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Dryinidae
Taxonomy
Dryini was named by Haliday (1833). It is extant. Its type is Dryinus.
It was corrected as Dryinidae by Rasnitsyn (1975), Carpenter (1992), Brothers (1999), Engel (2005), Sharkey (2007), Martynova et al. (2019) and Perkovsky et al. (2019).
It was assigned to Bethyloidea by Rasnitsyn (1975); to Apocrita by Carpenter (1992); and to Chrysidoidea by Brothers (1999), Engel (2005), Sharkey (2007), Martynova et al. (2019) and Perkovsky et al. (2019).
It was corrected as Dryinidae by Rasnitsyn (1975), Carpenter (1992), Brothers (1999), Engel (2005), Sharkey (2007), Martynova et al. (2019) and Perkovsky et al. (2019).
It was assigned to Bethyloidea by Rasnitsyn (1975); to Apocrita by Carpenter (1992); and to Chrysidoidea by Brothers (1999), Engel (2005), Sharkey (2007), Martynova et al. (2019) and Perkovsky et al. (2019).
Synonymy list
| Year | Name and author |
|---|---|
| 1975 | Dryinidae Rasnitsyn p. 104 |
| 1992 | Dryinidae Carpenter |
| 1999 | Dryinidae Brothers p. 243 |
| 2005 | Dryinidae Engel p. 486 |
| 2007 | Dryinidae Sharkey |
| 2019 | Dryinidae Martynova et al. p. 1013 |
| 2019 | Dryinidae Perkovsky et al. p. 170 |
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If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.
Fm. Dryinidae Haliday 1833
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Subfm. Anteoninae Kieffer 1907
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G. Deinodryinus Perkins 1907
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†Deinodryinus aptianus Olmi et al. 2010
†Deinodryinus areolatus Ponomareko 1975
†Deinodryinus velteni Guglielmino and Olmi 2011
Invalid names: Electrodryinus Ponomareko 1975 [synonym]
G. †Janzeniola Olmi 2009
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†Janzeniola baltica Olmi 2000
Invalid names: Janzenia Olmi 2000 [replaced]
G. Lonchodryinus Kieffer 1905
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†Lonchodryinus balticus Olmi and Guglielmino 2012
†Lonchodryinus groehni Olmi et al. 2021
Subfm. †Archaeodryininae Olmi et al. 2020
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Subfm. Bocchinae Richards 1939
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G. Bocchus Ashmead 1893
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†Bocchus primaevus Martins and Melo 2019
†Bocchus rex Olmi et al. 2022
†Bocchus schmalhauseni Perkovsky et al. 2020
†Bocchus vetustus Olmi 1991
Subfm. †Burmadryininae Olmi et al. 2014
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Subfm. Dryininae Haliday 1833
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G. Dryinus Latreille 1804
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†Dryinus acutifrons Martynova et al. 2019
†Dryinus alamellatus Olmi and Guglielmino 2011
†Dryinus antiquus Ponomarenko 1981
†Dryinus arakanesis Martynova et al. 2019
†Dryinus balticus Olmi 1984
†Dryinus bamar Martynova et al. 2019
†Dryinus brachytarsis Martynova et al. 2019
†Dryinus bruesi Olmi 1984
†Dryinus burmensis Martynova et al. 2019
†Dryinus canadensis Ponomarenko 1981
†Dryinus carsteni Olmi et al. 2022
†Dryinus chin Martynova et al. 2019
†Dryinus grimaldii Olmi 1995
†Dryinus hymenaeaphilus Olmi 1995
†Dryinus janzeni Olmi 2000
†Dryinus katyae Martynova et al. 2019
†Dryinus laminatus Martynova et al. 2019
†Dryinus latitarsis Martynova et al. 2019
†Dryinus maderai Guglielmino et al. 2018
†Dryinus mortuorum Brues 1933
†Dryinus muenchi Olmi and Bechly 2001
†Dryinus nadezhdae Martynova et al. 2019
†Dryinus palaeodominicanus Currado and Olmi 1983
†Dryinus palaeomexicanus Olmi 1995
†Dryinus parisei Martynova et al. 2019
†Dryinus poinari Olmi 1998
†Dryinus pristinus Olmi 1998
†Dryinus rasnitsyni Olmi and Guglielmino 2011
†Dryinus reifi Olmi and Bechly 2001
†Dryinus taron Martynova et al. 2019
†Dryinus velteni Olmi 2011
†Dryinus vetus Brues 1933
†Dryinus vetustus Olmi 1995
†Dryinus wunderlichi Olmi and Bechly 2001
Invalid names: Avodryinus Ponomarenko 1981 [synonym], Richardsidryinus Moczar 1965 [synonym]
G. †Harpactosphecion Haupt 1944
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†Harpactosphecion deletum Brues 1933
†Harpactosphecion filicorne Brues 1923
†Harpactosphecion filicornis Brues 1923
†Harpactosphecion gracile Brues 1933
†Harpactosphecion scheveni Olmi 2005
†Harpactosphecion sucinum Olmi 1987
G. †Hybristodryinus Engel 2005
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†Hybristodryinus anomalus Perkovsky et al. 2019
†Hybristodryinus castaneus Olmi et al. 2021
†Hybristodryinus concavifrons Perkovsky et al. 2019
†Hybristodryinus cretacicus Perkovsky et al. 2019
†Hybristodryinus karen Perkovsky et al. 2019
†Hybristodryinus kayin Perkovsky et al. 2019
†Hybristodryinus konbaung Perkovsky et al. 2019
†Hybristodryinus ligulatus Perkovsky et al. 2019
†Hybristodryinus magnificus Perkovsky et al. 2019
†Hybristodryinus mon Perkovsky et al. 2019
†Hybristodryinus moutesoe Tribull et al. 2020
†Hybristodryinus nalae Perkovsky et al. 2019
†Hybristodryinus pyu Perkovsky et al. 2019
†Hybristodryinus resinicolus Engel 2005
†Hybristodryinus shan Perkovsky et al. 2019
†Hybristodryinus viriosus Wang et al. 2021
†Hybristodryinus zaifui Chen et al. 2021
G. Pseudodryinus Olmi 1991
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†Pseudodryinus burmensis Olmi et al. 2014
†Pseudodryinus parisiensis Peinado et al. 2006
Invalid names: Dryinini Haliday 1833 [invalid subgroup]
Subfm. Gonatopodinae
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G. Gonatopus Ljungh 1810
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†Gonatopus cretacicus Olmi et al. 2010
Invalid names: Dicondylus Curtis 1829 [synonym], Laberius Kieffer 1914 [synonym]
Subfm. †Palaeoanteoninae Olmi and Bechly 2001
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G. †Palaeoanteon Olmi 2000
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†Palaeoanteon cenomanianum Perkovsky et al. 2020
†Palaeoanteon janzeni Olmi 2000
Subfm. †Protodryininae Olmi and Guglielmino 2012
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Subfm. †Raptodryininae Olmi et al. 2020
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Subfm. Thaumatodryininae Perkins 1905
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G. Thaumatodryinus Perkins 1905
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†Thaumatodryinus fuscescens Martins and Melo 2020
†Thaumatodryinus miocenicus Olmi 1995
†Thaumatodryinus priscus Olmi 1998
Diagnosis
No diagnoses are available
Measurements
No measurements are available
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| Source: c = class, p = phylum | |||||
| References: Kiessling 2004, Bush and Bambach 2015 | |||||
Age range
Maximum range based only on fossils: base of the Early/Lower Barremian to the top of the Early/Lower Miocene or 130.00000 to 15.97000 Ma
Minimum age of oldest fossil (stem group age): 125.45 Ma
Minimum age of oldest fossil (stem group age): 125.45 Ma
Collections (48 total)
| Time interval | Ma | Country or state | Original ID and collection number |
|---|---|---|---|
| Early/Lower Barremian | Lebanon | Aphelopus palaeophoenicius (124363) | |
| Aptian | Mongolia | Deinodryinus aptianus (113355) | |
| Early/Lower Cenomanian | Myanmar | Burmanteon olmii, Hybristodryinus moutesoe (111688) Dryinus burmensis, Dryinus katyae (161726) Dryinus carsteni (172900) Dryinus maderai, Hybristodryinus anomalus, Hybristodryinus concavifrons, Hybristodryinus cretacicus, Hybristodryinus karen, Hybristodryinus kayin, Hybristodryinus konbaung, Hybristodryinus ligulatus, Hybristodryinus magnificus, Hybristodryinus mon, Hybristodryinus nalae, Hybristodryinus pyu, Hybristodryinus shan, Dryinus acutifrons, Dryinus arakanesis, Dryinus bamar, Dryinus brachytarsis, Dryinus chin, Dryinus laminatus, Dryinus latitarsis, Dryinus nadezhdae, Dryinus parisei, Dryinus taron, Raptodryinus patrickmuelleri, Palaeoanteon cenomanianum, Hybristodryinus castaneus, Rasnitsynum burmense (194884) Dryinus taron (206272) Hybristodryinus cretacicus, Dryinus taron, Hybristodryinus zaifui, Hybristodryinus viriosus (165561) Hybristodryinus resinicolus (123116) Hybristodryinus resinicolus, Dryinus maderai, Hybristodryinus kayin, Hybristodryinus konbaung, Hybristodryinus nalae, Hybristodryinus pyu, Dryinus bamar, Dryinus katyae, Dryinus parisei (184692) Pseudodryinus burmensis, Burmadryinus cenomanianus (123053) Thaumatorrhinos athrix (216576) | |
| Late/Upper Cenomanian | Russian Federation | Laberius antiquus (35051) | |
| Late/Upper Santonian | Russian Federation | Cretodryinus zherichini (122590) | |
| Early/Lower Campanian | Russian Federation (Tenkinskii) | Gonatopus cretacicus, Anteonopsis antiquus (105375) | |
| Middle Campanian | Canada (Alberta) | Avodryinus canadensis (122384) | |
| Ypresian | France (Oise) | Pseudodryinus parisiensis (123850) | |
| Priabonian | Denmark | Alphadryinus balticus, Alphadryinus bruesi (123762) | |
| Priabonian | Ukraine | Bocchus rex (223355 227091) Bocchus schmalhauseni, Rovnodryinus khomychi (216854) Dryinus janzeni (196704) Dryinus reifi, Dryinus wunderlichi (207953) Palaeoanteon janzeni (185684) | |
| Priabonian | Russian Federation (Kaliningrad) | Bocchus primaevus (217266) Dryinus mortuorum, Harpactosphecion filicornis (156497) Dryinus muenchi, Dryinus reifi, Dryinus wunderlichi, Palaeoanteon janzeni, Harpactosphecion filicorne, Deinodryinus velteni, Dryinus bruesi, Dryinus janzeni, Lonchodryinus balticus, Protodryinus eocenicus (120814) Dryinus sp., Palaeoanteon janzeni, Janzenia baltica, Dryinus janzeni, Harpactosphecion filicornis (154633) Electrodryinus areolatus (122591) Harpactosphecion filicorne (123773 124318) Harpactosphecion filicornis (123757) Palaeodryinus groehni, Harpactosphecion gracile, Dryinus janzeni, Bocchus primaevus, Lonchodryinus groehni (110735) Thaumatodryinus gracilis, Thaumatodryinus filicornis, Dryinus filicornis, Thaumatodryinus deletus, Lestodryinus mortuorum, Lestodryinus vetus, Neodryinus somniatus (123927) | |
| Priabonian | Russian Federation | Harpactosphecion gracile, Dryinus janzeni (123419) | |
| Chattian | Russian Federation | Dryinidae indet. (125971) | |
| Early/Lower Miocene | Mexico (Chiapas) | Dryinus palaeomexicanus (124928) | |
| Burdigalian - Langhian | Dominican Republic | Bocchus vetustus (159268) Dryinus grimaldii, Dryinus hymenaeaphilus, Dryinus priscus, Dryinus pristinus, Dryinus vetustus, Pseudodryinus sucinus, Dryinidae indet. (122982) Dryinus palaeodominicanus (154719 166828) Dryinus palaeodominicanus, Alphadryinus sucinus (157305) Dryinus palaeodominicanus, Harpactosphecion scheveni (124846) Dryinus rasnitsyni, Harpactosphecion sucinum, Thaumatodryinus miocenicus, Dryinus alamellatus (122474) Dryinus sp., Dryinus grimaldii, Aphelopus poinari, Dryinus palaeodominicanus, Dryinus poinari, Harpactosphecion sucinum, Thaumatodryinus miocenicus (122472) Thaumatodryinus fuscescens (203120) Thaumatodryinus miocenicus (155866 155867) |