PBDB Taxon

Basic info	Taxonomic history	Classification	Included Taxa
Morphology	Ecology and taphonomy	External Literature Search	Age range and collections

Atriculicoides

Insecta - Diptera - Ceratopogonidae

Taxonomy

Atriculicoides was named by Remm (1976). It is not extant. Its type is Atriculicoides macrophthalmus. It is the type genus of Atriculicoidini.

It was synonymized subjectively with Protoculicoides by Szadziewski et al. (2016) and Szadziewski (2018).

It was assigned to Ceratopogonini by Remm (1976); to Ceratopogonidae by Szadziewski and Schlüter (1992) and Carpenter (1992); to Atriculicoidini by Szadziewski (1996) and Szadziewski (2004); to Forcipomyiinae by Borkent and Wirth (1997) and Borkent (2013); and to Atriculicoidinae by Borkent (2019) and Borkent and Dominiak (2020).

Species

A. cenomanensis, A. ciliatus, A. dasyheleis, A. globosus (syn. Lasiohelea cretea), A. hispanicus, A. incompletus, A. macrophthalmus (syn. A. squamiciliatus) (type species), A. sanjusti, A. sibiricus, A. swinhoei, A. szadziewskii, A. taimyricus

Synonymy list

Year	Name and author
1976	Atriculicoides Remm p. 345
1992	Atriculicoides Carpenter
1992	Atriculicoides Szadziewski and Schlüter p. 74
1996	Atriculicoides Szadziewski p. 51
1997	Atriculicoides Borkent and Wirth p. 29
2004	Atriculicoides Szadziewski p. 119
2013	Atriculicoides Borkent p. 27
2019	Atriculicoides Borkent p. 21
2020	Atriculicoides Borkent and Dominiak p. 40

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Rank	Name	Author
kingdom	Animalia	()
—	Bilateria
—	Eubilateria	Ax 1987
—	Protostomia
—	Ecdysozoa
—	Panarthropoda
phylum	Arthropoda	Latreille 1829
superclass	Hexapoda	Latreille 1825

Rank	Name	Author
class	Insecta
order	Diptera
infraorder	Culicomorpha
superfamily	Chironomoidea	Macquart 1838
family	Ceratopogonidae
subfamily	Atriculicoidinae	()
genus	Atriculicoides

If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.

G. †Atriculicoides Remm 1976

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†Atriculicoides cenomanensis Szadziewski and Schlüter 1992

†Atriculicoides ciliatus Borkent 2012

†Atriculicoides dasyheleis Szadziewski 1996

†Atriculicoides globosus Boesel 1937

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Invalid names: Lasiohelea cretea Boesel 1937 [synonym]

†Atriculicoides hispanicus Szadziewski and Arillo 2016

†Atriculicoides incompletus Szadziewski and Schlüter 1992

†Atriculicoides macrophthalmus Remm 1976

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Invalid names: Atriculicoides squamiciliatus Remm 1976 [synonym]

†Atriculicoides sanjusti Szadziewski and Arillo 2016

†Atriculicoides sibiricus Szadziewski 1996

†Atriculicoides swinhoei Cockerell 1919

†Atriculicoides szadziewskii Pérez-de la Fuente et al. 2011

†Atriculicoides taimyricus Szadziewski 1996

Diagnosis

No diagnoses are available

Measurements

No measurements are available

Composition:

chitinc

Environment:

terrestrialc

Locomotion:

actively mobilep

Created:

2017-04-17 09:56:18

Modified:

2017-04-17 09:56:18

Source: c = class, p = phylum

References: Bush and Bambach 2015, Kiessling 2004

Age range: base of the Early/Lower Albian to the top of the Late/Upper Campanian or 112.03000 to 70.60000 Ma

Collections (19 total)

Time interval	Ma	Country or state	Original ID and collection number
Early/Lower Albian	112.03 - 109.0	Spain	A. szadziewskii (109438)
Late/Upper Albian	105.3 - 99.6	Spain (Aragón)	Protoculicoides hispanicus, Protoculicoides sanjusti (91626)
Early/Lower Cenomanian	99.6 - 93.5	France	Ceratopogonidae indet. A, Ceratopogonidae indet. B (123018)
Early/Lower Cenomanian	99.6 - 93.5	Myanmar	A. sp., Johannsenomyia swinhoei (123020)
Early/Lower Cenomanian	99.6 - 93.5	Myanmar (Kachin)	A. swinhoei (104053)
Late/Upper Cenomanian	99.6 - 93.5	Russian Federation	A. sp. (35051)
Turonian	93.9 - 89.8	USA (New Jersey)	A. globosus (123016) A. incompletus, A. globosus (110895)
Late/Upper Santonian	85.8 - 83.5	Russian Federation	A. sp., A. macrophthalmus, A. squamiciliatus (122590) A. sp., A. spp., A. macrophthalmus, A. dasyheleis, A. sibiricus, A. taimyricus (35056)
Late/Upper Campanian	83.5 - 70.6	Canada (Manitoba)	A. sp., A. globosus (122353 122354) A. sp., Lasiohelea globosa, Lasiohelea cretea (122356)
Middle Campanian	83.5 - 70.6	Canada (Alberta)	A. globosus (104932 122383 122388) A. globosus, Protoculicoides depressus (122384) A. sp. (34896) A. sp., A. globosus (122389)