name	Author
superclass	Hexapoda	Latreille 1825
class	Insecta
order	Diptera
infraorder	Culicomorpha
superfamily	Chironomoidea	Macquart 1838
family	Ceratopogonidae
family	Ceratopogonidae

If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.

Fm. Ceratopogonidae Newman 1834 [biting midge]

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G. †Adelohelea Borkent 1995

†Adelohelea burmitica Szadziewski and Poinar 2005

†Adelohelea glabra Borkent 1995

†Adelohelea magyarica Borkent 1997

G. †Alautunmyia Borkent 1996

†Alautunmyia elongata Borkent 1996

G. †Archiculicoides Szadziewski 1996

†Archiculicoides acraorum Borkent 2000

†Archiculicoides schleei Szadziewski 1996

†Archiculicoides unus Borkent 2000

Subfm. †Atriculicoidinae Szadziewski 1996

G. †Atriculicoides Remm 1976

†Atriculicoides cenomanensis Szadziewski and Schlüter 1992

†Atriculicoides ciliatus Borkent 2012

†Atriculicoides dasyheleis Szadziewski 1996

†Atriculicoides globosus Boesel 1937

Invalid names: Lasiohelea cretea Boesel 1937 [synonym]

†Atriculicoides hispanicus Szadziewski and Arillo 2016

†Atriculicoides incompletus Szadziewski and Schlüter 1992

†Atriculicoides macrophthalmus Remm 1976

Invalid names: Atriculicoides squamiciliatus Remm 1976 [synonym]

†Atriculicoides sanjusti Szadziewski and Arillo 2016

†Atriculicoides sibiricus Szadziewski 1996

†Atriculicoides swinhoei Cockerell 1919

†Atriculicoides szadziewskii Pérez-de la Fuente et al. 2011

†Atriculicoides taimyricus Szadziewski 1996

G. †Burmahelea Szadziewski and Sontag 2019

†Burmahelea neli Szadziewski and Sontag 2019

Subfm. Ceratopogoninae Newman 1834

Tr. Ceratopogonini Newman 1834

G. Alluaudomyia Kieffer 1913

†Alluaudomyia succinea Szadziewski 1988

G. Atyphohelea Borkent 1998

†Atyphohelea cothurnata Meunier 1904

Invalid names: Ceratopogon flagellus Meunier 1904 [synonym]

G. Austrohelea Wirth and Grogan 1988

G. Baeodasymyia Clastrier and Raccurt 1979

†Baeodasymyia dominicana Szadziewski and Grogan 1994

G. Baeohelea Wirth and Blanton 1970

G. †Brachycretacea Szadziewski 1996

†Brachycretacea taimyrica Remm 1976

G. Brachypogon Kieffer 1899

Subg. Brachypogon (Brachypogon) Kieffer 1899

†Brachypogon (Brachypogon) americanus Szadziewski and Grogan 1998

†Brachypogon (Brachypogon) balticus Szadziewski 1988

†Brachypogon (Brachypogon) eocenicus Szadziewski 1988

†Brachypogon (Brachypogon) gedanicus Szadziewski 1988

†Brachypogon (Brachypogon) miocaenicus Szadziewski 1993

Subg. Brachypogon (Isohelea) Kieffer 1917

†Brachypogon (Isohelea) dominicanus Szadziewski and Grogan 1998

†Brachypogon (Isohelea) henningseni Szadziewski 1988

†Brachypogon (Isohelea) polonicus Szadziewski 1988

†Brachypogon (Isohelea) prominuloides Szadziewski and Grogan 1998

†Brachypogon (Isohelea) prominulus Meunier 1904

†Brachypogon balticus Szadziewski 1988

G. Camptopterohelea Wirth and Hubert 1960

†Camptopterohelea odora Stebner et al. 2017

G. Ceratoculicoides Wirth and Ratanaworabhan 1971

†Ceratoculicoides danicus Szadziewski 1988

G. †Ceratopalpomyia Szadziewski 1988

†Ceratopalpomyia eocenica Szadziewski 1988

G. Ceratopogon Meigen 1803

†Ceratopogon bitterfeldi Szadziewski 1993

†Ceratopogon ceranowiczi Szadziewski 1988

†Ceratopogon crypticus Szadziewski 1988

†Ceratopogon eminens Meunier 1904

†Ceratopogon forcipiformis Meunier 1904

Invalid names: Ceratopogon defectus Meunier 1904 [synonym]

†Ceratopogon gedanicus Szadziewski 1988

†Ceratopogon grogani Szadziewski 1988

†Ceratopogon hennigi Szadziewski 1988

†Ceratopogon kotejai Szadziewski 1993

†Ceratopogon margaritae Szadziewski 1988

†Ceratopogon miocaenicus Szadziewski 1993

†Ceratopogon nanalobus Borkent and Grogan 1995

†Ceratopogon paraeminens Borkent and Grogan 1995

†Ceratopogon piotrowskii Szadziewski 1988

†Ceratopogon pisinnus Borkent and Grogan 1995

†Ceratopogon remmicola Szadziewski 1988

†Ceratopogon remmicolus Szadziewski 1988

†Ceratopogon ritzkowskii Szadziewski 1988

†Ceratopogon spiniger Loew 1850

†Ceratopogon subeminens Szadziewski 1993

†Ceratopogon succinicola Szadziewski 1993

†Ceratopogon tertiaricus Szadziewski 1988

Invalid names: Ceratopogon (Ceratopogon) Meigen 1803 [invalid subgroup]

G. †Eohelea Petrunkevitch 1957

†Eohelea fossicola Szadziewski 1993

†Eohelea gedanica Szadziewski 1988

†Eohelea grogani Szadziewski 1988

†Eohelea indica Stebner and Szadziewski 2017

†Eohelea miocaenea Szadziewski 1993

†Eohelea petrunkevitchi Szadziewski 1985

†Eohelea sakhalinica Szadziewski 1990

†Eohelea sinuosa Meunier 1904

Invalid names: Eohelea stridulans Petrunkevitch 1957 [synonym]

G. Fanthamia de Meillon 1939

G. †Fossihelea Szadziewski 1988

†Fossihelea gracilitarsis Meunier 1904

†Fossihelea miocaenica Szadziewski 1993

G. †Gedanohelea Szadziewski 1988

†Gedanohelea fushunensis Stebner et al. 2016

†Gedanohelea gerdesorum Stebner and Szadziewski 2017

†Gedanohelea liaoningensis Stebner et al. 2016

†Gedanohelea loewi Szadziewski 1988

†Gedanohelea succinea Szadziewski 1988

†Gedanohelea wirthi Szadziewski 1988

G. †Mantohelea Szadziewski 1988

†Mantohelea gedanica Szadziewski 1988

†Mantohelea laca Meunier 1904

†Mantohelea sinica Stebner et al. 2016

G. Meunierohelea Szadziewski 1988

†Meunierohelea anglesensis Peñalver et al. 2021

†Meunierohelea borkenti Stebner and Szadziewski 2017

†Meunierohelea cambayana Stebner and Szadziewski 2017

†Meunierohelea fudalai Szadziewski et al. 2020

†Meunierohelea gedanicola Szadziewski 1988

†Meunierohelea miocaenica Szadziewski 1993

†Meunierohelea nielseni Szadziewski 1988

†Meunierohelea orientalis Stebner and Szadziewski 2017

†Meunierohelea wirthi Szadziewski 1988

Invalid names: Chimaerohelea Debenham 1988 [synonym]

G. †Monogedania Szadziewski et al. 2022

†Monogedania clunipes Loew 1850

G. Monohelea Kieffer 1917

G. Nannohelea Grogan and Wirth 1980

†Nannohelea eocenica Szadziewski 1988

†Nannohelea grogani Szadziewski 1988

G. †Nelohelea Szadziewski and Sontag 2019

†Nelohelea neli Szadziewski and Sontag 2019

G. †Palaeobrachypogon Borkent 1995

†Palaeobrachypogon aquilonius Boesel 1937

†Palaeobrachypogon grandiforceps Borkent 2000

†Palaeobrachypogon macronyx Remm 1976

†Palaeobrachypogon remmi Borkent 1995

†Palaeobrachypogon taimyricus Szadziewski 1996

†Palaeobrachypogon vetus Borkent 1995

G. †Peronehelea Borkent 1995

†Peronehelea chrimikalydia Borkent 1995

†Peronehelea frigida Remm 1976

†Peronehelea sibirica Szadziewski 1996

G. Schizohelea Kieffer 1917

†Schizohelea baltica Szadziewski 1988

G. Serromyia Meigen 1818

†Serromyia alphea von Heyden 1870

Invalid names: Serromyia austera Statz 1944 [synonym], Serromyia colorata Statz 1944 [synonym], Serromyia spinofemorata Statz 1944 [synonym]

†Serromyia anomalicornis Loew 1850

†Serromyia errata Szadziewski 2018

†Serromyia polonica Szadziewski 1988

†Serromyia ryszardi Borkent 1990

†Serromyia sinuosa Borkent 1990

†Serromyia spinigera Loew 1850

Invalid names: Ceratopogon elongatus Meunier 1904 [synonym]

†Serromyia succinea Szadziewski 1988

G. Stilobezzia Kieffer 1911

Subg. Stilobezzia (Acanthohelea) Kieffer 1917

†Stilobezzia (Acanthohelea) dominicana Szadziewski and Grogan 1998

†Stilobezzia (Acanthohelea) falcata Meunier 1904

Invalid names: Ceratopogon spinosus Meunier 1904 [synonym]

†Stilobezzia (Acanthohelea) kutscheri Szadziewski 1993

†Stilobezzia (Acanthohelea) saxonica Szadziewski 1993

†Stilobezzia (Acanthohelea) veterana Meunier 1920

Invalid names: Stilobezzia goetghebueri Statz 1944 [synonym]

†Stilobezzia (Acanthohelea) wirthicola Szadziewski and Grogan 1998

Invalid names: Stilobezzia (Acanthohelea) succinea Szadziewski 1993 [replaced]

Subg. Stilobezzia (Stilobezzia) Kieffer 1911

†Stilobezzia (Stilobezzia) antilleana Szadziewski and Grogan 1998

†Stilobezzia kurthi Borkent 2000

†Stilobezzia pikei Borkent 2012

†Stilobezzia roggeroi Choufani and Nel 2013

G. †Wirthohelea Szadziewski 1988

†Wirthohelea trifida Szadziewski 1988

Invalid names: Stilobezziini Wirth 1952 [synonym]

Tr. Culicoidini Kieffer 1911

G. Culicoides Latreille 1809

Subg. Culicoides (Groganomyia) Szadziewski and Dominiak 2019

†Culicoides (Groganomyia) bojarskii Szadziewski and Dominiak 2019

†Culicoides (Groganomyia) burmiticus Szadziewski and Dominiak 2019

†Culicoides (Groganomyia) ellenbergeri Szadziewski and Dominiak 2019

†Culicoides (Groganomyia) myanmaricus Szadziewski and Dominiak 2019

Subg. Culicoides (Oecacta) Poey 1853

†Culicoides (Oecacta) dasyheleiformis Szadziewski 1988

†Culicoides (Oecacta) succivarius Szadziewski 1988

†Culicoides abbreviatipennis Statz 1944

†Culicoides agamus Borkent 1995

†Culicoides ambericus Szadziewski and Grogan 1998

†Culicoides annosus Borkent 1995

†Culicoides antilleanus Szadziewski and Grogan 1998

†Culicoides atratus Statz 1944

†Culicoides austerus Statz 1944

†Culicoides balticus Szadziewski 1988

†Culicoides bicolor Statz 1944

†Culicoides bifidus Borkent 1996

†Culicoides brisaci Choufani and Nel 2013

†Culicoides brodzinskyi Szadziewski and Grogan 1998

†Culicoides bullus Borkent 1995

†Culicoides canadensis Boesel 1937

†Culicoides casei Grogan and Szadziewski 1988 [biting midge]

†Culicoides ceranowiczi Szadziewski 1988

†Culicoides dasyheleiformis Szadziewski 1988

†Culicoides doyeni Choufani et al. 2014

†Culicoides elongatulus Statz 1944

†Culicoides eoselficus Szadziewski 1988

†Culicoides filipalpis Remm 1976

†Culicoides gedanensis Szadziewski 1988

†Culicoides gracilior Statz 1944

†Culicoides grandibocus Borkent 1996

†Culicoides hispanicola Szadziewski and Grogan 1998

†Culicoides jucundus Statz 1944

†Culicoides kaluginae Remm 1976

†Culicoides liliputanus Statz 1944

†Culicoides mammalicola Szadziewski and Grogan 1998

†Culicoides megacanthus Palmer 1957

†Culicoides obesus Statz 1944

†Culicoides obscuratus Statz 1944

†Culicoides obuncus Borkent 1995

†Culicoides paleopestis Peñalver et al. 2021

†Culicoides prussicus Szadziewski 1988

†Culicoides speciosus Meunier 1904

†Culicoides sphenostylus Remm 1976

†Culicoides subgedanensis Szadziewski 1993

†Culicoides succineus Remm 1976

†Culicoides succivarius Szadziewski 1988

†Culicoides tenuipennis Statz 1944

†Culicoides truncatus Borkent 2000

†Culicoides tyrrelli Boesel 1937

†Culicoides ventralis Borkent and Wirth 1997

Invalid names: Culicoides abdominalis Statz 1944 [replaced]

†Culicoides yoosti Borkent 2000

Invalid names: Devalquia Choufani and Nel 2013 [synonym]

G. Washingtonhelea Wirth and Grogan 1988

Tr. Heteromyiini Wirth 1962

G. Heteromyia Say 1825

†Heteromyia dominicana Szadziewski and Grogan 1997

G. Metahelea Edwards 1929

†Metahelea serafini Szadziewski 1998

G. Neurohelea Kieffer 1925

G. Physohelea Grogan and Wirth 1979

†Physohelea obtusa Meunier 1904

Invalid names: Ceratopogon cothurnatulus Meunier 1904 [synonym]

Tr. Johannsenomyiini Crampton 1925

G. Johannsenomyia Malloch 1915

G. Macropeza Meigen 1818

G. Mallochohelea Wirth 1962

†Mallochohelea martae Szadziewski 2005

Tr. Palpomyiini Enderlein 1936

G. Bezzia Kieffer 1899

†Bezzia eocenica Szadziewski 1988

†Bezzia longipennis Statz 1944

G. Palpomyia Meigen 1818

†Palpomyia edwardsi Cockerell 1921

†Palpomyia erikae Szadziewski 1993

†Palpomyia freyi Pierce 1966

†Palpomyia jantari Szadziewski 1988

†Palpomyia multispinosa Pierce 1966

†Palpomyia riedeli Szadziewski 1988

†Palpomyia ryshkoffi Pierce 1966

†Palpomyia shilo Pierce 1966

†Palpomyia succinea Szadziewski 1988

†Palpomyia unca Hong 1981

†Palpomyia wirthorum Szadziewski and Grogan 1997

Invalid names: Eopalpomyitis Hong et al. 2000 [synonym], Miopalpomyia Pierce 1966 [synonym], Neopalpomyia Pierce 1966 [synonym], Parapalpomyia Pierce 1966 [synonym]

G. Phaenobezzia Haeselbarth 1965

†Phaenobezzia wirthi Szadziewski and Grogan 1997

Invalid names: Ceratopogon escheri Giebel 1856 [nomen dubium], Culicoides carri Pierce 1966 [nomen dubium], Culicoides fossilis Pierce 1966 [nomen dubium], Culicoides laurae Pierce 1966 [nomen dubium], Culicoides mioceneus Pierce 1966 [nomen dubium], Johannsenomyia hotchkissae Pierce 1966 [nomen dubium], Neoculicoides Pierce 1966 [nomen dubium], Neoculicoides jeanneae Pierce 1966 [nomen dubium], Sphaeromiini Newman 1834 [empty]

Subfm. Forcipomyiinae Lenz 1934

Tr. Dasyheleini

G. Dasyhelea Kieffer 1913

†Dasyhelea antilleana Szadziewski and Grogan 1998

†Dasyhelea antiqua Palmer 1957

†Dasyhelea browneae Pierce 1966

†Dasyhelea dara Pierce 1966

†Dasyhelea dominicana Szadziewski and Grogan 1998

†Dasyhelea eodicryptoscenica Szadziewski 1988

†Dasyhelea gedanica Szadziewski 1988

†Dasyhelea hispaniolae Szadziewski and Grogan 1998

†Dasyhelea judithae Pierce 1966

†Dasyhelea kanakoffi Pierce 1966

†Dasyhelea minuticola Szadziewski and Grogan 1998

†Dasyhelea miocaenica Szadziewski 1993

†Dasyhelea stanislavi Szadziewski 1988

†Dasyhelea stenoceras Palmer 1957

Invalid names: Sebessia Remm 1979 [synonym]

Tr. Forcipomyiini Lenz 1934

G. Atrichopogon Kieffer 1906

†Atrichopogon brunnescens Statz 1944

†Atrichopogon dominicanus Szadziewski and Grogan 1998

†Atrichopogon eocenicus Szadziewski 1988

Invalid names: Atrichopogon (Atrichopogon) Kieffer 1906 [invalid subgroup]

G. Forcipomyia Meigen 1818

Subg. Forcipomyia (Euprojoannisia) Brèthes 1914

†Forcipomyia (Euprojoannisia) berendti Szadziewski 1988

†Forcipomyia (Euprojoannisia) henningseni Szadziewski 1988

†Forcipomyia (Euprojoannisia) miocaenica Szadziewski 1993

†Forcipomyia (Euprojoannisia) piriformis Meunier 1904

†Forcipomyia (Euprojoannisia) tuberculosa Szadziewski 1993

Subg. Forcipomyia (Forcipomyia) Meigen 1818

†Forcipomyia (Forcipomyia) eobreviflagellata Szadziewski 1988

†Forcipomyia (Forcipomyia) eocostata Szadziewski 1988

†Forcipomyia (Forcipomyia) fusiparamera Szadziewski and Grogan 1998

†Forcipomyia (Forcipomyia) gedanicola Szadziewski 1988

†Forcipomyia (Forcipomyia) grimaldii Szadziewski and Grogan 1998

†Forcipomyia (Forcipomyia) krzeminskii Szadziewski 1988

†Forcipomyia (Forcipomyia) kulickae Szadziewski 1988

†Forcipomyia (Forcipomyia) lyneborgi Szadziewski 1988

†Forcipomyia (Forcipomyia) pseudomicrohelea Szadziewski 1988

†Forcipomyia (Forcipomyia) rasnitsyni Szadziewski et al. 2021

†Forcipomyia (Forcipomyia) subgedanicola Szadziewski 1993

†Forcipomyia (Forcipomyia) turbinata Meunier 1904

†Forcipomyia (Forcipomyia) uncula Meunier 1904

†Forcipomyia (Forcipomyia) unculiformis Szadziewski 1993

Subg. Forcipomyia (Lasiohelea) Kieffer 1921

†Forcipomyia (Lasiohelea) americana Szadziewski and Grogan 1998

†Forcipomyia (Lasiohelea) succinea Szadziewski 1988

†Forcipomyia (Lasiohelea) woodruffi Szadziewski and Grogan 1998

Subg. Forcipomyia (Lepidohelea) Kieffer 1917

†Forcipomyia (Lepidohelea) antilleana Szadziewski and Grogan 1998

†Forcipomyia (Lepidohelea) chrysosuccinea Szadziewski and Grogan 1998

†Forcipomyia (Lepidohelea) domibicolor Szadziewski and Grogan 1998

†Forcipomyia (Lepidohelea) lepidosuccinea Szadziewski and Grogan 1998

Subg. Forcipomyia (Phytohelea) Remm 1971

†Forcipomyia (Phytohelea) eophytoheleana Szadziewski 1988

Subg. Forcipomyia (Synthyridomyia) Saunders 1957

†Forcipomyia (Synthyridomyia) tertiaricola Szadziewski and Grogan 1998

Subg. Forcipomyia (Thyridomyia) Saunders 1925

†Forcipomyia (Thyridomyia) frutetosuccinea Szadziewski and Grogan 1998

Subg. Forcipomyia (Trichohelea) Goetghebuer 1920

†Forcipomyia (Trichohelea) bifidicola Szadziewski 1993

†Forcipomyia (Trichohelea) eotrichoheleana Szadziewski 1988

†Forcipomyia (Trichohelea) succinicola Szadziewski 1993

†Forcipomyia eobreviflagellata Szadziewski 1988

†Forcipomyia krzeminskii Szadziewski 1988

†Forcipomyia kulickae Szadziewski 1988

†Forcipomyia nadicola Szadziewski 2013

†Forcipomyia orientalus Hong 1981

Invalid names: Amberaspinus Evenhuis 1994 [synonym]

G. †Gerontodacus Borkent 2019

†Gerontodacus krzeminskii Choufani et al. 2015

†Gerontodacus punctus Borkent 2000

†Gerontodacus skalskii Szadziewski and Arillo 1998

†Gerontodacus succineus Szadziewski 1996

G. †Heleageron Borkent 1995

†Heleageron arenatus Borkent 1995

†Heleageron grimaldii Borkent 1996

Subfm. †Lebanoculicoidinae Borkent 2000

G. †Lebanoculicoides Szadziewski 1996

†Lebanoculicoides bloudani Choufani et al. 2015

†Lebanoculicoides daheri Choufani et al. 2015

†Lebanoculicoides excantabris Pérez-de la Fuente et al. 2011

†Lebanoculicoides mesozoicus Szadziewski 1996

Subfm. Leptoconopinae Noe 1907

G. †Archiaustroconops Szadziewski 1996

†Archiaustroconops alavensis Szadziewski and Arillo 1998

†Archiaustroconops andersoni Szadziewski et al. 2015

†Archiaustroconops annae Choufani et al. 2015

†Archiaustroconops besti Borkent et al. 2013

†Archiaustroconops bocaparvus Borkent 2000

†Archiaustroconops borkenti Pérez-de la Fuente et al. 2011

†Archiaustroconops ceratoformis Szadziewski 1996

†Archiaustroconops cretaceous Szadziewski 1996

†Archiaustroconops dominiakae Choufani et al. 2015

†Archiaustroconops gracilis Szadziewski and Poinar 2005

†Archiaustroconops hammanensis Choufani et al. 2015

†Archiaustroconops hamus Borkent 2000

†Archiaustroconops kotejai Szadziewski and Poinar 2005

†Archiaustroconops szadziewskii Borkent 2000

G. Austroconops Wirth and Lee 1958

†Austroconops asiaticus Szadziewski 2004

†Austroconops borkenti Szadziewski and Schlüter 1992

†Austroconops fossilis Szadziewski 1996

†Austroconops gladius Borkent 2000

†Austroconops gondwanicus Szadziewski 1996

†Austroconops megaspinus Borkent 2000

†Austroconops perrichoti Dominiak et al. 2018

†Austroconops sibiricus Szadziewski 1996

G. †Baskintoconops Pielowska-Ceranowska 2023

†Baskintoconops maaloufi Pielowska-Ceranowska 2023

G. †Fossileptoconops Szadziewski 1996

†Fossileptoconops lebanicus Szadziewski 1996

G. †Jordanoconops Szadziewski 2000

†Jordanoconops weitschati Szadziewski 2000

G. Leptoconops Skuse 1889

Subg. Leptoconops (Leptoconops) Skuse 1889

†Leptoconops (Leptoconops) boreus Kalugina 1991

†Leptoconops (Leptoconops) burmiticus Szadziewski 2004

†Leptoconops (Leptoconops) clava Borkent 1997

†Leptoconops (Leptoconops) copiosus Borkent 1996

†Leptoconops (Leptoconops) curvachelus Borkent 1996

†Leptoconops (Leptoconops) daugeroni Choufani et al. 2011

†Leptoconops (Leptoconops) ellenbergeri Szadziewski 2015

†Leptoconops (Leptoconops) gravesi Choufani et al. 2014

†Leptoconops (Leptoconops) myanmaricus Szadziewski 2004

†Leptoconops (Leptoconops) nosopheris Poinar 2008

†Leptoconops (Leptoconops) primaevus Borkent 1995

†Leptoconops (Leptoconops) rossi Szadziewski 2004

†Leptoconops (Leptoconops) rovnensis Sontag and Szadziewski 2011

†Leptoconops (Leptoconops) sibiricus Szadziewski 1996

†Leptoconops (Leptoconops) subrossicus Szadziewski and Poinar 2005

†Leptoconops (Leptoconops) succineus Szadziewski 1988

†Leptoconops (Leptoconops) zherikhini Szadziewski and Arillo 2003

Subg. †Leptoconops (Palaeoconops) Borkent 2001

†Leptoconops (Palaeoconops) amplificatus Borkent 2001

†Leptoconops (Palaeoconops) antiquus Borkent 2001

G. †Minyohelea Borkent 1995

†Minyohelea bacula Borkent 2000

†Minyohelea casca Borkent 1997

†Minyohelea falcata Borkent 2000

†Minyohelea lebanica Szadziewski 1996

†Minyohelea minuta Szadziewski 1996

†Minyohelea nexuosa Pielowska-Ceranowska 2021

†Minyohelea pumilis Borkent 1995

†Minyohelea schleei Szadziewski 1996

†Minyohelea wirthi Szadziewski 1996

Invalid names: Lebanoconops Szadziewski 1996 [synonym]

Invalid names: Austroconopinae Borkent et al. 1987 [synonym]

G. †Protoculicoides Boesel 1937

†Protoculicoides depressus Boesel 1937

†Protoculicoides revelatus Borkent 2019

Invalid names: Ceratopogon crassipalpis Meunier 1912 [nomen dubium], Ceratopogon madagascariensis Meunier 1912 [nomen dubium], Ceratopogon tenuipes Meunier 1912 [nomen dubium], Ceratopogon viscatus Meunier 1912 [nomen dubium], Heleidae [synonym], Simulia pasithea von Heyden 1870 [nomen dubium], Simulium terribilis Förster 1891 [nomen dubium], Sinopogonitinae Hong 2002 [invalid subgroup]

Diagnosis

No diagnoses are available

Measurements

No measurements are available

Composition:

chitinc

Environment:

terrestrialc

Locomotion:

actively mobilep

Created:

2017-04-17 09:56:18

Modified:

2017-04-17 09:56:18

Source: c = class, p = phylum

References: Bush and Bambach 2015, Kiessling 2004

Age range

Maximum range based only on fossils: base of the Late/Upper Berriasian to the top of the Holocene or 145.00000 to 0.00000 Ma
Minimum age of oldest fossil (stem group age): 140.2 Ma

Collections (164 total)

Time interval	Ma	Country or state	Original ID and collection number
Late/Upper Berriasian	145.0 - 140.2	United Kingdom (England)	Archiaustroconops besti (154198)
Hauterivian	132.9 - 129.4	Austria	Minyohelea casca (119479)
Early/Lower Barremian	130.0 - 125.45	Lebanon	Archiaustroconops ceratoformis (124925 154704) Baskintoconops maaloufi (229464) Lebanoculicoides bloudani (164939) Lebanoculicoides daheri, Protoculicoides krzeminskii, Archiaustroconops annae, Archiaustroconops hammanensis, Archiaustroconops dominiakae, Minyohelea nexuosa (122471) Leptoconops amplificatus, Leptoconops antiquus, Austroconops sp., Protoculicoides punctus, Archiaustroconops ceratoformis, Archiaustroconops szadziewskii, Archiaustroconops hamus, Austroconops gondwanicus, Austroconops fossilis, Minyohelea falcata (123017) Minyohelea nexuosa (229465) Minyohelea sp., Archiaustroconops ceratoformis, Archiaustroconops bocaparvus (124363) Minyohelea sp., Austroconops sp., Lebanoculicoides mesozoicus, Fossileptoconops lebanicus, Protoculicoides succineus, Archiculicoides schleei, Archiaustroconops ceratoformis, Archiaustroconops hamus, Austroconops cretaceous, Austroconops gondwanicus, Austroconops fossilis, Minyohelea schleei, Lebanoconops lebanicus, Lebanoconops minutus, Lebanoconops wirthi (124276) Protoculicoides punctus (124617) Protoculicoides punctus, Archiaustroconops ceratoformis, Archiaustroconops hamus, Archiaustroconops hammanensis (123416) Protoculicoides sp., Minyohelea sp., Austroconops sp., Archiaustroconops sp., Lebanoculicoides mesozoicus, Protoculicoides succineus, Protoculicoides unus, Protoculicoides acraorum, Archiaustroconops ceratoformis, Archiaustroconops szadziewskii, Archiaustroconops hamus, Archiaustroconops bocaparvus, Austroconops gladius, Austroconops fossilis, Austroconops megaspinus, Minyohelea schleei, Minyohelea bacula, Minyohelea lebanica, Minyohelea minuta (122359)
Early/Lower Barremian	130.0 - 125.45	Syria	Lebanoculicoides bloudani (164938)
Late/Upper Aptian	122.46 - 113.0	Australia (Victoria)	Leptoconops sp. (77192)
Albian	113.0 - 100.5	Jordan	Archiaustroconops sp., Jordanoconops weitschati (124848) Ceratopogonidae indet. (124847)
Early/Lower Albian	112.03 - 109.0	Spain	Austroconops sp., Lebanoculicoides excantabris, Protoculicoides skalskii, Archiaustroconops borkenti, Atriculicoides szadziewskii (109438)
Late/Upper Albian	105.3 - 99.6	Spain (Álava)	Leptoconops zherikhini, Protoculicoides skalskii, Archiaustroconops alavensis (108769)
Late/Upper Albian	105.3 - 99.6	France (Poitou-Charentes)	Leptoconops daugeroni, Austroconops perrichoti (99114)
Late/Upper Albian	105.3 - 99.6	Spain (Aragón)	Leptoconops zherikhini, Protoculicoides skalskii, Protoculicoides hispanicus, Protoculicoides sanjusti (91626)
Late/Upper Albian - Early/Lower Cenomanian	105.3 - 93.5	France (Charente-Maritime)	Ceratopogonidae indet. (107076)
Early/Lower Cenomanian	99.6 - 93.5	France (Poitou-Charentes)	Ceratopogonidae indet. (65422)
Early/Lower Cenomanian	99.6 - 93.5	Myanmar (Kachin)	Protoculicoides sp., Leptoconops sp., Leptoconops burmiticus, Atriculicoides swinhoei, Protoculicoides burmiticus, Archiaustroconops gracilis, Archiaustroconops kotejai, Leptoconops subrossicus, Leptoconops nosopheris (104053)
Early/Lower Cenomanian	99.6 - 93.5	Myanmar	Atriculicoides sp., Austroconops asiaticus, Leptoconops burmiticus, Leptoconops myanmaricus, Leptoconops rossi, Johannsenomyia swinhoei (123020) Austroconops sp., Leptoconops myanmaricus, Archiaustroconops sp., Archiculicoides andersoni (123054) Ceratopogonidae indet. (111688) Culicoides bojarskii, Culicoides myanmaricus (161726) Culicoides burmiticus, Burmahelea neli, Nelohelea neli (187929) Leptoconops ellenbergeri, Culicoides ellenbergeri (165560) Protoculicoides revelatus (123116)
Early/Lower Cenomanian	99.6 - 93.5	France	Ceratopogonidae indet. A, Ceratopogonidae indet. B (123018) Ceratopogonidae indet. D, Ceratopogonidae indet. C (123019)
Late/Upper Cenomanian	99.6 - 93.5	USA (New Jersey)	Culicoides casei (110959)
Late/Upper Cenomanian	99.6 - 93.5	Russian Federation	Atriculicoides sp., Leptoconops boreus (35051)
Turonian	93.9 - 89.8	France	Leptoconops sp., Leptoconops gravesi, Culicoides doyeni (164386)
Turonian	93.9 - 89.8	USA (New Jersey)	Atriculicoides globosus, Palaeobrachypogon sp., Leptoconops copiosus, Leptoconops curvachelus, Culicoides bifidus (123016) Culicoides casei, Atriculicoides incompletus, Atriculicoides globosus, Culicoides filipalpis, Palaeobrachypogon remmi, Leptoconops copiosus, Leptoconops curvachelus, Alautunmyia elongata, Heleageron grimaldii, Culicoides grandibocus, Culicoides truncatus, Culicoides yoosti, Palaeobrachypogon grandiforceps, Stilobezzia kurthi (110895)
Santonian	86.3 - 83.6	Hungary	Leptoconops clava, Adelohelea magyarica (119480)
Early/Lower Santonian	85.8 - 83.5	France	Devalquia brisaci, Metahelea roggeroi (152462)
Late/Upper Santonian	85.8 - 83.5	Russian Federation	Atriculicoides sp., Atriculicoides spp., Culicoides sp., Culicoides filipalpis, Ceratopogon macronyx, Ceratopogon frigidus, Atriculicoides macrophthalmus, Culicoides kaluginae, Culicoides sphenostylus, Culicoides succineus, Austroconops sibiricus, Leptoconops sibiricus, Atriculicoides dasyheleis, Atriculicoides sibiricus, Atriculicoides taimyricus, Washingtonhelea taimyrica, Baeohelea taimyrica, Peronehelea sibirica (35056) Atriculicoides sp., Leptoconops sp., Atriculicoides macrophthalmus, Atriculicoides squamiciliatus, Austroconops sibiricus (122590)
Late/Upper Campanian	83.5 - 70.6	Canada (Manitoba)	Atriculicoides sp., Atriculicoides globosus, Adelohelea glabra, Culicoides filipalpis, Culicoides canadensis, Culicoides obuncus, Peronehelea chrimikalydia (122353) Atriculicoides sp., Leptoconops primaevus, Atriculicoides globosus, Heleageron arenatus, Culicoides filipalpis, Culicoides tyrrelli, Culicoides canadensis, Culicoides agamus, Culicoides obuncus, Culicoides annosus, Palaeobrachypogon remmi, Palaeobrachypogon vetus, Peronehelea chrimikalydia (122354) Atriculicoides sp., Protoculicoides depressus, Lasiohelea globosa, Lasiohelea cretea, Heleageron arenatus, Culicoides filipalpis, Dasyhelea tyrrelli, Atrichopogon canadensis, Culicoides obuncus, Ceratopogon aquilonius, Palaeobrachypogon remmi, Palaeobrachypogon vetus (122356)
Late/Upper Campanian	83.5 - 70.6	Myanmar	Protoculicoides sp. (195455)
Middle Campanian	83.5 - 70.6	Canada (Alberta)	Atriculicoides sp., Culicoides sp., Adelohelea glabra, Peronehelea chrimikalydia (34896) Ceratopogonidae indet., Atriculicoides globosus, Heleageron arenatus, Culicoides filipalpis, Culicoides tyrrelli, Culicoides bullus, Palaeobrachypogon aquilonius, Peronehelea chrimikalydia (122388) Ceratopogonidae indet., Atriculicoides sp., Culicoides sp., Leptoconops primaevus, Protoculicoides depressus, Atriculicoides globosus, Heleageron arenatus, Culicoides filipalpis, Palaeobrachypogon sp., Palaeobrachypogon vetus, Peronehelea sp., Peronehelea chrimikalydia, Stilobezzia pikei (122389) Culicoides filipalpis (122385) Culicoides filipalpis, Culicoides bullus (123143) Leptoconops primaevus, Atriculicoides globosus, Adelohelea glabra, Heleageron arenatus, Culicoides filipalpis, Culicoides tyrrelli, Culicoides canadensis, Culicoides obuncus, Palaeobrachypogon remmi, Palaeobrachypogon vetus, Protoculicoides depressus (122384) Leptoconops primaevus, Atriculicoides globosus, Culicoides filipalpis, Culicoides tyrrelli, Culicoides annosus, Peronehelea chrimikalydia (104932) Leptoconops primaevus, Atriculicoides globosus, Culicoides filipalpis, Culicoides tyrrelli, Culicoides canadensis, Minyohelea pumilis, Palaeobrachypogon vetus, Peronehelea chrimikalydia (122383) Minyohelea pumilis, Peronehelea chrimikalydia (122358)
Ypresian	56.0 - 47.8	China (Liaoning)	Mantohelea sinica, Gedanohelea fushunensis, Gedanohelea liaoningensis (125934) Palpomyia unca, Aspinus orientalus (124530) Sinopogonites eocenicus (124820)
Ypresian	56.0 - 47.8	India (Gujarat)	Ceratopogonidae indet., Forcipomyia sp. (183743) Ceratopogonidae indet., Mantohelea sp., Leptoconops sp., Stilobezzia sp., Forcipomyia sp., Brachypogon sp., Serromyia sp., Camptopterohelea odora, Eohelea indica, Gedanohelea gerdesorum, Meunierohelea cambayana, Meunierohelea borkenti, Meunierohelea orientalis (124463) Forcipomyia sp., Meunierohelea sp., Gedanohelea gerdesorum (183742)
Lutetian	47.8 - 41.3	Russian Federation	Forcipomyia nadicola (151918) Leptoconops sp., Stilobezzia sp., Ceratopogoninae indet., Eohelea sakhalinica, Forcipomyia sp. indet., Forcipomyia sp., Forcipomyia sp. (107340)
Lutetian	47.8 - 41.3	USA (Montana)	Alluaudomyia sp. (224195)
Bartonian	41.3 - 38.0	Australia (Victoria)	Monohelea sp., Meunierohelea anglesensis, Culicoides paleopestis (224365)
Priabonian	38.0 - 33.9	Germany (Halle)	Ceratopogon sp., Culicoides sp., Dasyhelea sp., Forcipomyia sp., Culicoides speciosus, Ceratopogon bitterfeldi, Ceratopogon forcipiformis, Ceratopogon hennigi, Eohelea fossicola, Eohelea miocaenea, Eohelea sinuosa, Fossihelea sp., Fossihelea miocaenica, Monohelea clunipes, Serromyia sp., Serromyia spinigera, Forcipomyia sp., Stilobezzia succinea, Forcipomyia miocaenica, Forcipomyia tuberculosa, Forcipomyia gedanicola, Forcipomyia subgedanicola, Forcipomyia unculiformis, Forcipomyia succinicola, Culicoides ceranowiczi, Culicoides subgedanensis, Brachypogon miocaenicus, Ceratopogon miocaenicus, Ceratopogon subeminens, Ceratopogon succinicolus, Chimaerohelea miocaenica, Palpomyia erikae, Serromyia alphea (129227) Ceratopogon sp., Culicoides sp., Mantohelea gedanica, Forcipomyia sp., Forcipomyia sp., Culicoides speciosus, Ceratopogon forcipiformis, Ceratopogon hennigi, Eohelea sinuosa, Fossihelea sp., Fossihelea sp. B, Monohelea clunipes, Nannohelea sp., Serromyia sp., Forcipomyia sp., Brachypogon sp., Brachypogon sp., Forcipomyia tuberculosa, Forcipomyia bifidicola, Dasyhelea miocaenica, Brachypogon prominulus, Ceratopogon kotejai, Stilobezzia falcata, Stilobezzia kutscheri, Stilobezzia saxonica, Chimaerohelea nielseni, Serromyia alphea (129228)
Priabonian	38.0 - 33.9	Russian Federation (Kaliningrad)	Ceratopogon escheri (147344) Ceratopogon lacus, Ceratopogon speciosus, Ceratopogon prominulus, Ceratopogon eminens, Ceratopogon forcipiformis, Fossihelea sp. A, Ceratopogon gracilitarsis, Ceratopogon clunipes, Ceratopogon falcatus, Ceratopogon obtusus, Ceratopogon cothurnatus, Ceratopogon turbinatus, Ceratopogon unculus, Ceratopogon piriformis, Ceratopogon defectus, Ceratopogon flagellus, Ceratopogon elongatus, Ceratopogon cothurnatulus, Ceratopogon spiniger, Ceratopogon sinuosus, Ceratopogon spinosus (123927) Ceratopogon sp., Culicoides sp., Ceratopogoninae indet., Forcipomyia sp., Eohelea sinuosa, Monohelea clunipes, Forcipomyia uncula, Brachypogon sp., Forcipomyia pseudomicrohelea, Culicoides ceranowiczi, Alluaudomyia succinea, Culicoides dasyheleiformis (123221) Ceratopogon sp., Culicoides sp., Ceratopogoninae indet., Forcipomyia spp., Forcipomyia sp., Ceratopogonini indet., Ceratopogon tertiaricus, Monohelea sp. A, Monohelea clunipes, Forcipomyia turbinata, Forcipomyia uncula, Dasyhelea eodicryptoscenica, Brachypogon balticus, Brachypogon prominulus, Forcipomyia krzeminskii (123993) Ceratopogon sp., Culicoides sp., Stilobezzia sp. A, Stilobezzia sp. B, Ceratopogoninae indet., Forcipomyia spp., Forcipomyia sp., Ceratopogonini indet., Ceratopogon forcipiformis, Eohelea sinuosa, Fossihelea gracilitarsis, Monohelea clunipes, Nannohelea sp., Serromyia sp., Forcipomyia piriformis, Brachypogon prominulus (123962) Ceratopogonidae indet. (201682)
Priabonian	38.0 - 33.9	Poland	Ceratopogon sp., Culicoides sp., Dasyhelea sp., Mantohelea lacus, Mantohelea gedanica, Stilobezzia sp., Stilobezzia sp. C, Stilobezzia sp. D, Stilobezzia sp. E, Forcipomyia spp., Forcipomyia sp., Forcipomyia sp. A, Forcipomyia sp., Ceratopogonini indet., Ceratopogon eminens, Ceratopogon forcipiformis, Ceratopogon grogani, Ceratopogon hennigi, Ceratopogon margaritae, Ceratopogon tertiaricus, Eohelea gedanica, Eohelea grogani, Eohelea petrunkevitchi, Fossihelea gracilitarsis, Gedanohelea loewi, Meunierohelea sp., Meunierohelea sp. B, Meunierohelea sp. C, Meunierohelea nielseni, Monohelea clunipes, Nannohelea sp., Serromyia sp., Serromyia sp. A, Serromyia sp. B, Serromyia spinigera, Serromyia succinea, Stilobezzia falcata, Physohelea obtusa, Neurohelea cothurnata, Bezzia eocenica, Eohelea stridulans, Forcipomyia turbinata, Forcipomyia uncula, Forcipomyia sp., Forcipomyia piriformis, Culicoides sp. A, Culicoides sp. B, Culicoides sp. C, Brachypogon sp., Brachypogon sp. A, Atrichopogon eocenicus, Forcipomyia eocostata, Forcipomyia gedanicola, Dasyhelea eodicryptoscenica, Dasyhelea gedanica, Dasyhelea stanislavi, Culicoides ceranowiczi, Culicoides eoselficus, Culicoides gedanensis, Alluaudomyia succinea, Brachypogon balticus, Brachypogon eocenicus, Brachypogon gedanicus, Brachypogon henningseni, Brachypogon prominulus, Ceratopogon ceranowiczi, Ceratopogon crypticus, Ceratopogon gedanicus, Ceratopogon ritzkowskii, Gedanohelea succinea, Meunierohelea gedanicola, Monohelea baltica, Serromyia anomalicornis, Serromyia polonica, Serromyia ryszardi, Serromyia sinuosa, Wirthohelea trifida, Palpomyia jantari, Culicoides speciosus, Culicoides dasyheleiformis, Culicoides succivarius, Culicoides balticus, Ceratopogon remmicolus, Forcipomyia krzeminskii, Forcipomyia kulickae, Forcipomyia eobreviflagellata, Eohelea sp. A, Eohelea sp. B (120813) Ceratopogon sp., Culicoides sp., Forcipomyia sp., Forcipomyia sp., Eohelea sinuosa, Meunierohelea sp., Serromyia sp., Culicoides speciosus (154301) Ceratopogon sp., Dasyhelea sp. A, Forcipomyia spp., Forcipomyia sp., Fossihelea gracilitarsis, Meunierohelea sp. C, Ceratopogon clunipes, Serromyia sp., Neurohelea cothurnata, Ceratopogon spiniger, Forcipomyia uncula, Forcipomyia berendti, Forcipomyia gedanicola, Forcipomyia eophytoheleana, Forcipomyia eotrichoheleana, Dasyhelea eodicryptoscenica, Culicoides prussicus, Brachypogon balticus, Brachypogon prominulus, Ceratopogon anomalicornis, Culicoides dasyheleiformis (109625) Ceratopogon sp., Dasyhelea sp., Fossihelea gracilitarsis, Monohelea clunipes, Brachypogon sp., Brachypogon balticus, Brachypogon polonicus, Brachypogon prominulus, Palpomyia riedeli (153481) Mantohelea gedanica, Metahelea serafini, Mallochohelea martae, Monogedania clunipes, Schizohelea baltica (110737)
Priabonian	38.0 - 33.9	Ukraine	Ceratopogon sp. (122999) Ceratopogon sp., Stilobezzia sp., Forcipomyia sp., Brachypogon sp., Eohelea sinuosa, Meunierohelea sp., Monohelea clunipes (122996) Culicoides sp., Stilobezzia sp., Eohelea sinuosa, Meunierohelea sp. (122995) Palpomyia sp., Ceratopogon sp., Culicoides sp., Dasyhelea sp., Mantohelea laca, Stilobezzia sp., Forcipomyia sp., Leptoconops rovnensis, Brachypogon sp., Culicoides speciosus, Brachypogon balticus, Brachypogon prominulus, Ceratopogon bitterfeldi, Ceratopogon forcipiformis, Ceratopogon grogani, Ceratopogon hennigi, Ceratopogon margaritae, Ceratopogon tertiaricus, Eohelea fossicola, Eohelea gedanica, Eohelea miocaenea, Eohelea petrunkevitchi, Eohelea sinuosa, Fossihelea sp., Fossihelea gracilitarsis, Fossihelea miocaenica, Gedanohelea loewi, Meunierohelea sp., Meunierohelea nielseni, Monohelea sp., Monohelea clunipes, Nannohelea sp., Nannohelea grogani, Serromyia sp., Serromyia spinigera, Serromyia succinea, Stilobezzia falcata, Physohelea obtusa, Neurohelea sp., Neurohelea cothurnata, Bezzia eocenica (122997) Palpomyia sp., Ceratopogon sp., Culicoides sp., Stilobezzia sp., Forcipomyia sp., Brachypogon sp., Culicoides speciosus, Brachypogon balticus, Brachypogon prominulus, Ceratopogon eminens, Ceratopogon forcipiformis, Eohelea gedanica, Eohelea grogani, Eohelea sinuosa, Meunierohelea sp., Monohelea clunipes, Stilobezzia falcata, Bezzia eocenica (113621)
Priabonian	38.0 - 33.9	United Kingdom (England)	Dasyhelea sp., Stilobezzia sp., Palpomyia edwardsi (123959) Stilobezzia sp. (21976)
Priabonian	38.0 - 33.9	Denmark	Ceratopogon sp., Culicoides sp., Dasyhelea sp., Stilobezzia sp., Stilobezzia sp. A, Ceratopogoninae indet., Forcipomyia spp., Forcipomyia sp., Forcipomyia sp., Forcipomyia sp. B, Ceratopogonini indet., Culicoides speciosus, Ceratopogon eminens, Ceratopogon forcipiformis, Ceratopogon hennigi, Ceratopogon tertiaricus, Eohelea gedanica, Eohelea petrunkevitchi, Eohelea sinuosa, Fossihelea gracilitarsis, Meunierohelea sp., Meunierohelea sp. A, Meunierohelea nielseni, Monohelea clunipes, Nannohelea sp., Nannohelea grogani, Serromyia sp., Stilobezzia falcata, Neurohelea cothurnata, Bezzia eocenica, Eohelea stridulans, Forcipomyia uncula, Forcipomyia piriformis, Culicoides sp. B, Brachypogon sp. A, Forcipomyia henningseni, Forcipomyia gedanicola, Forcipomyia lyneborgi, Forcipomyia succinea, Forcipomyia eophytoheleana, Culicoides eoselficus, Alluaudomyia sp. A, Alluaudomyia succinea, Brachypogon balticus, Brachypogon eocenicus, Brachypogon gedanicus, Brachypogon henningseni, Brachypogon prominulus, Ceratoculicoides danicus, Ceratopalpomyia eocenica, Ceratopogon crypticus, Ceratopogon piotrowskii, Ceratopogon ritzkowskii, Gedanohelea wirthi, Meunierohelea wirthi, Monohelea baltica, Nannohelea eocenica, Palpomyia succinea, Culicoides balticus, Stilobezziini indet., Forcipomyia krzeminskii, Leptoconops succineus, Ceratopogon sp. B, Ceratopogon sp. A, Ceratopogon sp. C (123762)
Priabonian	38.0 - 33.9	Russian Federation	Ceratopogon lacus (123419)
Early/Lower Oligocene	33.9 - 28.4	France	Simulium terribilis (121474)
MP 30	28.4 - 23.03	Germany (Nordrhein-Westfalen)	Tetragoneura veterana (138696)
MP 30	28.4 - 23.03	Germany	Ceratopogon alpheus, Simulia pasithea (123945)
Chattian	28.1 - 23.03	Germany (North Rhine-Westphalia)	Ceratopogon pupa 13, Ceratopogon pupa 12, Ceratopogon pupa 11, Ceratopogon pupa 10, Ceratopogon pupa 9, Ceratopogon pupa 8, Ceratopogon pupa 7, Ceratopogon pupa 6, Ceratopogon pupa 5, Ceratopogon pupa 4, Ceratopogon pupa 3, Ceratopogon pupa 2, Ceratopogon pupa 1, Ceratopogon sp., Serromyia sp., Culicoides abbreviatipennis, Culicoides atratus, Culicoides austerus, Culicoides bicolor, Culicoides elongatulus, Culicoides gracilior, Culicoides jucundus, Culicoides liliputanus, Culicoides obesus, Culicoides tenuipennis, Culicoides abdominalis, Serromyia austera, Serromyia colorata, Serromyia spinofemorata, Stilobezzia goetghebueri, Bezzia longipennis, Atrichopogon brunnescens, Culicoides obscuratus, Stilobezzia veterana (122010)
Chattian	28.1 - 23.03	New Zealand (Otago)	Austrohelea sp. (191651)
Chattian	28.1 - 23.03	France (Provence-Alpes-Côte d'Azur)	Ceratopogon sp. (122488)
Early/Lower Miocene	23.03 - 15.97	Ethiopia	Forcipomyia rasnitsyni (222682)
Early/Lower Miocene	23.03 - 15.97	USA (California)	Dasyhelea dara (152225) Dasyhelea judithae (152224) Dasyhelea stenoceras (152227)
Early/Lower Miocene	23.03 - 15.97	Mexico	Culicoides sp., Forcipomyia sp., Forcipomyia sp. 1, Forcipomyia sp. 2, Brachypogon sp. (21977)
Early/Lower Miocene	23.03 - 15.97	Mexico (Chiapas)	Dasyhelea sp. (138882 140834) Dasyhelea sp., Forcipomyia sp., Nannohelea sp. (140217) Forcipomyia sp. (106432 124467 140215 140833) Forcipomyia sp., Brachypogon sp. (124474) Nannohelea sp. (140212)
Burdigalian	20.44 - 15.97	New Zealand (Otago)	Dasyhelea sp., Forcipomyia sp. (191650)
Burdigalian - Langhian	20.44 - 13.82	Dominican Republic	Brachypogon dominicanus, Forcipomyia sp., Forcipomyia antilleana, Forcipomyia chrysosuccinea, Forcipomyia domibicolor (125970) Dasyhelea sp., Forcipomyia sp., Forcipomyia sp., Forcipomyia sp., Culicoides brodzinskyi, Culicoides ambericus, Brachypogon dominicanus, Stilobezzia dominicana, Stilobezzia antilleana, Baeodasymyia dominicana, Phaenobezzia wirthi, Culicoides sp., Brachypogon sp., Dasyhelea minuticola, Forcipomyia grimaldii, Forcipomyia woodruffi, Forcipomyia sp., Forcipomyia antilleana, Forcipomyia domibicolor, Forcipomyia sp. (122982) Dasyhelea sp., Forcipomyia sp., Forcipomyia sp., Nannohelea sp. A, Culicoides antilleanus, Culicoides ambericus, Brachypogon dominicanus, Brachypogon prominuloides, Stilobezzia dominicana, Stilobezzia antilleana, Culicoides mammalicolus, Baeodasymyia dominicana, Palpomyia wirthorum, Culicoides sp., Brachypogon sp., Stilobezzia sp. A, Dasyhelea antilleana, Dasyhelea dominicana, Atrichopogon sp., Atrichopogon dominicanus, Forcipomyia grimaldii, Forcipomyia americana, Forcipomyia woodruffi, Forcipomyia sp., Forcipomyia antilleana, Forcipomyia domibicolor, Forcipomyia lepidosuccinea, Forcipomyia sp., Forcipomyia frutetosuccinea, Forcipomyia tertiaricola, Forcipomyia sp. (122473) Dasyhelea sp., Forcipomyia sp., Forcipomyia sp., Nannohelea sp., Culicoides hispanicolus, Brachypogon americanus, Culicoides sp., Brachypogon sp., Brachypogon sp., Dasyhelea dominicana, Dasyhelea hispaniolae, Forcipomyia fusiparamera, Forcipomyia grimaldii, Forcipomyia woodruffi, Forcipomyia sp., Forcipomyia antilleana, Forcipomyia domibicolor (123857) Forcipomyia sp., Forcipomyia sp. (123393) Forcipomyia sp., Forcipomyia sp., Stilobezzia dominicana, Brachypogon sp., Forcipomyia antilleana (140835) Heteromyia dominicana (122474) Meunierohelea fudalai (124427)
Hemingfordian	20.43 - 15.97	USA (California)	Culicoides carri (152228) Culicoides fossilis (144522) Culicoides miocenea (152207 152230) Dasyhelea antiqua (123023 123025 123026 123027 123028 123029 123032 123033 123036 123037 123038 123039 123040 123041 123043 152213 152218) Dasyhelea antiqua, Culicoides megacanthus (123034) Dasyhelea antiqua, Dasyhelea dara (152215) Dasyhelea antiqua, Dasyhelea kanakoffi (152214) Dasyhelea antiqua, Dasyhelea stenoceras (123031 123042) Dasyhelea antiqua, Dasyhelea stenoceras, Culicoides megacanthus (123022 123030) Dasyhelea antiqua, Dasyhelea stenoceras, Culicoides megacanthus, Dasyhelea dara, Dasyhelea judithae, Dasyhelea kanakoffi, Dasyhelea browneae, Culicoides miocenea, Neopalpomyia freyi, Parapalpomyia ryshkoffi, Johannsenomyia hotchkissae (152222) Dasyhelea antiqua, Dasyhelea stenoceras, Dasyhelea dara, Dasyhelea judithae, Dasyhelea kanakoffi, Culicoides laurae, Culicoides miocenea, Neopalpomyia freyi, Parapalpomyia ryshkoffi, Johannsenomyia hotchkissae (152229) Dasyhelea antiqua, Dasyhelea stenoceras, Dasyhelea dara, Dasyhelea judithae, Dasyhelea kanakoffi, Dasyhelea browneae, Culicoides miocenea, Neopalpomyia freyi, Parapalpomyia ryshkoffi, Johannsenomyia hotchkissae (152216 152217) Dasyhelea antiqua, Dasyhelea stenoceras, Dasyhelea dara, Dasyhelea judithae, Dasyhelea kanakoffi, Neopalpomyia freyi, Parapalpomyia ryshkoffi (152219) Dasyhelea antiqua, Dasyhelea stenoceras, Dasyhelea dara, Dasyhelea judithae, Dasyhelea kanakoffi, Parapalpomyia ryshkoffi, Johannsenomyia hotchkissae (152221) Dasyhelea antiqua, Dasyhelea stenoceras, Dasyhelea kanakoffi, Neopalpomyia freyi, Parapalpomyia ryshkoffi, Johannsenomyia hotchkissae (152220) Dasyhelea antiqua, Neopalpomyia freyi (152212) Dasyhelea australis (123035) Dasyhelea browneae (152223) Dasyhelea dara, Dasyhelea judithae, Johannsenomyia hotchkissae, Neoculicoides jeanneae (144521) Dasyhelea dara, Dasyhelea kanakoffi (152226) Miopalpomyia sp., Dasyhelea antiqua, Culicoides megacanthus, Johannsenomyia hotchkissae (106431) Neopalpomyia multispinosa (152232) Parapalpomyia ryshkoffi, Miopalpomyia shilo (152231)
Middle Miocene	15.97 - 11.608	Australia (New South Wales)	Ceratopogonidae indet. (224044)
Holocene	0.0117 - 0.0	New Zealand	Heleidae indet. (164242)
Holocene	0.0117 - 0.0	Madagascar	Ceratopogon viscatus, Ceratopogon madagascariensis, Ceratopogon crassipalpis, Ceratopogon tenuipes (151372)
Holocene	0.0117 - 0.0	Tanzania	Ceratopogonidae indet. (234540)