PBDB Taxon

Basic info	Taxonomic history	Classification	Included Taxa
Morphology	Ecology and taphonomy	External Literature Search	Age range and collections

Cleiothyridina grossula

Rhynchonellata - Athyridida - Athyrididae

Taxonomy

Athyris grossula was named by Waagen (1883). It is a 3D body fossil. Its type locality is Chidru, "Cephalopod Bed", Salt Range, which is in a Wuchiapingian carbonate limestone in the Chhidru Formation of Pakistan.

It was recombined as Cleiothyridina grossula by Clapham (2015).

Sister species lacking formal opinion data

C. angelicoides, C. coloradoensis, C. crassicardinalis, C. deroissii, C. elegans, C. fimbriata, C. glabristria, C. humerosa, C. incrassata, C. media, C. pecossi, C. segmentata, C. squamosa, C. transversa

Synonymy list

Year	Name and author
1883	Athyris grossula Waagen p. 474 figs. Pl 38, figs 9,10
2015	Cleiothyridina grossula Clapham

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Rank	Name	Author
kingdom	Animalia	()
—	Bilateria
—	Eubilateria	Ax 1987
—	Protostomia
—	Spiralia
superphylum	Lophotrochozoa
—	Lophophorata
—	Panbrachiopoda	Carlson and Cohen 2020
phylum	Brachiopoda	Cuvier 1805

Rank	Name	Author
subphylum	Rhynchonelliformea	Williams et al. 1996
class	Rhynchonellata	Williams et al. 1996
order	Athyridida	Boucot et al. 1964
suborder	Athyrididina	Boucot et al. 1964
superfamily	Athyridoidea	McCoy 1844
family	Athyrididae	McCoy 1841
subfamily	Cleiothyridininae
genus	Cleiothyridina	Buckman 1906
species	grossula	()

If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.

†Cleiothyridina grossula Waagen 1883

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Diagnosis

No diagnoses are available

Measurements

part	N	mean	minimum	maximum	median	s.d.
shell height	2	10.2	10.0	10.5	10.2	0.354

Composition:

low Mg calcitep

Environment:

marineuc

Locomotion:

stationarysubo

Attached:

yesp

Life habit:

low-level epifaunalsubo

Diet:

suspension feedersubo

Vision:

blindc

Created:

2018-11-10 18:06:04

Modified:

2018-11-10 18:06:04

Source: subo = suborder, c = class, p = phylum, uc = unranked clade

References: Nesnidal et al. 2013, Aberhan et al. 2004

Age range: base of the Wuchiapingian to the top of the Changhsingian or 259.90000 to 252.17000 Ma

Collections (41 total)

Time interval	Ma	Country or state	Original ID and collection number
Capitanian	264.28 - 259.51	Pakistan	Athyris grossula (125883 127001 127032 127033)
Wuchiapingian	259.9 - 254.17	Pakistan	Athyris grossula (type locality: 88933 90253 125901 125918 125963 126003 126008 126264 126268 126272 126275 126279 126293 126296 126298 126299 126342 126343 126345 126458 126692 126693 126937 127007 127025 127028)
Wuchiapingian - Changhsingian	259.9 - 252.17	Pakistan	Athyris grossula (126704 126740 126750 126753 126755 126762 126886 127014)
Changhsingian	254.17 - 252.17	Pakistan	Athyris grossula (126896 126934 126981)