PBDB

Basic info	Taxonomic history	Classification	Included Taxa
Morphology	Ecology and taphonomy	External Literature Search	Age range and collections

Neoptera

Insecta

Taxonomy

Neoptera was named by Martynov (1923). It is extant.

It was assigned to Pterygota by Martynov (1928), Carpenter (1992), Kukalová-Peck and Lawrence (2004) and Misof et al. (2014); and to Neopterygota by Sroka et al. (2015).

Subtaxa

Ampelipteridae, Apheloneuridae, Archaemiopteridae, Archipanorpidae, Asiuropidae, Baryshnyalidae, Blattoneoptera, Boltonocostidae (syn. Orthocostidae), Choristosialidae, Clareocercaria, Controversala, Dictyodipteridae, Dictyophara scudderi, Endoneoptera, Eopanorpella, Exopterygota, Gelasopteridae, Glosselytrodea (syn. Jurinida), Hemineoptera, Holometabola (syn. Endopterygota), Mesophthiridae, Metropatoridae, Metropatorites, Micropalentomidae, Mixotermitoidea, Naucoris crassus, Oiophassus, Orthoneoptera, Palaeoheteroptera, Palaeonepidoideus, Paraneoptera, Perielytridae, Permembiidae, Permosialidae (syn. Permonkidae, Tologopteridae, Epimastacidae, Perloblattidae), Polyneoptera, Prosigara, Psocodea (syn. Psocoptera, Copeognatha), Saurodectidae, Sibosmylina, Sinembiidae, Sphalmatoblattina, Sycopteridae, Sycopteron, Synomaloptilidae, Tillyardites, Trachopterygidae, Triadochorista, Trichorthophlebia, Uninervidae

Synonyms

Paraneoptera is an unranked clade. It is extant.

It was considered an invalid subgroup of Neoptera by Misof et al. (2014).

It was assigned to Neoptera by Nel et al. (2012).
Sycopteridae was named by Tillyard (1918). It is not extant. Its type is Sycopteron.

It was considered an invalid subgroup of Neoptera by Carpenter (1992).

It was assigned to Psocoptera by Tillyard (1918).
Dictyophara scudderi was named by Piton (1940). Its type specimen is Coll. Piton 925, an exoskeleton, and it is a compression fossil. Its type locality is Menat (Piton collection), which is in a Selandian crater lake diatomite in the Menat Formation of France.

It was considered a nomen dubium by Szwedo et al. (2004).
Naucoris crassus was named by Piton (1935). It is a compression fossil. Its type locality is Lac Chambon (Piton coll), which is in a Gelasian crater lake diatomite/volcaniclastic in France.

It was considered a nomen dubium by Nel and Paicheler (1992).

Synonymy list

Year	Name and author
1918	Sycopteridae Tillyard
1923	Neoptera Martynov
1928	Neoptera Martynov p. 10
1935	Naucoris crassus Piton p. 90 figs. Pl 4, fig 4
1940	Dictyophara scudderi Piton p. 162 figs. Pl XVIII fig 8
1992	Neoptera Carpenter
2004	Neoptera Kukalová-Peck and Lawrence
2012	Paraneoptera Nel et al. p. 385
2014	Neoptera Misof et al.
2015	Neoptera Sroka et al. p. 979

Is something missing? Join the Paleobiology Database and enter the data

Rank	Name	Author
kingdom	Animalia	()
—	Bilateria
—	Eubilateria	Ax 1987
—	Protostomia
—	Ecdysozoa
—	Panarthropoda
phylum	Arthropoda	Latreille 1829
—	Deuteropoda
—	Mandibulata
—	Pancrustacea	Zrzavý and Štys 1997
—	Altocrustacea	Regier et al. 2010
superclass	Allotriocarida

Rank	Name	Author
—	Athalassocarida
—	Labiocarida	Schwentner et al. 2017
superclass	Hexapoda	Latreille 1825
—	Cercofilata
class	Insecta
—	Dicondylia
—	Paranotalia
subclass	Pterygota	()
—	Neopterygota	Crampton 1924
infraclass	Neoptera	Martynov 1923
infraclass	Neoptera	Martynov 1923

If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.

Infracl. Neoptera Martynov 1923

show all | hide all

Fm. †Ampelipteridae Haupt 1941

G. †Ampeliptera Pruvost 1927

†Ampeliptera limburgica Pruvost 1927

G. †Tshekardobia Rasnitsyn 1977

†Tshekardobia lurida Novokshonov 2004

†Tshekardobia maculitincta Novokshonov 2004

†Tshekardobia magnifica Novokshonov 2004

†Tshekardobia osmylina Rasnitsyn 1977

†Tshekardobia panorpitincta Novokshonov 2004

G. †Tshunicola Rasnitsyn 1977

†Tshunicola carbonarius Rasnitsyn 1977

Fm. †Apheloneuridae Carpenter 1976

G. †Apheloneura Carpenter 1976

†Apheloneura amplia Carpenter 1976

†Apheloneura minutissima Carpenter 1976

†Apheloneura uralensis Novokshonov 2000

Fm. †Archaemiopteridae Guthörl 1939

G. †Archaemioptera Guthörl 1939

†Archaemioptera carbonaria Guthörl 1939

Fm. †Archipanorpidae Tillyard 1917

G. †Archipanorpa Tillyard 1917

†Archipanorpa magnifica Tillyard 1917

Fm. †Asiuropidae Novokshonov 1997

G. †Asiuropa Novokshonov 1997

†Asiuropa uralensis Novokshonov 1997

Fm. †Baryshnyalidae Ilger and Brauckmann 2011

G. †Baryshnyala Ilger and Brauckmann 2011

†Baryshnyala occulta Ilger and Brauckmann 2011

Fm. †Boltonocostidae Ross et al. 2013

G. †Boltonocosta Carpenter 1986

†Boltonocosta splendens Bolton 1912

Invalid names: Orthocosta Bolton 1912 [replaced]

Invalid names: Orthocostidae Bolton 1912 [replaced]

Fm. †Choristosialidae Tillyard 1932

G. †Choristosialis Tillyard 1932

†Choristosialis enigmatica Tillyard 1932

Unr. Clareocercaria Prokop et al. 2017

Unr. Acercaria Börner 1904

Superor. Pancondylognatha Yoshizawa and Leinhard 2016

Unr. Condylognatha Börner 1904

Or. Hemiptera Linnaeus 1758 [true bug]

Invalid names: Proboscidea Scopoli 1763 [synonym], Ryngota Fabricius 1775 [synonym]

Superor. Thripida Fallen 1814

Or. †Permopsocida Tillyard 1926

Fm. †Archipsyllidae Handlirsch 1906

G. †Archipsylla Handlirsch 1906

G. †Archipsyllodes Vishnyakova 1976

G. †Archipsyllopsis Vishnyakova 1976

G. †Bittacopsocus Beutel et al. 2019

G. †Burmopsylla Liang et al. 2016

G. †Dinmopsylla Lambkin 2018

G. †Eopsylla Vishnyakova 1976

G. †Mydiognathus Yoshizawa and Leinhard 2016

Invalid names: Psocorrhyncha Huang et al. 2016 [synonym]

Fm. †Permopsocidae Tillyard 1926

G. †Lithopsocidium Carpenter 1932

G. †Orthopsocus Carpenter 1932

G. †Permopsocus Tillyard 1926

Invalid names: Ancylopsocus Tillyard 1926 [synonym]

G. †Progonopsocus Tillyard 1926

Fm. †Psocidiidae Tillyard 1926

G. †Austropsocidium Tillyard 1935

G. †Dichentomum Tillyard 1926

Invalid names: Chaetopsocidium Tillyard 1926 [synonym], Metapsocidium Tillyard 1926 [synonym], Pentapsocidium Tillyard 1926 [synonym], Permentomum Tillyard 1926 [synonym], Psocidium Tillyard 1926 [synonym]

G. †Hypopsylla Prokop et al. 2016

G. †Liassopsocus Ansorge 1996

G. †Megapsocidium Tillyard 1935

G. †Parapsocidium Zalessky 1937

G. †Stenopsocidium Tillyard 1935

Invalid names: Dichentomidae Carpenter 1932 [synonym]

Fm. †Westphalopsocidae Azar et al. 2013

G. †Westphalopsocus Azar et al. 2013

†Westphalopsocus pumilio Azar et al. 2013

Or. †Hypoperlida Martynov 1928

Fm. †Hypoperlidae Martynov 1928

G. †Boreopsocus Shcherbakov 1994

†Boreopsocus danksae Shcherbakov 1994

†Boreopsocus ficticius Novokshonov 1995

Invalid names: Psocivena Novokshonov 1995 [synonym]

G. †Hypoperla Martynov 1928

†Hypoperla elegans Martynov 1928

†Hypoperla grata Novokshonova 1998

†Hypoperla nobilis Novokshonov 1995

†Hypoperla vaulevi Novokshonov 2001

G. †Idelopsocus Zalessky 1929

†Idelopsocus arcuatus Martynov 1928

†Idelopsocus diradiatus Rasnitsyn 1996

†Idelopsocus galinae Novokshonov 2001

†Idelopsocus incommendatus Novokshonov et al. 2002

†Idelopsocus levis Novokshonov 1995

†Idelopsocus mutovinus Rasnitsyn and Aristov 2013

†Idelopsocus splendens Zalessky 1948

†Idelopsocus tataricus Zalessky 1929

Invalid names: Hypoperlopsis Zalessky 1948 [synonym], Martynopsocus Karny 1930 [synonym]

G. †Kaltanelmoa Rohdendorf 1961

†Kaltanelmoa sibirica Rohdendorf 1961

G. †Onthomastax Rasnitsyn and Aristov 2013

†Onthomastax coprinus Rasnitsyn and Aristov 2013

Invalid names: Farianopteridae Martynova 1961 [synonym], Kaltanelmoidae Rohdendorf 1961 [synonym], Martynopsocidae Karny 1930 [synonym]

Fm. †Sojanoperidae Novokshonov 2002

G. †Sojanopus Novokshonov 2002

†Sojanopus festivum Novokshonov 2002

Invalid names: Eukulojina Sinitshenkova 1981 [empty], Hypoperlina [invalid subgroup], Strephocladina [invalid subgroup]

Or. †Miomoptera Martynov 1927

G. †Miomina Martins-Neto and Gallego 1999

†Miomina mendozina Martins-Neto and Gallego 1999

†Miomina riojana Martins-Neto and Gallego 2006

Fm. †Palaeomanteidae Handlirsch 1906

G. †Archisialis Martynov 1932

†Archisialis praecox Martynov 1932

G. †Balymotikha Rasnitsyn and Aristov 2013

†Balymotikha deterior Rasnitsyn and Aristov 2013

G. †Belmomantis Prokop et al. 2017

†Belmomantis azari Prokop et al. 2017

G. †Delopsocus Tillyard 1928

†Delopsocus elongatus Sellards 1909

†Delopsocus fasciatus Tillyard 1928

†Delopsocus furcatulus Martynov 1930

†Delopsocus kamensis Martynov 1938

Invalid names: Delopterum kamense Carpenter 1939 [objective synonym]

†Delopsocus kansanum Carpenter 1939

†Delopsocus latus Sellards 1909

†Delopsocus lepidus Kukalová 1963

†Delopsocus sinuosus Kukalová 1963

†Delopsocus stenopterus Rasnitsyn 2004

G. †Delopteriella Zalessky 1956

†Delopteriella graciosa Zalessky 1956

G. †Delopterinus Rasnitsyn 2004

†Delopterinus bicolor Rasnitsyn 2004

†Delopterinus zebra Rasnitsyn 2004

G. †Delopterum Sellards 1909

†Delopterum anale Martynov 1928

†Delopterum candidum Zhuzhgova 2002

†Delopterum commune Rasnitsyn 2004

†Delopterum hangardi Guthorl 1950

†Delopterum iljinskiense Martynova 1961

†Delopterum incertum Martynov 1928

†Delopterum insigne Martynov 1928

†Delopterum kaltanicum Martynova 1961

†Delopterum minutum Sellards 1909

Invalid names: Delopterum dunbari Tillyard 1928 [synonym]

†Delopterum pantherinum Rasnitsyn 2004

†Delopterum radtshenkoi Martynova 1961

†Delopterum rasnitsyni Novokshonov 2000

†Delopterum truncatum Kukalová 1963

†Delopterum zonatum Rasnitsyn 2004

Invalid names: Leptoneurula Martynov 1928 [synonym], Pseudomantis Martynov 1928 [synonym]

G. †Elmomantis Prokop et al. 2017

†Elmomantis engeli Prokop et al. 2017

G. †Mazonopsocus Prokop et al. 2017

†Mazonopsocus testai Prokop et al. 2017

G. †Miomatoneura Martynov 1927

†Miomatoneura angusta Kukalová 1963

†Miomatoneura candida Kukalová 1963

†Miomatoneura frigida Martynov 1927

†Miomatoneura intermedia Martynov 1928

†Miomatoneura palaeozoica Kukalová 1963

†Miomatoneura parvula Martynov 1930

†Miomatoneura permica Kukalová 1963

†Miomatoneura rotundata Martynov 1928

G. †Miomatoneurella Martynova 1958

†Miomatoneurella americana Rasnitsyn 2004

†Miomatoneurella boskovicensis Kukalová 1963

†Miomatoneurella reducta Martynova 1958

†Miomatoneurella rossoshana Rasnitsyn and Aristov 2013

G. †Miomatoneurites Zalessky 1956

†Miomatoneurites sylvaensis Zalessky 1956

G. †Neodelopterum Rasnitsyn 2004

†Neodelopterum anomalum Rasnitsyn 2004

G. †Palaeomantina Rasnitsyn 1977

†Palaeomantina pentamera Rasnitsyn 1977

G. †Palaeomantis Handlirsch 1904

†Palaeomantis aestiva Novokshonov 2000

†Palaeomantis apicalis Rasnitsyn 2004

†Palaeomantis hangardi Guthörl 1962

†Palaeomantis laeta Novokshonov and Zhuzhgova 2002

†Palaeomantis ostertali Guthörl 1962

†Palaeomantis schmidti Handlirsch 1904

Invalid names: Palaeomantis aequalis Martynov 1928 [synonym]

†Palaeomantis sylvaensis Martynov 1940

†Palaeomantis tshekardensis Novokshonov 2000

Invalid names: Stefanomioptera Guthörl 1962 [synonym]

G. †Permodelopterum Kukalová 1963

†Permodelopterum obscurum Kukalová 1963

G. †Permonia Kukalová 1963

†Permonia arta Kukalová 1963

†Permonia insolida Kukalová 1963

†Permonia permoni Kukalová 1963

G. †Permonikia Kukalová 1963

†Permonikia permoniki Kukalová 1963

G. †Perunopterum Kukalová 1963

†Perunopterum corium Kukalová 1963

†Perunopterum lepidum Kukalová 1963

†Perunopterum lumbiforme Kukalová 1963

†Perunopterum macrum Kukalová 1963

†Perunopterum mirum Kukalová 1963

†Perunopterum peruni Kukalová 1963

†Perunopterum saxeum Kukalová 1963

G. †Saaromioptera Guthörl 1963

†Saaromioptera jordani Guthörl 1963

G. †Sellardsiopsis Zalessky 1939

†Sellardsiopsis conspicua Zalessky 1939

Invalid names: Miomantisca clara Zalessky 1956 [synonym], Miomoptilon pusillus Zalessky 1956 [synonym], Palaeomantisca lata Martynov 1940 [synonym], Paramioma palida Zalessky 1956 [synonym]

Invalid names: Miomantisca Zalessky 1956 [synonym], Miomoptilon Zalessky 1956 [synonym], Palaeomantisca Martynov 1940 [synonym], Paramioma Zalessky 1956 [synonym]

G. †Stigmodelopterum Rasnitsyn 2004

†Stigmodelopterum pterostigmalis Rasnitsyn 2004

G. †Tridelopterum Rasnitsyn 2004

†Tridelopterum indebitum Rasnitsyn and Aristov 2013

†Tridelopterum variegatum Rasnitsyn 2004

Invalid names: Archisialidae Martynov 1932 [synonym], Delopteridae Sellards 1909 [synonym], Palaeomantiscidae Rasnitsyn 1977 [synonym], Stefanomiopterinae Guthörl 1962 [invalid subgroup]

G. †Controversala Brauckmann and Herd 2005

†Controversala miomopteroides Brauckmann and Herd 2005

Fm. †Dictyodipteridae Rohdendorf 1961

G. †Dictyodiptera Rohdendorf 1961

†Dictyodiptera multinervis Rohdendorf 1961

G. †Eopanorpella Schmidt 1962

†Eopanorpella ernsti Schmidt 1962

Unr. Exopterygota Sharp 1899

Or. Blattaria Latreille 1810 [cockroach]

G. †Barona Calisto and Piñeiro 2019

†Barona arcuata Calisto and Piñeiro 2019

Superfm. †Caloblattinoidea Vrsansky and Ansorge 2000

Fm. †Caloblattinidae Vršanský and Ansorge 2000

G. †Aktassoblatta Vishnyakova 1971

G. †Asioblatta Vishnyakova 1968

G. †Caloblattina Handlirsch 1906

Invalid names: Apistoblattula Bode 1953 [synonym], Macroblattina Bode 1953 [synonym], Pachyneuroblattina Handlirsch 1906 [synonym], Palmoblattina Bode 1953 [synonym], Polyphleboblatta Bode 1953 [synonym], Strebloblattula Bode 1953 [synonym]

G. †Dazhublattella Fang et al. 2013

G. †Decomposita Vršanský 2008

G. †Etapia Vishnyakova 1983

G. †Euryblattula Martynov 1937

G. †Fusiblatta Hong 1980

G. †Ijablatta Vishnyakova 1983

G. †Itchetuja Vishnyakova 1983

G. †Kemerowia Vishnyakova 1983

G. †Nuurcala Vršanský 2003

G. †Okras Vršanský 2020

G. †Paleovia Vršanský 2008

G. †Rhipidoblatta Vishnyakova 1968

G. †Rhipidoblattina Handlirsch 1906

G. †Samaroblatta Tillyard 1919

Invalid names: Striatotegmen Haughton 1924 [synonym]

G. †Samaroblattula Martynov 1937

G. †Shartegoblattina Vršanský 2004

G. †Sogdoblatta Martynov 1937

G. †Solemnia Vršanský 2008

G. †Soliblatta Lin 1986

G. †Srdiecko Vršanský 2008

G. †Taublatta Martynov 1937

G. †Taublattopsis Vishnyakova 1985

G. †Thuringoblatta Kuhn 1938

Fm. †Eadiidae Vršanský 2009

G. †Eadia Vršanský 2009

Fm. †Fuziidae Vršanský et al. 2009

G. †Arcofuzia Wei et al. 2012

G. †Colorifuzia Wei et al. 2013

G. †Fuzia Vršanský et al. 2009

G. †Longifuzia Liang et al. 2019

G. †Parvifuzia Guo and Ren 2011

Fm. †Raphidiomimidae Vishnyakova 1973

G. †Cameloblatta Vishniakova 1973

G. †Cretacechorista Hong et al. 1989

G. †Divocina Liang et al. 2012

G. †Falcatusiblatta Liang et al. 2018

G. †Fortiblatta Liang et al. 2009

G. †Graciliblatta Liang et al. 2012

G. †Liadoblattina Handlirsch 1906

Invalid names: Mesoblattopsis Handlirsch 1906 [synonym], Ptyctoblattina Bode 1953 [synonym], Trirhabdoblattina Bode 1953 [synonym]

G. †Paekthoblatta So and Won 2021

G. †Raphidiomima Vishniakova 1973

G. †Raphidiomimula Grimaldi and Ross 2004

Invalid names: Caloblattinidea Vrsansky and Ansorge 2000 [empty]

Fm. †Cratovitismidae Bechly 2007

G. †Antophiloblatta Sendi 2020

†Antophiloblatta hispida Sendi 2020

G. †Jantaropterix Vršanský and Grimaldi 2003

†Jantaropterix newjersey Vršanský and Grimaldi 2003

G. †Lepidopterix Sendi 2020

†Lepidopterix vegrandis Sendi 2020

G. †Nadveruzenie Vršanský et al. 2021

†Nadveruzenie postava Vršanský et al. 2021

G. †Perspicuus Koubová 2020

†Perspicuus pilosus Koubová 2020

†Perspicuus vrsanskyi Mlynský 2020

G. †Vzrkadlenie Vršanský 2020

†Vzrkadlenie miso Vršanský 2020

G. †Irreblatta Vršanský 2008

†Irreblatta relicta Vršanský 2008

Fm. †Lygobiidae Mitchell 1908

G. †Lygobius Mitchell 1908

†Lygobius knowltoni Mitchell 1908

Fm. Nocticolidae Bolivar 1892

G. †Crenocticola Li and Huang 2019

†Crenocticola burmanica Li and Huang 2019

†Crenocticola svadba Podstrelená 2020

G. †Mulleriblattina Sendi and Vidlička 2020

†Mulleriblattina bowangi Sendi and Vidlička 2020

G. †Petroblattina Scudder 1879

Fm. †Phyllodromiidae Burnner 1865

Subfm. †Phyllodromiinae Kirby 1904

G. †Phantocephalus Zhang 1989

Superfm. Polyphagoidea Walker 1868

Fm. †Holocompsidae Rehn 1951

Subfm. †Vitisminae Vršanský and Ansorge 2001

Fm. †Skokidae Vršanský 2007 [jumping cockroach]

G. †Skok Vršanský 2007

Fm. †Socialidae Vršanský 2010

G. †Sociala Vršanský 2010

Invalid names: Blattariae kliveri Handlirsch 1920 [nomen dubium], Blattariae reisi Handlirsch 1920 [nomen dubium], Mesoblattinoidea Handlirsch 1906 [empty]

Or. †Protelytroptera Tillyard 1931

Fm. †Apachelytridae Carpenter and Kukalová 1964

G. †Apachelytron Carpenter and Kukalová 1964

†Apachelytron transversum Carpenter and Kukalová 1964

Fm. †Archelytridae Carpenter 1933

G. †Archelytron Carpenter 1933

†Archelytron superbum Carpenter 1933

G. †Ortelytron Kukalová 1965

†Ortelytron artum Kukalová 1965

†Ortelytron europaeum Kukalová 1965

G. †Arctocoleus Martynov 1932

†Arctocoleus ivensis Martynov 1932

Fm. †Bardocoleidae Zalessky 1947

G. †Uralelytron Rohdendorf 1939

†Uralelytron insignis Zalessky 1947

†Uralelytron martynovi Rohdendorf 1939

Invalid names: Bardocoleus Zalessky 1947 [synonym]

G. †Crasselytron Kukalová 1965

†Crasselytron convexum Kukalová 1965

Fm. †Dermelytridae Kukalová 1966

G. †Chanoselytron Kukalová 1966

†Chanoselytron gingiva Kukalová 1966

G. †Dermelytron Kukalová 1966

†Dermelytron conservativum Kukalová 1966

†Dermelytron pigmentatum Kukalová 1966

G. †Psychelytron Kukalová 1966

†Psychelytron progressivum Kukalová 1966

Fm. †Elytroneuridae Carpenter 1933

G. †Elytroneura Carpenter 1933

†Elytroneura permiana Carpenter 1933

Invalid names: Blattocoleidae Zeuner 1933 [synonym]

G. †Glabelytron Kukalová 1965

†Glabelytron lativenosum Kukalová 1965

Fm. †Labidelytridae Kukalová-Peck 1987

G. †Labidelytron Kukalová-Peck 1987

†Labidelytron enervatum Kukalová 1966

Invalid names: Stenelytron Kukalová 1966 [replaced]

G. †Xenelytron Kukalová 1966

†Xenelytron ligula Kukalová 1966

Invalid names: Stenelytridae Kukalová 1966 [replaced]

Fm. †Megelytridae Carpenter 1933

G. †Megelytron Tillyard 1931

†Megelytron robustum Tillyard 1931

Fm. †Permelytridae Tillyard 1931

G. †Blattelytron Tillyard 1931

†Blattelytron permianum Tillyard 1931

G. †Parablattelytron Tillyard 1931

†Parablattelytron delicatum Tillyard 1931

†Parablattelytron elongatum Tillyard 1931

Invalid names: Acosmelytron acutum Tillyard 1931 [synonym]

†Parablattelytron latum Carpenter 1933

†Parablattelytron rectum Tillyard 1931

†Parablattelytron subincisum Tillyard 1931

Invalid names: Acosmelytron Tillyard 1931 [synonym]

G. †Permelytron Tillyard 1931

†Permelytron schucherti Tillyard 1931

Invalid names: Acosmelytridae Tillyard 1931 [synonym], Blattelytridae Tillyard 1931 [synonym]

Fm. †Permophilidae Tillyard 1924

G. †Elytrathrix Kukalová 1966

†Elytrathrix hirsuta Kukalová 1966

G. †Permophilus Tillyard 1924

†Permophilus capulus Kukalová 1966

†Permophilus hirtus Kukalová 1966

†Permophilus minor Tillyard 1924

†Permophilus pincombei Tillyard 1924

Fm. †Planelytridae Kukalová 1965

G. †Planelytron Kukalová 1965

†Planelytron planum Kukalová 1965

Fm. †Protelytridae Tillyard 1931

G. †Blanzelytron Langiaux and Parriat 1975

†Blanzelytron vetteri Langiaux and Parriat 1975

G. †Permelytropsis Carpenter 1933

†Permelytropsis cubitalis Carpenter 1933

G. †Protelytron Tillyard 1931

†Protelytron angustum Carpenter 1933

†Protelytron dunbari Tillyard 1931

†Protelytron furcatum Carpenter 1939

†Protelytron permianum Tillyard 1931

Invalid names: Protelytron tillyardi Carpenter 1933 [synonym]

G. †Protelytropsis Tillyard 1931

†Protelytropsis grandis Tillyard 1931

Fm. †Protocoleidae Tillyard 1924

G. †Austrelytron Kukalová 1966

†Austrelytron tillyardi Kukalová 1966

G. †Phyllelytron Kukalová 1966

†Phyllelytron acuminatum Riek 1976

†Phyllelytron folium Kukalová 1966

†Phyllelytron granulatum Kukalová 1966

†Phyllelytron melinum Kukalová 1966

†Phyllelytron petalon Kukalová 1966

G. †Protocoleus Tillyard 1924

†Protocoleus mitchelli Tillyard 1924

G. †Rugelytron Kukalová 1965

†Rugelytron fuscum Kukalová 1965

G. †Venelytron Kukalová 1965

†Venelytron tuberculatum Kukalová 1965

Fm. †Gelasopteridae Carpenter 1976

G. †Gelasopteron Carpenter 1976

†Gelasopteron gracile Carpenter 1976

Or. †Glosselytrodea Martynov 1938

Fm. †Archoglossopteridae Martynova 1958

G. †Archoglossopterum Martynova 1958

†Archoglossopterum shoricum Martynova 1958

Fm. †Glosselytridae Martynov 1938

G. †Glosselytron Martynov 1938

†Glosselytron multivenosum Martynov 1938

G. †Karajurina Vilesov and Novokshonov 1994

†Karajurina desperata Rasnitsyn and Aristov 2013

†Karajurina unica Vilesov and Novokshonov 1994

G. †Mongolojurina Ponomarenko 1988

†Mongolojurina altaica Ponomarenko 1988

†Mongolojurina linguale Ponomarenko 2000

†Mongolojurina martynovae Ponomarenko 2000

G. †Sinojurina Huang et al. 2007

†Sinojurina permiana Huang et al. 2007

G. †Uskatelytrum Martynova 1952

†Uskatelytrum sibiricum Martynova 1952

Invalid names: Uskatelytridae Martynova 1952 [synonym]

Fm. †Glossopteridae Sharov 1966

G. †Glossopterum Sharov 1966

†Glossopterum martynovae Sharov 1966

†Glossopterum sharovi Novokshonov 1998

Fm. †Jurinidae Zalessky 1929

G. †Eoglosselytrum Martynova 1952

†Eoglosselytrum anomale Martynova 1961

†Eoglosselytrum biarmicum Rasnitsyn and Aristov 2013

†Eoglosselytrum kaltanicum Martynova 1952

†Eoglosselytrum kondomense Martynova 1952

†Eoglosselytrum kultshumovense Rasnitsyn and Aristov 2013

†Eoglosselytrum kuznetzkiense Martynova 1952

†Eoglosselytrum perfectum Martynova 1961

†Eoglosselytrum perplexum Riek 1953

†Eoglosselytrum pictum Martynova 1961

†Eoglosselytrum sharovi Martynova 1961

†Eoglosselytrum usinskiense Martynova 1961

†Eoglosselytrum zalesskyi Vilesov and Novokshonov 1994

Invalid names: Permoberothella Riek 1953 [synonym]

G. †Isadelytron Rasnitsyn and Aristov 2013

†Isadelytron planum Rasnitsyn and Aristov 2013

†Isadelytron speciosum Rasnitsyn and Aristov 2013

G. †Jurina Zalessky 1929

†Jurina marginata Martynov 1938

†Jurina reducta Martynov 1938

Invalid names: Anorthoneura insignis Martynov 1938 [synonym]

†Jurina scutulata Zalessky 1929

†Jurina sojanensis Martynov 1938

Invalid names: Anorthoneura asymmetrica Martynov 1938 [synonym]

Invalid names: Anorthoneura Martynov 1938 [synonym]

G. †Ladinoglosselytron Hong 2007

†Ladinoglosselytron martynovae Hong 2007

G. †Mesojurina Martynova 1943

†Mesojurina sogjutensis Martynova 1943

G. †Protojurina Martynova 1958

†Protojurina cellulosa Martynova 1958

†Protojurina duplex Martynova 1961

†Protojurina juchnevskae Martynova 1961

†Protojurina triramosa Martynova 1961

G. †Shaanxiglosselytron Hong 2007

†Shaanxiglosselytron longata Hong 2007

G. †Sinoglosselytron Hong 2007

†Sinoglosselytron qishuiheensis Hong 2007

G. †Surijoka Martynova 1958

†Surijoka grandicella Martynova 1958

†Surijoka lutevensis Béthoux et al. 2001

Invalid names: Anorthoneuridae Martynov 1938 [synonym], Eoglosselytridae Martynova 1952 [synonym]

Fm. †Permoberothidae Tillyard 1932

G. †Moscheloptera Pérez-de la Fuente et al. 2022

†Moscheloptera phantasma Pérez-de la Fuente et al. 2022

G. †Permoberotha Tillyard 1932

†Permoberotha villosa Tillyard 1932

Invalid names: Dictyobiella nervosa Tillyard 1937 [synonym], Permoberotha convergens Tillyard 1937 [synonym]

Invalid names: Dictyobiella Tillyard 1937 [synonym]

G. †Sylvaelytron Novokshonov 1998

†Sylvaelytron latipennatum Novokshonov 1998

Invalid names: Permoberothinae Tillyard 1937 [empty]

Fm. †Polycytellidae Tillyard 1922

G. †Argentinoglosselytrina Martins-Neto and Gallego 2001

†Argentinoglosselytrina pulchella Martins-Neto and Gallego 2001

G. †Moltenojurina Béthoux and Anderson 2021

†Moltenojurina parva Béthoux and Anderson 2021

G. †Polycytella Tillyard 1922

†Polycytella rasnitsyni Béthoux and Anderson 2021

†Polycytella triassica Tillyard 1922

Invalid names: Jurinida [synonym]

Unr. Holometabola

Unr. Aparaglossata Peters et al. 2014

Unr. Mecopterida

Superor. Amphiesmenoptera Kiriakoff 1948

G. †Aphryganoneura Tillyard 1926

Unr. Endymenoptera Engel 2022

Fm. †Eocoronidae Tindale 1980

G. †Liadoptilia Handlirsch 1939

Or. †Protomeroptera Engel 2022

Fm. †Stereochoristidae Tillyard 1919

G. †Trichopteridium Geinitz 1880

Invalid names: Necrotaulius handlirschi Fischer 1962 [nomen dubium], Necrotaulius regularis Handlirsch 1939 [nomen dubium], Necrotaulius vicinus Handlirsch 1939 [nomen dubium], Papilionidea Laicharting 1781 [synonym]

Superor. Antliophora Hennig 1969

Or. Diptera Linnaeus 1758 [fly]

Invalid names: Antliata Fabricius 1775 [synonym], Halterata Scopoli 1763 [synonym], Nakridletia Vršanský et al. 2010 [synonym]

Or. Mecoptera Packard 1886 [scorpionfly]

Invalid names: Panorpata Brauer [synonym]

Or. Siphonaptera Latreille 1825 [flea]

G. †Curvilarva Novokshonov 2004

†Curvilarva tshekardensis Novokshonov 2004

Or. †Permotrichoptera Martynova 1958

Fm. †Microptysmatidae Martynova 1958

Invalid names: Uraloptysmatidae Ivanov 1992 [synonym]

Unr. †Neuropteroidea

Unr. Coleopterida Kukalová-Peck and Lawrence 2004

Or. Coleoptera Linnaeus 1758 [beetle]

Invalid names: Eleuterata Fabricius 1775 [synonym]

Or. †Skleroptera Kirejtshuk and Nel 2013

Or. Strepsiptera Kirby 1813 [twisted-winged parasite]

Superor. Neuropterida

Unr. Eidoneuroptera Engel et al. 2018

Fm. †Permithonidae Tillyard 1922

Invalid names: Palaemerobiidae Martynov 1928 [synonym], Permegalomidae Martynova 1952 [synonym], Permopsychopsidae Riek 1953 [synonym], Permosisyridae Martynov 1932 [synonym], Sialidopsidae Zalessky 1928 [synonym]

Subor. †Priscaenigmatomorpha Engel 2002

Or. Raphidioptera Handlirsch 1908 [snakefly]

Fm. †Avioxyelidae Nel et al. 2013

G. †Avioxyela Nel et al. 2013

†Avioxyela gallica Nel et al. 2013

G. †Cavalarva Aristov and Rasnitsyn 2015

†Cavalarva caudata Aristov and Rasnitsyn 2015

Or. Hymenoptera Linnaeus 1758

Unr. Aculeati Latreille 1802

Unr. Nemoglossata Latreille 1802

Fm. Apiariae Latreille 1802

Unr. Platiglossata Latreille 1802

Invalid names: Andrenetae Latreille 1802 [empty], Crabronites Latreille 1802 [empty], Formicariae Latreille 1802 [empty], Mutillariae Latreille 1802 [empty], Philantores Latreille 1802 [empty], Scolietae Latreille 1802 [empty], Sphegimae Latreille 1802 [empty]

Superfm. Anaxyeloidea Martynov 1925

Fm. Anaxyelidae Martynov 1925

Subfm. †Anaxyelinae Martynov 1925

Invalid names: Dolichostigmatinae Rasnitsyn 1968 [synonym], Kempendajinae Rasnitsyn 1980 [synonym]

Subfm. Syntexinae Benson 1935

Invalid names: Paraxiphydriinae Gao et al. 2022 [synonym]

Fm. †Bryopompilidae Engel and Grimaldi 2006

Invalid names: Bryopompilidae Rodriguez et al. 2016 [objective synonym]

G. †Nygmatius Hong 1983

†Nygmatius beipiaoensis Hong 1983

Superfm. Pamphilioidea Cameron 1890 [sawfly]

Fm. Megalodontesidae Konow 1897

Subfm. †Archoxyelydinae Wang et al. 2013

Subfm. †Decorisiricinae Wang et al. 2015

G. †Jibaissodes Ren 1995

Subfm. Megalodontesinae Konow 1897

Invalid names: Rudisiriciinae Gao et al. 2010 [synonym]

Subfm. †Praesiricinae Rasnitsyn 1968

Invalid names: Praesiricidae Rasnitsyn 1968 [synonym]

Fm. †Mirolydidae Wang et al. 2017

G. †Mirolyda Wang et al. 2017

Fm. Pamphiliidae Cameron 1890

Subfm. Cephalciinae Benson 1945

G. Cephaleia Jurine 1801

Subfm. Juralydinae Rasnitsyn 1977

Subfm. Pamphiliinae Cameron 1890

G. †Ulteramus Archibald and Rasnitsyn 2015

Invalid names: Lydites Newman 1834 [replaced]

G. †Prolyda Rasnitsyn 1968

†Prolyda depressa Rasnitsyn 1969

†Prolyda dimidia Wang et al. 2016

†Prolyda elegantula Wang et al. 2016

†Prolyda karatavica Rasnitsyn 1968

†Prolyda xyelocera Rasnitsyn 1968

Fm. †Xyelydidae Rasnitsyn 1968

G. †Brevilyda Wang and Rasnitsyn 2016

G. †Chifengilyda Zheng and Chen 2018

G. †Ferganolyda Rasnitsyn 1983

G. †Fissilyda Wang et al. 2015

G. †Medilyda Wang and Rasnitsyn 2016

G. †Mesolyda Rasnitsyn 1963

Invalid names: Alloxyelula Ren 1995 [synonym]

G. †Novalyda Gao et al. 2013

G. †Rectilyda Wang et al. 2014

G. †Sagulyda Rasnitsyn 1983

G. †Sinoprolyda Hong 1987

G. †Sogutia Rasnitsyn 1977

G. †Strenolyda Wang and Rasnitsyn 2016

G. †Strophandria Rasnitsyn 1968

G. †Xyelyda Rasnitsyn 1968

Invalid names: Megalodontoidea Konow 1897 [synonym]

Superfm. Tenthredinoidea Latreille 1802 [wood wasp]

Fm. Argidae Konow 1890 [argid sawfly]

Subfm. Arginae Konow 1890

Subfm. Sterictiphorinae Rohwer 1911

G. †Atefia Krogmann et al. 2013

†Atefia rasnitsyni Krogmann et al. 2013

Fm. Athaliidae Rohwer 1911

G. Athalia Leach 1817

Fm. Blasticotomidae Thomson 1871

G. Blasticotoma Klug 1834

G. †Paremphytus Brues 1908

G. Runaria Malaise 1931

G. †Zippelia Rasnitsyn and Müller 2023

Invalid names: Xyelotomidae Rasnitsyn 1968 [synonym]

Fm. Cimbicidae Kirby 1837 [cimbicid sawfly]

Subfm. Abiinae Thomson 1871

Subfm. Cimbicinae Kirby 1837

Subfm. Corynidinae Benson 1938

G. †Sinocimbex Zhang et al. 1994

G. †Trichiosomites Brues 1908

Invalid names: Pachylostictinae Benson 1938 [empty]

Fm. Diprionidae Rohwer 1910 [conifer sawfly]

Subfm. Diprioninae Rohwer 1910

G. †Eodiprion Schedl 2007

G. †Paleomonoctenus Nel 2004

Fm. †Electrotomidae Rasnitsyn 1977

G. †Electrotoma Rasnitsyn 1977

G. †Manevalia Piton 1940

†Manevalia pachyliformis Piton 1940

Fm. Pergidae Ashmead 1898

Subfm. Perginae Ashmead 1898

Subfm. Perreyiinae Cameron 1883

Invalid names: Euryinae Konow 1898 [synonym]

Invalid names: Syzygoniinae Cameron 1883 [empty]

Fm. Tenthredinidae Latreille 1802

Subfm. Allantinae Newman 1834

Subfm. Blennocampinae Konow 1890

G. †Emprionopsis Zhang et al. 1994

Subfm. Heterarthrinae Benson 1952

G. †Leptocampus Zhang et al. 1994

Subfm. Nematinae

G. †Phthonocampus Zhang et al. 1994

G. Rhogogaster Konow 1884

Subfm. Selandriinae Thomson 1871

Subfm. Tenthredininae Latreille 1802

Invalid names: Nematus ellipticus Fritsch 1901 [nomen dubium]

Invalid names: Pteronus prodigus Brues 1908 [nomen dubium]

Unr. Terebrantes Latreille 1802

Unr. Ventropedonculati Latreille 1802

Fm. Ichneumonides Latreille 1802 [ichneumon wasp]

Invalid names: Chrysidides Latreille 1802 [empty], Cleptiosa Latreille 1802 [empty], Cynipsera Latreille 1802 [empty], Evaniales Latreille 1802 [empty]

Unr. Ventrosessiles Latreille 1802

Invalid names: Tenthredines Latreille 1802 [empty], Urocerata Latreille 1802 [empty]

Unr. Unicalcarida Schulmeister et al. 2002

Subor. Apocrita Gerstaecker 1867 [wasp]

Unr. Aculeata Latreille 1802

G. †Apocritites Rasnitsyn and Jarzembowski 1998

Fm. †Armaniidae Dlussky 1983

Infraor. Brachycleistogastromorpha Kovalev 1994

Superfm. Ceraphronoidea Haliday 1833

Superfm. Evanioidea Latreille 1802

Superfm. Ichneumonoidea Latreille 1802

Fm. †Khutelchalcididae Rasnitsyn et al. 2004

Fm. †Kuafuidae Rasnitsyn and Zhang 2010

Fm. †Limnetidae Hong 1983

G. †Longhuaia Hong 1984

Superfm. Megalyroidea Schletterer 1889

Superfm. †Panguoidea Li et al. 2019

Infraor. Proctotrupomorpha

Invalid names: Cynipomorpha Kovalev 1994 [synonym]

Superfm. Stephanoidea Leach 1815

Invalid names: Ephialtitoidea Handlirsch 1906 [synonym]

Superfm. Trigonalyoidea Cresson 1887

Invalid names: Apiaria lapidea Germar 1842 [nomen dubium], Asiacynipites Hong 2002 [nomen nudum], Dendrocerus dubitatus Brues 1937 [nomen dubium], Eucynipites Hong 2002 [nomen nudum], Evaniomorpha Rasnitsyn 1988 [invalid subgroup], Stephanomorpha Leach 1815 [invalid subgroup]

Superfm. Cephoidea Newman 1834 [stem sawfly]

Fm. Cephidae Newman 1834 [stem sawfly]

Fm. †Sepulcidae Rasnitsyn 1968

G. †Cratoenigma Krogmann and Nel 2012

†Cratoenigma articulata Krogmann and Nel 2012

Superfm. Orussoidea Newman 1834 [wood wasp]

Fm. †Burmorussidae Zhang et al. 2020

Fm. Orussidae Newman 1834 [parasitic wood wasp]

Fm. †Paroryssidae Martynov 1925

Invalid names: Karatavitoidea Rasnitsyn 1963 [synonym]

Superfm. Siricoidea Billberg 1820

Fm. †Protosiricidae Rasnitsyn and Zhang 2004

Fm. †Pseudosiricidae Handlirsch 1906

Invalid names: Myrmiciidae Maa 1949 [synonym]

G. †Sinanxyela Hong 1982

Fm. †Sinosiricidae Hong 1975

Fm. Siricidae Billberg 1820 [horntail]

Subor. Vespina Laicharting 1781

Invalid names: Euhymenoptera [synonym]

Superfm. Xiphydrioidea Leach 1819

Invalid names: Xiphydriidae Leach 1819 [empty]

Invalid names: Orussomorpha Rasnitsyn 1980 [invalid subgroup]

Superfm. Xyeloidea Newman 1834 [sawfly]

Fm. †Syspastoxyelidae Engel and Huang 2016

G. †Burmoxyela Zheng and Rasnitsyn 2021

G. †Deltoxyela Wang et al. 2021

G. †Grandixyela Zheng 2021

G. †Pinguixyela Zheng and Rasnitsyn 2021

G. †Striaexyela Zheng 2019

G. †Syspastoxyela Engel and Huang 2016

Fm. Xyelidae Newman 1834 [sawfly]

Subfm. †Archexyelinae Rasnitsyn 1964

Subfm. †Daohugoinae Rasnitsyn and Zhang 2004

Subfm. Macroxyelinae Ashmead 1898

Subfm. †Madygellinae Rasnitsyn 1969

Subfm. Xyelinae Newman 1834

Invalid names: Aculeata Scopoli 1763 [synonym], Cleptes steenstrupi Beck 1836 [nomen nudum], Piezata Fabricius 1798 [synonym], Pseudisobrachium oligocenicum Theobald 1937 [nomen dubium], Shandongodes necrodes Zhang 1985 [nomen dubium], Siricomorpha Rasnitsyn 1980 [invalid subgroup], Symphyta Gerstaecker 1867 [invalid subgroup]

G. †Metabolarva Kirejtshuk et al. 2013

†Metabolarva bella Kirejtshuk et al. 2013

Fm. †Srokalarvidae Haug et al. 2016

G. †Srokalarva Haug et al. 2016

†Srokalarva berthei Haug et al. 2016

Invalid names: Endopterygota Sharp 1899 [synonym], Panorpida [empty]

Fm. †Mesophthiridae Gao et al. 2019

G. †Mesophthirus Gao et al. 2019

†Mesophthirus engeli Gao et al. 2019

Fm. †Metropatoridae Handlirsch 1906

G. †Metropator Handlirsch 1906

†Metropator pusillus Handlirsch 1906

G. †Metropatorites Keller 1934

†Metropatorites kassenbergensis Keller 1934

Fm. †Micropalentomidae Schmidt 1962

G. †Micropalentomum Schmidt 1962

†Micropalentomum minusculum Schmidt 1962

Or. †Mixotermitoidea Handlirsch 1906

Fm. †Kliveriidae Handlirsch 1919

G. †Kliveria Handlirsch 1906

†Kliveria incerta Kliver 1886

Fm. †Mixotermitidae Handlirsch 1906

G. †Mixotermes Sterzel 1881

†Mixotermes lugauensis Sterzel 1881

G. †Oiophassus Zhang 1989

†Oiophassus nycterus Zhang 1989

G. †Palaeoheteroptera Meunier 1900

†Palaeoheteroptera carinata Meunier 1900

†Palaeoheteroptera lapidaria Weyenbergh 1869

G. †Palaeonepidoideus Meunier 1900

Fm. †Perielytridae Zalessky 1948

G. †Perielytron Zalessky 1943

†Perielytron mirabile Zalessky 1943

Fm. †Permembiidae Tillyard 1937

G. †Kungurmica Novokshonov 1998

†Kungurmica tshekardensis Novokshonov 1998

Subfm. †Letopalopterinae Martynova 1961

G. †Issapaloptera Rasnitsyn and Aristov 2013

†Issapaloptera infracta Rasnitsyn and Aristov 2013

G. †Letopaloptera Martynova 1961

†Letopaloptera albardiana Martynova 1961

†Letopaloptera carpenteriana Martynova 1961

G. †Neembia Rasnitsyn and Aristov 2013

†Neembia ampla Rasnitsyn and Aristov 2013

G. †Permindigena Novokshonov 1998

†Permindigena lientericus Novokshonov 1998

G. †Permembia Tillyard 1928

†Permembia delicatula Tillyard 1928

Fm. †Permosialidae Martynov 1928

G. †Epimastax Martynov 1928

†Epimastax celer Kukalová 1965

†Epimastax hesterae Rasnitsyn and van Dijk 2011

†Epimastax mutovinensis Rasnitsyn and Aristov 2013

†Epimastax parvulus Martynov 1928

†Epimastax sojanensis Rasnitsyn 1977

†Epimastax tshepanikha Rasnitsyn and Aristov 2013

Invalid names: Sindomioptera Rasnitsyn 1977 [synonym]

G. †Palaeomantopsis Martynov 1928

†Palaeomantopsis furcatella Martynov 1928

Invalid names: Permosialis quadriramosa Martynova 1952 [synonym]

G. †Permosialis Martynov 1928

†Permosialis asiatica Martynova 1961

†Permosialis bayda Aristov and Rasnitsyn 2022

†Permosialis belebei Aristov and Rasnitsyn 2022

†Permosialis belmontensis Riek 1968

†Permosialis frivola Storozhenko 1991

†Permosialis lata Martynov 1928

†Permosialis latiformis Martynova 1952

†Permosialis martynovae Aristov and Rasnitsyn 2023

†Permosialis matutina Martynova 1961

†Permosialis mongolica Storozhenko 1992

†Permosialis paucinervis Martynov 1928

Invalid names: Permosialis bifasciata Martynov 1932 [synonym], Permosialis brevifurcata Martynova 1952 [synonym], Permosialis cauleoides Martynova 1952 [synonym], Permosialis defurcata Martynova 1952 [synonym], Permosialis fasciata Martynova 1952 [synonym], Permosialis immaculata Martynova 1952 [synonym], Permosialis marmorata Martynova 1952 [synonym]

†Permosialis perfecta Martynova 1952

†Permosialis sibirica Martynova 1961

†Permosialis triassica Novokshonov and Zhuzhgova 2004

†Permosialis ualentovae Novokshonov and Zhuzhgova 2004

†Permosialis udmurtensis Rasnitsyn and Aristov 2013

†Permosialis vizzya Aristov and Rasnitsyn 2023

†Permosialis zavialovensis Rasnitsyn and Aristov 2013

Invalid names: Sarbalopterodes Storozhenko 1991 [synonym], Tologoptera Storozhenko 1992 [synonym]

Invalid names: Epimastacidae Martynov 1928 [synonym], Perloblattidae Storozhenko 1992 [synonym], Permonkidae Rasnitsyn 1977 [synonym], Tologopteridae Storozhenko 1992 [synonym]

Unr. Polyneoptera Martynov 1938

Unr. Anartioptera Engel and Grimaldi 2004

Unr. Polyorthoptera Engel and Grimaldi 2004

Superor. Orthopterida Boudreaux 1979

Fm. †Cheliphlebiidae Handlirsch 1906

Superor. Archaeorthoptera Béthoux and Nel 2002

G. †Aviologus Coty et al. 2014

†Aviologus duquesnei Coty et al. 2014

G. †Bethouxia Prokop et al. 2015

†Bethouxia ariegensis Prokop et al. 2015

Fm. †Chresmodidae Haase 1890

G. †Chresmoda Germar 1839

†Chresmoda aquatica Martínez-Delclòs 1989

†Chresmoda chikuni Zhang and Ge 2017

†Chresmoda libanica Nel et al. 2004

†Chresmoda mongolica Ponomarenko 1986

†Chresmoda multinervis Zhang et al. 2009

†Chresmoda neotropica Engel and Heads 2008

†Chresmoda obscura Germar 1839

Invalid names: Halometra gigantea Germar 1839 [synonym], Halometra minor Oppenheim 1888 [synonym], Propygolampis bronni Weyenbergh 1874 [synonym], Pygolampis gigantea Germar 1839 [synonym], Sternarthron zitteli Haase 1890 [synonym]

†Chresmoda orientalis Esaki 1949

†Chresmoda oweni Westwood 1854

†Chresmoda shihi Zhang et al. 2009

Invalid names: Gryllidium Westwood 1854 [synonym], Halometra Oppenheim 1888 [synonym], Propygolampis Weyenbergh 1874 [synonym], Saurophthiroides Ponomarenko 1986 [synonym], Sternarthron Haase 1890 [synonym]

G. †Jurachresmoda Zhang et al. 2008

†Jurachresmoda gaskelli Zhang et al. 2008

†Jurachresmoda sanyica Zhang et al. 2009

G. †Sinochresmoda Zhang et al. 2008

†Sinochresmoda magnicornia Zhang et al. 2008

Or. †Cnemidolestodea Handlirsch 1937

G. †Aviocladus Prokop et al. 2014

†Aviocladus pectinatus Prokop et al. 2014

G. †Aviohapaloptera Prokop et al. 2014

†Aviohapaloptera bethouxi Prokop et al. 2014

Subor. †Cnemidolestina Handlirsch 1937

Fm. †Cnemidolestidae Handlirsch 1906

Invalid names: Aetophlebiidae Handlirsch 1906 [synonym], Narkemidae Handlirsch 1911 [synonym], Narkeminidae Storozhenko 1996 [synonym]

Fm. †Ctenoptilidae Aristov 2014

Fm. †Cymenophlebiidae Pruvost 1919

Fm. †Gerapompidae Handlirsch 1906

Fm. †Issatermidae Aristov 2018

Fm. †Neraphidiidae Aristov 2014

Fm. †Prygidae Aristov and Rasnitsyn 2014

Fm. †Psoropteridae Carpenter 1976

Fm. †Spanioderidae Handlirsch 1906

Invalid names: Anthraconeuridae Laurentiaux and Laurentiaux-Vieira 1980 [synonym], Carpenteropteridae Pinto and Ornellas 1991 [synonym]

Fm. †Sylvabestiidae Aristov 2000

Fm. †Tillyardembiidae Zalessky 1938

Invalid names: Heteroptilidae Carpenter 1976 [synonym], Permocapniidae Martynov 1940 [synonym]

Subor. †Parmapterina Aristov 2016

Fm. †Necrophasmidae Martynov 1925

Invalid names: Juraperlidae Huang and Nel 2007 [synonym]

Fm. †Parmapteridae Aristov and Rasnitsyn 2015

Fm. †Pinideliidae Storozhenko 1997

Fm. †Protembiidae Tillyard 1937

Invalid names: Sojanoraphidiidae Martynova 1952 [synonym], Sylvardembiidae Novokshonov 2000 [synonym], Telactinopterygidae Carpenter 1943 [synonym], Tshekardominidae Novokshonov and Aristov 2002 [synonym]

Fm. †Roqueiidae Nel et al. 2022

G. †Roqueia Nel et al. 2022

Fm. †Tococladidae Carpenter 1966

G. †Elmocladus Rasnitsyn and Aristov 2021

G. †Tococladus Carpenter 1966

G. †Tshekardocladus Aristov and Rasnitsyn 2015

G. †Xixia Gu et al. 2014

†Xixia huban Gu et al. 2014

G. †Contracladus Dvořák et al. 2021

†Contracladus impar Dvořák et al. 2021

G. †Coselia Bolton 1922

†Coselia palmiformis Bolton 1922

Fm. †Emphylopteridae Handlirsch 1922

G. †Emphyloptera Pruvost 1919

†Emphyloptera lecrivaini Pruvost 1919

Fm. †Eoblattidae Handlirsch 1906

G. †Anegertus Handlirsch 1911

†Anegertus cubitalis Handlirsch 1911

G. †Beloatta Nel et al. 2020

†Beloatta duquesnei Nel et al. 2020

G. †Eoblatta Handlirsch 1906

†Eoblatta robusta Brongniart 1894

†Eoblatta zeilleri Langiaux and Parriat 1974

G. †Lobeatta Béthoux 2005

†Lobeatta schneideri Béthoux 2005

G. †Lomovatka Aristov 2015

†Lomovatka udovichenkoi Aristov 2015

G. †Forfexala Béthoux and Herd 2009

†Forfexala kiarae Béthoux and Herd 2009

Fm. †Ischnoneuridae Handlirsch 1906

G. †Ischnoneura Brongniart 1894

†Ischnoneura brongniarti Lameere 1917

†Ischnoneura delicatula Brongniart 1894

†Ischnoneura oustaleti Brongniart 1885

Invalid names: Ischnoneurona Handlirsch 1919 [synonym], Leptoneura Brongniart 1885 [nomen nudum]

G. †Permarrhaphus Martynov 1931

†Permarrhaphus venosus Cockerell 1927

Invalid names: Adiphlebiidae Handlirsch 1906 [synonym], Stenoneurellidae Handlirsch 1919 [synonym]

G. †Lodevolongzhua Prokop et al. 2015

†Lodevolongzhua incompleta Prokop et al. 2015

Fm. †Nugonioneuridae Carpenter 1976

G. †Avionugonioneura Nel and Roques 2021

†Avionugonioneura jouaulti Nel and Roques 2021

G. †Duquesnia Nel and Roques 2021

†Duquesnia gallica Nel and Roques 2021

G. †Nugonioneura Tillyard 1937

†Nugonioneura problematica Tillyard 1937

Fm. †Omaliaidae Handlirsch 1906

G. †Anthrakoris Richardson 1956

†Anthrakoris aetherius Richardson 1956

G. †Omalia Van Beneden and Coemans 1867

†Omalia macroptera Van Beneden and Coemans 1867

Invalid names: Coseliidae Bolton 1922 [synonym]

G. †Omaliella Béthoux and Nel 2005

†Omaliella polonica Dvořák et al. 2019

†Omaliella ramosa Béthoux and Nel 2005

G. †Palaeomastax Handlirsch 1904

†Palaeomastax carbonis Handlirsch 1904

G. †Paleoischnoptera Schubnel et al. 2019

†Paleoischnoptera diaphanes Béthoux and Nel 2005

Invalid names: Ischnoptera Béthoux and Nel 2005 [replaced]

Unr. Panorthoptera Crampton 1928

G. †Bruaylogus Coty et al. 2014

†Bruaylogus magnificus Coty et al. 2014

Fm. †Cacurgidae Handlirsch 1911

G. †Cacurgus Handlirsch 1911

G. †Heterologopsis Brauckmann and Koch 1982

G. †Spilomastax Handlirsch 1911

Or. †Caloneurodea Martynov 1938

Fm. †Apsidoneuridae Carpenter 1961

G. †Aviogramma Prokop et al. 2014

Fm. †Caloneuridae Handlirsch 1906

Invalid names: Amboneuridae Carpenter 1980 [synonym], Anomalogrammatidae Carpenter 1943 [synonym], Eohymenidae Martynov 1937 [synonym], Euthygrammatidae Martynov 1928 [synonym], Paleuthygrammatidae Carpenter 1943 [synonym], Plesiogrammatidae Carpenter 1943 [synonym], Sthenaroceridae Handlirsch 1906 [synonym]

G. †Gallogramma Garrouste et al. 2017

G. †Gigagramma Béthoux et al. 2004

Fm. †Hapalopteridae Handlirsch 1906

Invalid names: Aenigmatodidae Handlirsch 1906 [synonym], Protokollariidae Handlirsch 1906 [synonym]

G. †Jarmiloptera Prokop et al. 2015

Fm. †Lodevogrammatidae Nel et al. 2023

G. †Lusitaneura Loureiro et al. 2010

G. †Nanogramma Béthoux et al. 2004

G. †Paleuthygramma Martynov 1930

G. †Paraspiculum Dvořák et al. 2021

Fm. †Permobiellidae Tillyard 1937

G. †Pruvostiella Handlirsch 1922

Invalid names: Euthyneura Pruvost 1919 [replaced]

Invalid names: Hapalopteroidea Handlirsch 1906 [synonym], Pleisiogramma kemerovoense Martynova 1952 [nomen dubium], Stenaroceridae [empty]

Fm. †Geraridae Scudder 1885

G. †Anepitedius Handlirsch 1911

G. †Axiologus Handlirsch 1906

G. †Cantabrala Kukalová-Peck and Brauckmann 1992

G. †Genentomum Scudder 1885

G. †Gerarulus Handlirsch 1911

G. †Gerarus Scudder 1885

Invalid names: Archaeacridites Meunier 1909 [synonym], Genopteryx Scudder 1885 [synonym], Rossites Richardson 1956 [synonym], Sthenaropoda Brongniart 1885 [synonym]

G. †Hirsutgerarus Zessin 2012

G. †Nacekomia Richardson 1956

G. †Osnogerarus Kukalová-Peck and Brauckmann 1992

G. †Owadpteron Dvořák et al. 2019

G. †Parapalaeomastax Dvořák et al. 2019

G. †Ploetzgerarus Zessin 2009

G. †Progenentomum Handlirsch 1906

Invalid names: Genopterygidae Richardson 1956 [synonym], Sthenaropodidae Handlirsch 1906 [synonym]

Fm. †Permostridulidae Béthoux et al. 2003

G. †Permostridulus Béthoux et al. 2003

G. †Sinogerarus Gu et al. 2017

†Sinogerarus pectinatus Gu et al. 2017

G. †Phtanomiamia Chen et al. 2020

†Phtanomiamia gui Chen et al. 2020

G. †Protodictyon Melander 1903

†Protodictyon pulchripenne Melander 1903

G. †Protomiamia Du et al. 2016

†Protomiamia yangi Du et al. 2016

G. †Sumixam Béthoux and Grossmann 2011

†Sumixam maximus Béthoux and Grossmann 2011

Invalid names: Lobeattida Béthoux 2005 [empty]

G. †Belebeus Aristov 2019

†Belebeus mutilus Aristov 2015

Invalid names: Belebey Aristov 2015 [replaced]

Unr. Eukinolabia Terry and Whiting 2005

Or. Embioptera Lameere 1900 [webspinners]

Fm. Anisembiidae Davis 1940

Subfm. Anisembiinae Davis 1940

Infraor. †Brachyphyllophagidea Rasnitsyn 2000

Fm. †Brachyphyllophagidae Rasnitsyn 2000

Fm. †Rasnalexiidae Gorochov 2021

Subor. Clothododea Engel and Grimaldi 2006

Infraor. Clothodomorpha Engel and Huang 2016

Fm. Embiidae Burmeister 1839

G. †Electroembia Ross 1956

G. Embia Latreille 1829

G. †Galloembia Falières et al. 2021

G. †Lithembia Ross 1984

Invalid names: Scelembiinae Ross 2001 [empty]

Infraor. Embiidea Lameere 1900

Superfm. †Alexarasnioidea Gorochov 2011

Subor. Neoembiodea Engel and Grimaldi 2006

Infraor. Oligotomorpha Ross 2007

Fm. Notoligotomidae Davis 1940

Subfm. †Burmitembiinae Engel and Grimaldi 2006

Fm. Scelembiidae Ross 2001

G. †Kumarembia Engel and Grimaldi 2011

Subfm. †Sorellembiinae Engel and Grimaldi 2006

Invalid names: Pachylembiidae Ross 2001 [synonym]

Fm. Teratembiidae Krauss 1911

G. Oligembia Davis 1939

Invalid names: Euembiaria Tillyard 1937 [invalid subgroup], Euembiodea Shcherbakov 2015 [invalid subgroup], Euembioptera Davis 1940 [invalid subgroup], Palembiodea Shcherbakov 2015 [invalid subgroup]

Unr. Holophasmatodea Grimaldi and Engel 2005

Fm. †Aerophasmidae Martynov 1928

Subfm. †Aerophasmatinae Martynov 1928

Subfm. †Chresmodellinae Bode 1953

Subfm. †Cretophasmatinae Sharov 1968

Fm. †Aeroplanidae Tillyard 1918

G. †Aeroplana Tillyard 1918

G. †Sharovoplana Gorochov and Maehr 2008

Invalid names: Paraplana Sharov 1968 [replaced]

Or. Phasmatodea Jacobson and Bianchi 1902 [stick insect]

Unr. Euphasmatodea Bradler 1999

Fm. †Pterophasmatidae Yang et al. 2019

G. †Raphidium Westwood 1854

Invalid names: Ela Giebel 1856 [objective synonym]

Subor. Timematodea Kevan 1982

Invalid names: Areolatae Redtenbacher 1906 [invalid subgroup], Chresmododea Martynov 1928 [empty], Necrophasmatodea Martynov 1925 [empty]

Superfm. †Prochresmodoidea Vishnyakova 1980

Fm. †Permophasmatidae Gorochov 1992

Fm. †Prochresmodidae Vishnyakova 1980

Superfm. †Susumanioidea Gorochov 1988

G. †Eoprephasma Archibald and Bradler 2015

Fm. †Susumaniidae Gorochov 1988

Superfm. †Xiphopteridea Sharov 1968

Fm. †Xiphopteridae Sharov 1968

Invalid names: Aeroplanoptera Tillyard 1918 [synonym]

Fm. †Gallophasmatidae Nel et al. 2010

G. †Gallophasma Nel et al. 2010

†Gallophasma longipalpis Nel et al. 2010

Unr. Haplocercata Terry and Whiting 2005

Or. Dermaptera De Geer 1773 [earwig]

Subor. †Archidermaptera Bey-Bienko 1936

Superfm. †Protodiplatyoidea Martynov 1925

Fm. †Turanoviidae Engel 2003

G. †Atopderma Zhao et al. 2010

†Atopderma elliptum Zhao et al. 2010

G. †Baseopsis Heer 1864

†Baseopsis forficulina Heer 1864

†Baseopsis sibirica Brauer et al. 1889

Fm. †Bellodermatidae Zhao et al. 2010

G. †Belloderma Zhao et al. 2010

G. †Chelisoficula Nel et al. 2003

†Chelisoficula caussaneli Nel et al. 2003

Subor. †Eodermaptera Engel 2003

G. †Archaeosoma Zhang 1994

Fm. †Semenoviolidae Vishnyakova 1980

Superfm. †Semenovioloidea Vishnyakova 1980

Invalid names: Turanodermidae Engel 2003 [empty]

G. †Leicarabus Hong 1983

†Leicarabus parvus Hong 1983

G. †Mesoforficula Ping 1935

†Mesoforficula sinkianensis Ping 1935

Subor. Neodermaptera Engel 2003

Infraor. Epidermaptera Engel 2003

G. †Laasbium Scudder 1900

Infraor. Protodermaptera Zacher 1910

G. †Rhadinolabis Engel et al. 2011

G. †Vendeenympha Engel and Perrichot 2014

Invalid names: Forficulina [invalid subgroup], Ocellia von Olfers 1907 [nomen dubium], Ocelliidae Erwin 1942 [invalid subgroup]

Invalid names: Labiidae [empty]

Or. Zoraptera Silvestri 1913

Fm. Zorotypidae Silvestri 1913

G. †Xenozorotypus Engel and Grimaldi 2002

G. Zorotypus Silvestri 1913

Invalid names: Paleospinosus Kaddumi 2005 [synonym]

Fm. †Hebeigrammidae Hong 2003

G. †Hebeigramma Hong 2003

†Hebeigramma divaricata Hong 1984

Invalid names: Mesogramma Hong 1984 [replaced]

Invalid names: Mesogrammatidae Hong 1984 [replaced]

Unr. Holopandictyoptera Kluge 2010

Superor. Dictyoptera Latreille 1829

Or. †Alienoptera Bai et al. 2016

Fm. †Alienopteridae Bai et al. 2016

Invalid names: Aethiocarenidae Poinar and Brown 2016 [synonym]

Fm. †Umenocoleidae Chen and Tan 1973

Invalid names: Ponopterixidae Nel et al. 2014 [synonym]

Invalid names: Aethiocarenodea Poinar and Brown 2016 [synonym], Umenocoleoidea Chen and Tan 1973 [invalid subgroup]

Fm. †Baissomantidae Gratshev and Zherikhin 1993

G. †Baissomantis Gratshev and Zherikhin 1993

G. †Labradormantis Demers-Potvin et al. 2021

Or. Blattodea Latreille 1810 [cockroach]

G. †Acriditis Germar 1842

G. †Anenev Vršanský et al. 2019

G. †Archimulacris Scudder 1868

Fm. †Archimylacridae Handlirsch 1906

G. †Balatronis Šmídová and Lei 2017

Superfm. Blaberoidea Saussure 1864

Fm. †Cainoblattinidae Ping 1932

Fm. †Compsoblattidae Schneider 1978

G. †Erucoblatta Gorochov and Anisyutkin 2007

Fm. †Gyroblattidae Durden 1969

Invalid names: Archoblattinidae Schneider 1983 [synonym], Phoberoblattidae Durden 1969 [synonym]

G. †Leptoblattina Woodward 1887

G. Morphna Shelford 1910

Fm. †Mutoviidae Vršanský and Aristov 2012

Fm. †Mylacridae Scudder 1879

Invalid names: Neorthroblattinidae Handlirsch 1906 [synonym]

Fm. †Necymylacridae Durden 1969

G. †Nehevblattella Vršanský 2004

G. †Oiratia Martynov 1930

G. †Oryctomastax Pruvost 1919

Fm. †Paucineuridae Hong 1980

Fm. †Poroblattinidae Handlirsch 1906

Invalid names: Acmaeoblattidae Durden 1969 [synonym], Diechoblattinidae Handlirsch 1906 [synonym]

G. †Prochaeradodis Piton 1940

G. †Progeotrupes Oppenheim 1888

G. †Pseudoplecta Vršanský and Anisyutkin 2008

G. †Rajharablatta Dutt 1977

Unr. Solumblattodea Evangelista and Wipfler 2019

G. †Ungoneurites Martynov 1930

Unr. Xylophagodea Engel 2011

Invalid names: Tutricablattae Wipfler and Evangelista 2019 [synonym]

Invalid names: Antemylacridae Durden 1972 [invalid subgroup], Archimylacris obovata Bolton 1911 [nomen dubium], Balatronis libanensis Sendi and Azar 2016 [nomen dubium], Blattida Latreille 1810 [synonym], Blattoidea Handlirsch 1906 [synonym], Blattoidea agilis Schlechtendal 1906 [nomen dubium], Blattoidea confusa Schlechtendal 1906 [nomen dubium], Blattoidea dictyoneura Schlechtendal 1906 [nomen dubium], Blattoidea excellens Schlechtendal 1906 [nomen dubium], Blattoidea propria Schlechtendal 1906 [nomen dubium], Blattoidea separata Schlechtendal 1906 [nomen dubium], Blattoidea singularis Schlechtendal 1906 [nomen dubium], Blattoidea venusta Schlechtendal 1906 [nomen dubium], Blattoptera Brunner 1882 [synonym], Coleopteron dubia Meunier 1895 [nomen dubium], Diaphanacridae Durden 1972 [invalid subgroup], Eotermes Haupt 1956 [nomen dubium], Etoblattina gerablattini Meunier 1921 [nomen dubium], Etoblattina inclusa Meunier 1921 [nomen dubium], Etoblattina incurvata Meunier 1921 [nomen dubium], Etoblattina mensa Meunier 1921 [nomen dubium], Haenoblattina tenuis Sellards 1908 [nomen dubium], Hemimylacris convexa Bolton 1911 [nomen dubium], Heterogamia antiqua Heer 1849 [nomen dubium], Lithomylacris kirkbyi Woodward 1887 [nomen dubium], Mesophonus maculatus Brauer et al. 1889 [nomen dubium], Oothecophora Kluge 2010 [synonym], Parallelophora Haupt 1956 [nomen dubium], Parallelophoridae Haupt 1956 [invalid subgroup], Parapanorpa Zalessky 1931 [nomen dubium], Poroblattina gratiosa Scudder 1895 [nomen dubium], Pycnoscelus gardneri Cockerell 1920 [nomen dubium], Schizoblattina minor Sellards 1908 [nomen dubium], Schizoblattina schucherti Sellards 1908 [nomen dubium], Sooblattella britannica Handlirsch 1920 [nomen dubium], Sysciophlebia uralica Zalessky 1953 [nomen dubium], Zetobora brunneri Scudder 1890 [nomen dubium]

G. †Kinklidoblatta Handlirsch 1906

†Kinklidoblatta corsini Laurentiaux 1950

†Kinklidoblatta lesquereuxi Scudder 1879

†Kinklidoblatta morini Pruvost 1912

Invalid names: Phyloblatta cuvelettei Pruvost 1912 [synonym], Phyloblatta lemayi Pruvost 1919 [synonym]

†Kinklidoblatta youhei Wei et al. 2013

†Kinklidoblatta zavjalovensis Becker-Migdisova 1961

Invalid names: Phyloblatta foveolata Becker-Migdisova 1961 [synonym]

Or. Mantodea Burmeister 1838 [mantis]

G. †Amorphoscelites Gratshev and Zherikhin 1993

G. †Aragonimantis Delclòs et al. 2016

G. †Burmantis Grimaldi 2003

Invalid names: Gryllomantis Gorochov 2006 [synonym]

G. †Chaeteessites Gratshev and Zherikhin 1993

Fm. †Cretomantidae Gratshev and Zherikhin 1993

G. †Cretophotina Gratshev and Zherikhin 1993

G. †Electromantis Gratshev and Zherikhin 1993

Fm. †Jantarimantidae Vršanský 2002

Invalid names: Archimantidae Vršanský 2002 [replaced]

G. †Jersimantis Grimaldi 1997

Fm. †Juramantidae Vršanský 2002

G. †Kazakhophotina Gratshev and Zherikhin 1993

Subor. Neomantodea Grimaldi 2003

Fm. †Santanmantidae Grimaldi 2003

G. †Vitimophotina Gratshev and Zherikhin 1993

Invalid names: Amorphoscelioidea Wang 1993 [empty], Chaeteessoidea Vickery and Kevan 1983 [empty], Gryllomantidae Gorochov 2006 [invalid subgroup], Metallyticoidea Vickery and Kevan 1983 [empty]

G. †Protoblattinopsis Laurentiaux 1953

†Protoblattinopsis stubblefieldi Laurentiaux 1953

G. †Pseudojantaropterix Gorochov 2006

†Pseudojantaropterix lebani Vršanský and Grimaldi 2003

G. †Qilianiblatta Zhang et al. 2013

†Qilianiblatta namurensis Zhang et al. 2013

G. †Raptoblatta Dittmann et al. 2015

†Raptoblatta waddingtonae Dittmann et al. 2015

G. †Sinothauma Hong and Li 2007

†Sinothauma ladinica Hong and Li 2007

Fm. †Subioblattidae Schneider 1983

G. †Potrerilloblatta Martins-Neto and Gallego 2007

G. †Samaroblattella Riek 1976

G. †Subioblatta Lin 1978

Invalid names: Architermes Haupt 1956 [nomen dubium], Blattuloidea Vishnyakova 1982 [invalid subgroup], Cryptovipositoria Kluge 2010 [synonym]

Unr. †Eoblattodea Laurentiaux 1959

Fm. †Ensiferoblattidae Li and Huang 2023

G. †Ensiferoblatta Li and Huang 2023

G. †Proceroblatta Li and Huang 2023

†Proceroblatta colossea Li and Huang 2023

Fm. †Spiloblattinidae Handlirsch 1906

G. †Ametroblatta Handlirsch 1906

G. †Arrhythmoblatta Handlirsch 1906

G. †Atactoblatta Handlirsch 1906

G. †Dicladoblatta Handlirsch 1906

G. †Doryblatta Handlirsch 1906

G. †Kinneyblatta Schneider et al. 2021

G. †Olethroblatta Handlirsch 1906

Subfm. †Permoblattininae Vishnyakova 1993

G. †Spiloblattina Scudder 1885

†Spiloblattinidae gardinerana Handlirsch 1906

†Spiloblattinidae zinkeniana Schlechtendal 1906

G. †Syscioblatta Handlirsch 1906

G. †Sysciophlebia Handlirsch 1906

Subfm. †Vivablattinae Vršanský and Mostovsky 2000

Invalid names: Kargaliinae Vishnyakova 1993 [replaced]

G. †Zavjaloblatta Becker-Migdisova 1961

Invalid names: Anthracoblattina triassica Scudder 1890 [nomen dubium], Anthracoblattina wagneri Kliver 1886 [nomen dubium], Etoblattina aperta Scudder 1895 [nomen dubium], Spiloblattina laxa Sellards 1908 [nomen dubium]

Invalid names: Mylacridina Anisyutkin 2007 [synonym], Palaeoblattariae Scudder 1879 [synonym]

G. †Ideliopsis Carpenter 1948

†Ideliopsis ovalis Carpenter 1948

G. †Kitshuga Aristov 2012

†Kitshuga ryzhkovae Aristov 2012

Fm. †Magicivenidae Yang et al. 2022

G. †Magicivena Yang et al. 2022

†Magicivena antennalis Yang et al. 2022

†Magicivena elegans Yang et al. 2022

†Magicivena sticta Yang et al. 2022

Fm. †Mantoblattidae Gorochov 2006

G. †Mantoblatta Gorochov 2006

†Mantoblatta mira Gorochov 2006

G. †Menatacridium Piton 1936

†Menatacridium eocenicum Piton 1936

Or. Notoptera Crampton 1915

Superfm. †Adeloblattoidea Durden 1969

Fm. †Adeloblattidae Durden 1969

G. †Acoblatta Cockerell 1927

G. †Adeloblatta Handlirsch 1906

Invalid names: Anthracoblattinidae Durden 1969 [empty]

Or. Orthoptera Olivier 1789 [grasshopper]

G. †Alekhosara Aristov 2008

†Alekhosara reticulata Aristov 2008

Subor. Caelifera Ander 1936

Infraor. Acrididea MacLeay 1821

Unr. Acridomorpha

Superfm. Tetrigoidea Rambur 1838

Infraor. Tridactylidea Brullé 1835

Superfm. †Dzhajloutshelloidea Gorochov 1994

Superfm. Tridactyloidea Brullé 1835

Invalid names: Ochrilidia lineata Piton 1940 [nomen dubium], Pneumoridea Uvarov 1966 [empty], Proscopiodea [synonym]

Subor. Ensifera Chopard 1920

G. †Araripemyrmecophilops Martins-Neto 1991

†Araripemyrmecophilops gracilis Martins-Neto 1991

Infraor. †Elcanidea Handlirsch 1906

Superfm. †Elcanoidea Handlirsch 1906

Superfm. †Permoraphidioidea Tillyard 1932

G. †Elcanopsis Tillyard 1918

†Elcanopsis sydneyensis Tillyard 1918

G. †Falsirameus Zhang 1985

†Falsirameus ravus Zhang 1985

Infraor. Gryllidea Laicharting 1781

Superfm. †Gryllavoidea Gorochov 1986

Superfm. Grylloidea Laicharting 1781

Superfm. Gryllotalpoidea Leach 1815

G. †Gryllobencain Haug et al. 2022

†Gryllobencain patrickmuelleri Haug et al. 2022

G. †Liassogrylloides Bode 1953

†Liassogrylloides basifastigatus Bode 1953

Infraor. †Oedischiidea Handlirsch 1906

Superfm. †Oedischioidea Handlirsch 1906

Invalid names: Tcholmanvissioidea Zalessky 1934 [synonym]

Fm. †Permotettigoniidae Nel and Garrouste 2016

G. †Rimnosentomon Zalessky 1955

Superfm. †Xenopteroidea Riek 1955

Invalid names: Triassomanteoidea [empty]

Superfm. †Phasmomimoidea Sharov 1968

Fm. †Phasmomimidae Sharov 1968

G. Terpandrus Saussure 1891

Terpandrus horridus Burmeister 1838 [Sydney gumleaf katydid]

Invalid names: Austrodictya corbouldi Tillyard 1922 [synonym]

†Terpandrus ikiensis Fujiyama 1970

Invalid names: Austrodictya Tillyard 1922 [synonym]

Fm. †Tettavidae Sharov 1968

G. †Protettavus Riek 1976

Infraor. Tettigoniidea

G. †Aenigmaraphidophora Azar et al. 2022

Superfm. Hagloidea Handlirsch 1906

Superfm. Rhaphidophoroidea Kirby 1906

Superfm. Schizodactyloidea Blanchard 1845

Superfm. Stenopelmatoidea Burmeister 1838

Invalid names: Gryllacridoidea Stal 1874 [synonym]

Superfm. Tettigonioidea Krauss 1902

G. †Tettoraptor Gorochov 2012

Invalid names: Locustariae Handlirsch 1926 [synonym]

G. †Thnetus Handlirsch 1904

†Thnetus stuckenbergi Handlirsch 1904

Fm. †Triassomanteidae Tillyard 1922

G. †Orichalcum Whalley 1985

G. †Triassomantis Tillyard 1922

Fm. †Vitimiidae Sharov 1968

G. †Deinovitimia Gorochov 1989

G. †Vitimia Sharov 1968

G. †Yuxiania Hong 1997

†Yuxiania jurassica Hong 1997

Invalid names: Acridiites Heer 1864 [nomen dubium], Acridiites liasinus Heer 1864 [nomen dubium], Kulmbachiellon fragile Kuhn 1938 [nomen dubium], Liadolocusta auscultans Handlirsch 1906 [nomen dubium], Locusta prisca Germar 1839 [nomen dubium], Locustopsites gigantea Theobald 1937 [nomen dubium]

Superfm. †Exaeretopteroidea Pierce 1944

Fm. †Exaeretopteridae Pierce 1944

G. †Exaeretoptera Pierce 1944

Fm. †Legendreiidae Huang et al. 2020

G. †Legendreia Huang et al. 2020

†Legendreia magnifica Huang et al. 2020

G. †Lithymnetes Scudder 1878

†Lithymnetes guttatus Scudder 1878

Fm. Locustidae Latreille 1802

G. †Acridomima Handlirsch 1906

†Acridomima deperditus Heer 1864

G. †Locustophanes Handlirsch 1939

†Locustophanes rhipidophorus Handlirsch 1939

G. †Loxoedischia Beckemeyer 2011

†Loxoedischia drewi Beckemeyer 2011

Fm. Mantides Latreille 1802 [praying mantis]

Invalid names: Raptoriae Latreille 1802 [empty]

G. †Nymphomorpha Henriksen 1922

†Nymphomorpha medialis Henriksen 1922

G. †Oligogetes Makarkin 1998

†Oligogetes relictum Makarkin 1998

G. †Permophyllum Prokop et al. 2015

†Permophyllum rotundatum Prokop et al. 2015

Fm. Phasgonuridae Karsch 1893

Invalid names: Phasgonurinae [empty]

G. †Pinegia Martynov 1928

†Pinegia meidigouensis Hong 1985

†Pinegia oknowae Martynov 1928

G. †Prophilaenites Bode 1953

†Prophilaenites hondelagensis Bode 1953

G. †Thnetodes Martynov 1928

†Thnetodes craticius Martynov 1928

G. †Tillyardiella Martynov 1930

†Tillyardiella distincta Martynov 1930

Subor. †Titanoptera Sharov 1968

Fm. †Deinotitanidae Gorochov 2011

G. †Deinotitan Gorochov 2007

G. †Monstrotitan Gorochov 2013

Subor. †Gerarina Scudder 1885

G. †Brachygerarus Gorochov 2004

G. †Permotitan Gorochov 2004

Subor. †Mesotitanina Tillyard 1925

Fm. †Mesotitanidae Tillyard 1925

Invalid names: Clathrotitanidae Riek 1954 [synonym]

Fm. †Paratitanidae Sharov 1968

Superfm. †Tettoedischioidea Gorochov 1987

Fm. †Tettoedischiidae Gorochov 1987

Fm. †Theiatitanidae Schubnel et al. 2021

G. †Theiatitan Schubnel et al. 2021

G. †Yinpingia Lin 1982

†Yinpingia caesia Lin 1982

Invalid names: Aboilodea Martynov 1925 [empty], Acrydiana Latreille 1802 [empty], Blattaceadup [empty], Eremobiinae [empty], Eremobiites Piton 1940 [nomen nudum], Locustoidea Handlirsch 1906 [empty], Orthophlebia longissima Giebel 1856 [nomen dubium], Saltatoria [synonym]

Or. †Paoliida Handlirsch 1906

Fm. †Anthracoptilidae Handlirsch 1922

G. †Adiphlebia Scudder 1885

†Adiphlebia lacoana Scudder 1885

Invalid names: Termes longitudinalis Scudder 1885 [synonym]

G. †Afrocladus Nel et al. 2015

†Afrocladus kenyaensis Nel et al. 2015

†Afrocladus pumilio Nel et al. 2015

G. †Anthracoptilus Lameere 1917

†Anthracoptilus perrieri Meunier 1909

Invalid names: Prostenoneura Handlirsch 1919 [objective synonym]

G. †Carrizocladus Rasnitsyn 2004

†Carrizocladus arroyo Rasnitsyn 2004

†Carrizocladus deter Rasnitsyn 2004

G. †Graticladus Novokshonov and Aristov 2004

†Graticladus apiatus Aristov and Rasnitsyn 2015

†Graticladus severus Novokshonov and Aristov 2004

G. †Homocladus Carpenter 1966

†Homocladus grandis Carpenter 1966

Invalid names: Homocladus ornatus Carpenter 1966 [synonym]

G. †Jarmilacladus Rasnitsyn and Aristov 2004

†Jarmilacladus patiens Rasnitsyn and Aristov 2013

†Jarmilacladus variabilis Rasnitsyn and Aristov 2004

G. †Lodevocladus Prokop et al. 2015

†Lodevocladus subtilis Prokop et al. 2015

G. †Mesoptilus Lameere 1917

†Mesoptilus carpenteri Guan et al. 2016

†Mesoptilus dolloi Lameere 1917

Invalid names: Stenoneurella fayoliana Handlirsch 1919 [objective synonym]

Invalid names: Stenoneurella Handlirsch 1919 [synonym]

G. †Mycteroptila Rasnitsyn 1977

†Mycteroptila armipotens Novokshonov 1998

†Mycteroptila dina Rasnitsyn 1977

G. †Paracladus Carpenter 1966

†Paracladus retardatus Carpenter 1966

G. †Pseudooedischia Handlirsch 1919

†Pseudooedischia sellardsi Lameere 1917

Invalid names: Pseudooedischia berthaudi Handlirsch 1919 [objective synonym]

Invalid names: Pseudomesoptilus Guan et al. 2016 [objective synonym]

G. †Rhinomaloptila Rasnitsyn 1977

†Rhinomaloptila polyneura Rasnitsyn 1977

G. †Spargoptilon Kukalová 1965

†Spargoptilon confertus Kukalová 1965

†Spargoptilon latericius Kukalová 1965

G. †Strephocladus Scudder 1885

†Strephocladus gandi Prokop et al. 2015

†Strephocladus mouralsensis Prokop et al. 2015

†Strephocladus permianus Guan et al. 2016

†Strephocladus subtilis Kliver 1883

G. †Strephoneura Martynov 1940

†Strephoneura pallida Novokshonov and Aristov 2004

†Strephoneura robusta Martynov 1940

†Strephoneura tenebrosa Novokshonov and Aristov 2004

†Strephoneura vasta Novokshonov and Aristov 2004

G. †Strephoptilus Rasnitsyn and Aristov 2013

†Strephoptilus borealis Rasnitsyn and Aristov 2013

†Strephoptilus longus Rasnitsyn and Aristov 2013

G. †Westphaloptilus Guan et al. 2016

†Westphaloptilus gallicus Guan et al. 2016

Invalid names: Prostenoneuridae Handlirsch 1919 [objective synonym], Strephocladidae Martynov 1938 [synonym], Strephoneuridae Martynov 1940 [synonym]

Fm. †Blattinopsidae Bolton 1925

G. †Alienus Handlirsch 1906

†Alienus lebachensis Goldenberg 1877

G. †Avionblattinopsis Quispe et al. 2021

†Avionblattinopsis oudardi Quispe et al. 2021

G. †Balduriella Meunier 1925

†Balduriella latissima Meunier 1925

G. †Blattinopsis Giebel 1867

†Blattinopsis americana Handlirsch 1906

†Blattinopsis anthracina Handlirsch 1906

†Blattinopsis antoniana Kukalová 1959

†Blattinopsis arnhardti Müller 1977

†Blattinopsis augustai Kukalová 1959

†Blattinopsis campestris Kukalová 1965

†Blattinopsis confusa Handlirsch 1937

†Blattinopsis ebersi Dohrn 1867

†Blattinopsis fimbriata Bolton 1925

†Blattinopsis goldenbergi Brongniart 1885

†Blattinopsis huysseni von Fritsch 1899

†Blattinopsis indefinitus Rasnitsyn and Aristov 2021

†Blattinopsis kukalovae Carpenter 1966

†Blattinopsis laqueata Scudder 1895

†Blattinopsis latipennis Handlirsch 1906

†Blattinopsis latissima Kukalová 1959

†Blattinopsis major Handlirsch 1937

†Blattinopsis martynovae Kukalová 1959

†Blattinopsis ornata Bolton 1925

†Blattinopsis ovalis Brongniart 1894

†Blattinopsis postuma Haupt 1952

†Blattinopsis postumus Haupt 1952

†Blattinopsis reticulata Germar 1851

†Blattinopsis seberi Hörnschemeyer and Stapf 2001

†Blattinopsis tardefurcata Müller 1977

†Blattinopsis taschenbergi Schlechtendal 1906

Invalid names: Anadyomene von Fritsch 1899 [synonym], Anadyomenella Strand 1929 [synonym], Blattinopsis (Blattinopsis) Giebel 1867 [invalid subgroup], Megablattina Brongniart 1885 [synonym], Oryctoblattina Scudder 1879 [objective synonym], Oryctomylabris Handlirsch 1906 [synonym], Palaeoricanites Haupt 1941 [synonym], Pseudofulgora Handlirsch 1906 [synonym]

G. †Glaphyrophlebia Handlirsch 1906

†Glaphyrophlebia anderhalterorum Beckemeyer 2013

†Glaphyrophlebia arnulfi Hörnschemeyer and Stapf 2001

†Glaphyrophlebia borea Aristov et al. 2021

†Glaphyrophlebia clava Kukalová 1965

†Glaphyrophlebia delicatula Bolton 1934

†Glaphyrophlebia glinka Aristov et al. 2022

†Glaphyrophlebia iva Aristov and Rasnitsyn 2021

†Glaphyrophlebia jeckenbachi Hörnschemeyer and Stapf 2001

†Glaphyrophlebia kamaensis Aristov and Rasnitsyn 2022

†Glaphyrophlebia komia Aristov et al. 2021

†Glaphyrophlebia oblonga Deichmüller 1882

†Glaphyrophlebia ovata Sellards 1909

†Glaphyrophlebia parvavena Hörnschemeyer and Stapf 2001

†Glaphyrophlebia popasnaya Aristov and Rasnitsyn 2022

†Glaphyrophlebia pusilla Handlirsch 1906

†Glaphyrophlebia pygmaea Meunier 1907

†Glaphyrophlebia rara Aristov and Rasnitsyn 2022

†Glaphyrophlebia rohwedderi Hörnschemeyer and Stapf 2001

†Glaphyrophlebia rossica Martynov 1931

†Glaphyrophlebia semipenna Aristov et al. 2022

†Glaphyrophlebia speciosa Sellards 1909

†Glaphyrophlebia subcostalis Martynov 1928

Invalid names: Sindonopsis reducta Martynov 1928 [synonym]

†Glaphyrophlebia uralensis Martynov 1940

†Glaphyrophlebia vaga Aristov and Rasnitsyn 2022

†Glaphyrophlebia victoiriensis Nel et al. 2022

†Glaphyrophlebia vorkutensis Aristov et al. 2021

†Glaphyrophlebia wettinensis Handlirsch 1906

Invalid names: Blattinopsiella Meunier 1907 [synonym], Prisca Handlirsch 1906 [synonym], Protociccus Brongniart 1885 [nomen nudum], Pursa Sellards 1909 [synonym], Sindon Sellards 1909 [synonym], Sindonopsis Martynov 1928 [synonym]

G. †Klebsiella Meunier 1908

†Klebsiella exstincta Meunier 1908

G. †Microblattina Scudder 1895

†Microblattina perdita Scudder 1895

G. †Rhipidioptera Brongniart 1894

†Rhipidioptera elegans Brongniart 1894

G. †Stephanopsis Kukalova 1958

†Stephanopsis elegans Schlechtendal 1906

†Stephanopsis incerta Laurentiaux 1950

†Stephanopsis mirandus Richardson 1956

Invalid names: Glaphyrokoris Richardson 1956 [synonym]

G. †Westphaloblattinopsis Béthoux and Jarzembowski 2010

†Westphaloblattinopsis edwardsae Béthoux and Jarzembowski 2010

Invalid names: Klebsiellidae Handlirsch 1919 [synonym], Oryctoblattinidae Handlirsch 1906 [replaced]

Fm. †Katerinkidae Prokop and Nel 2007

G. †Katerinka Prokop and Nel 2007

†Katerinka hilaris Prokop and Nel 2007

G. †Protoblattiniella Meunier 1912

†Protoblattiniella minutissima Meunier 1912

Invalid names: Protoptera Rasnitsyn 1977 [synonym]

G. †Parablattogryllus Storozhenko 1988

†Parablattogryllus obscurus Storozhenko 1988

G. †Permofossilis Aristov 2013

†Permofossilis commasticatus Aristov 2013

Or. Plecoptera Burmeister 1839 [stonefly]

G. †Afroperla van Dijk and Geertsema 2004

†Afroperla permiana van Dijk and Geertsema 2004

Subor. Antarctoperlaria Zwick 1973

Superfm. Eusthenioidea Banks 1913

Fm. Eustheniidae Banks 1913

Superfm. Gripopterygoidea Enderlein 1909

Fm. Gripopterygidae Enderlein 1909

Invalid names: Austroperlidae Tillyard 1921 [empty]

Invalid names: Gripopterygomorpha Enderlein 1909 [invalid subgroup]

Subor. Arctoperlaria Zwick 1973

Infraor. Euholognatha Zwick 1973

Superfm. Nemouroidea Billberg 1820

Fm. †Kachinoperlidae Chen 2022

G. †Kachinoperla Chen 2022

Fm. †Perspicuusoperlidae Chen 2022

G. †Perspicuusoperla Chen 2022

Infraor. Systellognatha Enderlein 1969

Superfm. Perloidea Latreille 1802

Fm. †Petroperlidae Sroka et al. 2018

Superfm. Pteronarcyoidea Newman 1853

Invalid names: Perlomorpha Latreille 1802 [synonym]

Invalid names: Nemourina Newman 1853 [synonym]

G. †Capitiperla Lin 1992

†Capitiperla tonicopoda Lin 1992

Fm. †Fatjanopteridae Martynova 1961

G. †Avionptera Schubnel et al. 2019

†Avionptera communeaui Schubnel et al. 2019

G. †Fatjanoptera Martynova 1961

†Fatjanoptera mnemonica Martynova 1961

G. †Fluminiperla Lin 1980

†Fluminiperla hastis Lin 1980

Fm. †Gulouidae Béthoux et al. 2011

G. †Gulou Béthoux et al. 2011

†Gulou carpenteri Béthoux et al. 2011

†Gulou oudardi Schubnel et al. 2019

G. †Kaptsheranga Sinitshenkova 1985

†Kaptsheranga porrecta Sinitshenkova 1985

G. †Marciperla Lin 1986

†Marciperla curta Lin 1986

Subor. †Perlopseina Martynov 1940

Fm. †Perlopseidae Martynov 1940

G. †Perlopsis Martynov 1940

G. †Sinoperla Hong 1992

†Sinoperla liaoningensis Hong 1992

Invalid names: Perlaria Latreille 1802 [synonym], Perlina Latreille 1802 [invalid subgroup]

Or. †Protorthoptera Handlirsch 1906

G. †Acridites Germar 1842

†Lithosialis carbonaria Germar 1842

Fm. †Adeloneuridae Carpenter 1938

G. †Adeloneura Carpenter 1938

†Adeloneura thompsoni Carpenter 1938

G. †Agogoblattina Handlirsch 1906

†Agogoblattina occidua Scudder 1885

G. †Anthracomastax Handlirsch 1904

Fm. †Anthracothremmidae Handlirsch 1906

G. †Anthracothremma Scudder 1885

†Anthracothremma robusta Scudder 1885

G. †Melinophlebia Handlirsch 1911

†Melinophlebia analis Handlirsch 1911

G. †Pericalyphe Handlirsch 1911

†Pericalyphe longa Handlirsch 1911

G. †Silphion Handlirsch 1911

†Silphion latipenne Handlirsch 1911

Fm. †Apithanidae Handlirsch 1911

G. †Apithanus Handlirsch 1911

†Apithanus jocularis Handlirsch 1911

Fm. †Appertiidae Paulian 1965

G. †Appertia Paulian 1965

†Appertia besairiei Paulian 1965

G. †Archaeologus Handlirsch 1906

†Archaeologus falcatus Handlirsch 1906

G. †Archimastax Handlirsch 1906

†Archimastax americanus Handlirsch 1906

G. †Asiopompus Sharov 1961

†Asiopompus tomicus Sharov 1961

G. †Boutakovia Pruvost 1934

†Boutakovia saleei Pruvost 1934

G. †Cacurgellus Pruvost 1919

†Cacurgellus barryi Pruvost 1919

G. †Chrestotella Handlirsch 1919

†Chrestotella danae Scudder 1868

G. †Chrestotes Scudder 1868

†Chrestotes lapidea Scudder 1868

G. †Commentrya Lameere 1917

†Commentrya maximae Brongniart 1894

Invalid names: Oedischiopsis Handlirsch 1919 [objective synonym]

Fm. †Cymbopsidae Kukalová 1965

G. †Cymbopsis Kukalová 1965

†Cymbopsis excelsa Kukalová 1965

G. †Danielsiella Cockerell 1916

†Danielsiella priscula Cockerell 1916

G. †Distasis Handlirsch 1904

†Distasis rhipiphora Handlirsch 1904

Fm. †Eucaenidae Handlirsch 1906

G. †Eucaenus Scudder 1885

†Eucaenus ovalis Scudder 1885

Invalid names: Eucaenus attenuatus Melander 1903 [synonym], Eucaenus mazonus Melander 1903 [synonym], Eucaenus minor Handlirsch 1911 [synonym], Teneopteron mirabile Carpenter 1943 [synonym]

Invalid names: Teneopteron Carpenter 1943 [synonym]

Invalid names: Teneopteridae Richardson 1956 [synonym]

G. †Gerarianus Handlirsch 1919

†Gerarianus commentryanus Handlirsch 1919

G. †Gerarites Handlirsch 1919

†Gerarites commentryi Brongniart 1894

G. †Geraroides Handlirsch 1906

†Geraroides maximus Melander 1903

Fm. †Hadentomidae Handlirsch 1906

G. †Hadentomum Handlirsch 1906

†Hadentomum americanum Handlirsch 1906

Invalid names: Palaeocixiidae Handlirsch 1919 [synonym]

G. †Hadentomoides Riek 1974

†Hadentomoides dwykensis Riek 1974

G. †Hemeristia Dana 1864

†Hemeristia occidentalis Dana 1864

Fm. †Herdinidae Carpenter and Richardson 1971

G. †Herdina Carpenter and Richardson 1971

†Herdina mirificus Carpenter and Richardson 1971

Fm. †Homalophlebiidae Handlirsch 1906

G. †Homalophlebia Brongniart 1894

†Homalophlebia finoti Brongniart 1894

G. †Parahomalophlebia Handlirsch 1906

†Parahomalophlebia courtini Brongniart 1894

Fm. †Homoeodictyidae Martynov 1937

G. †Homoeodictyon Martynov 1937

†Homoeodictyon elongatum Martynov 1937

G. †Hyaloptilus Lameere 1917

†Hyaloptilus minimus Lameere 1917

G. †Kelleropteron Brauckmann and Hahn 1980

†Kelleropteron kaelberbergense Brauckmann and Hahn 1980

G. †Lecopterum Sellards 1909

†Lecopterum delicosum Sellards 1909

G. †Limburgina Laurentiaux 1950

†Limburgina antiqua Laurentiaux 1950

G. †Macrophlebium Goldenberg 1869

†Macrophlebium hollebeni Goldenberg 1869

Fm. †Megagnathidae Tillyard 1918

G. †Megagnatha Bolton 1917

†Megagnatha odonatiformis Bolton 1917

G. †Megalometer Handlirsch 1906

†Megalometer lata Handlirsch 1906

G. †Melanoblattula Cockerell 1927

†Melanoblattula nigrescens Cockerell 1927

G. †Metacheliphlebia Handlirsch 1906

†Metacheliphlebia elongata Scudder 1885

G. †Notoblattites Tillyard 1916

†Notoblattites mitchelli Tillyard 1918

†Notoblattites subcostalis Tillyard 1916

†Notoblattites wianamattensis Tillyard 1918

G. †Ochetopteron Cockerell 1927

†Ochetopteron canaliculatum Cockerell 1927

G. †Orthoneurites Martynov 1928

†Orthoneurites regularis Martynov 1928

Fm. †Pachytylopsidae Handlirsch 1904

G. †Pachytylopsis de Borre 1875

†Pachytylopsis deriodoti Pruvost 1930

†Pachytylopsis melas Handlirsch 1904

†Pachytylopsis persenairei de Borre 1875

†Pachytylopsis stainieri Pruvost 1930

Invalid names: Palorthopteron Handlirsch 1904 [synonym]

G. †Protopachytylopsis Laurentiaux and Laurentiaux-Vieira 1981

†Protopachytylopsis leckwycki Laurentiaux and Laurentiaux-Vieira 1981

G. †Symballophlebia Handlirsch 1904

†Symballophlebia latipennis Handlirsch 1904

G. †Palaeocarria Cockerell 1917

†Palaeocarria ornata Cockerell 1917

G. †Palaeoedischia Meunier 1914

†Palaeoedischia boulei Meunier 1914

G. †Polyernus Scudder 1885

†Polyernus complanatus Scudder 1885

G. †Polyetes Handlirsch 1906

†Polyetes furcifer Handlirsch 1906

†Polyetes problematica Bolton 1934

G. †Prototettix Giebel 1856

†Prototettix lithanthraca Goldenberg 1854

Invalid names: Lithophasma Brongniart 1885 [objective synonym]

G. †Pruvostia Bolton 1921

†Pruvostia spectabilis Bolton 1921

G. †Pseudetoblattina Handlirsch 1906

†Pseudetoblattina reliqua Scudder 1893

G. †Pseudogerarus Handlirsch 1906

†Pseudogerarus scudderi Handlirsch 1906

G. †Pseudopolyernus Handlirsch 1906

†Pseudopolyernus laminarum Scudder 1885

G. †Ptenodera Bolton 1922

†Ptenodera dubius Bolton 1922

Fm. †Rigattopteridae Pinto 1996

G. †Rigattoptera Pinto 1996

†Rigattoptera ornellasae Pinto 1996

G. †Roomeria Meunier 1914

†Roomeria carbonaria Meunier 1914

G. †Schuchertiella Handlirsch 1911

†Schuchertiella gracilis Handlirsch 1911

G. †Sharovia Pinto and de Ornellas 1978

†Sharovia permiafricana Pinto and de Ornellas 1978

Fm. †Sheimiidae Martynova 1958

G. †Magniscapa Shcherbakov 2015

†Magniscapa alligans Aristov and Rasnitsyn 2008

G. †Parasheimia Aristov 2004

†Parasheimia kazanica Aristov and Rasnitsyn 2008

†Parasheimia rotundata Aristov 2004

†Parasheimia truncata Aristov 2004

G. †Pseudosheimia Aristov 2004

†Pseudosheimia caudata Aristov 2004

†Pseudosheimia caudissima Aristov and Rasnitsyn 2008

G. †Sheimia Martynova 1958

†Sheimia sojanensis Martynova 1958

†Sheimia tshekardensis Aristov 2004

G. †Soyanembia Aristov and Rasnitsyn 2008

†Soyanembia sharovi Aristov and Rasnitsyn 2008

G. †Visherifera Novokshonov et al. 2002

†Visherifera camura Novokshonov et al. 2002

†Visherifera sylvaensis Aristov 2004

Invalid names: Visheriferinae Novokshonov et al. 2002 [synonym]

Fm. †Soyaniidae Shcherbakov 2015

G. †Soyania Aristov and Rasnitsyn 2008

†Soyania logica Aristov and Rasnitsyn 2008

Fm. †Stephanotermopsidae Laurentiaux 1966

G. †Stephanotermopsis Laurentiaux 1966

†Stephanotermopsis rodendorfi Laurentiaux 1966

Fm. †Stygnidae Handlirsch 1906

G. †Stygne Handlirsch 1906

†Stygne roemeri Handlirsch 1906

Fm. †Thoronysididae Handlirsch 1919

G. †Thoronysis Handlirsch 1906

†Thoronysis ingbertensis von Ammon 1903

Invalid names: Asiopompidae Sharov 1961 [invalid subgroup], Atava Sellards 1909 [nomen dubium], Atava ovata Sellards 1909 [nomen dubium], Eucaenus pusillus Handlirsch 1911 [nomen dubium], Eucaenus rotundatus Handlirsch 1906 [nomen dubium], Hadentomoidea Handlirsch 1906 [synonym], Protoblattinopsidae Laurentiaux 1966 [empty], Protoperlaria Tillyard 1928 [synonym], Prototettigidae Handlirsch 1906 [invalid subgroup], Roomeriidae Handlirsch 1919 [invalid subgroup], Schuchertiellidae Handlirsch 1911 [invalid subgroup], Stenaropodidae Handlirsch 1906 [empty]

G. †Suksunus Aristov 2015

†Suksunus bicodex Aristov 2015

Unr. Xenonomia Terry and Whiting 2005

Or. Grylloblattodea Walker 1914 [ice crawler]

G. †Angaroptera Wachtler 2017

†Angaroptera nicolaswachtleri Wachtler 2017

Fm. †Aristoviidae Storozhenko and Gröhn 2023

G. †Aristovia Storozhenko and Gröhn 2023

Fm. †Euchauliodidae Riek 1974

G. †Euchauliodes Riek 1974

G. †Lodevoisadia Nel et al. 2022

†Lodevoisadia coheni Nel et al. 2022

Invalid names: Grylloblattida Walker 1914 [replaced]

Or. Mantophasmatodea Zompro et al. 2002 [gladiator]

Fm. Mantophasmatidae Zompro et al. 2002

G. †Adicophasma Engel and Grimaldi 2004

G. †Ensiferophasma Zompro 2005

G. †Raptophasma Zompro 2001

Invalid names: Ensiferophasmatidae Zompro 2005 [synonym], Raptophasmatidae Zompro 2005 [synonym]

Invalid names: Blattopteroidea Martynov 1924 [empty], Dermapterida [invalid subgroup], Mystroptera [invalid subgroup], Orthacanthacris incertus Piton 1940 [nomen dubium], Orthopteroidea [empty], Plecopterida [invalid subgroup], Polyplecoptera [invalid subgroup], Spectra Latreille 1802 [empty]

G. †Prosigara Scudder 1890

†Prosigara flabellum Scudder 1890

Or. Psocodea Hennig 1966 [bark louse]

Subor. Psocomorpha Roesler 1944

Infraor. Archipsocetae Yoshizawa 2002

Fm. Archipsocidae Enderlein 1903 [ancient barklouse]

Subfm. Archipsocinae Enderlein 1903

G. †Awightipsocus Azar 2014

†Awightipsocus jarzembowskii Azar 2014

†Awightipsocus megaloi Azar 2014

†Awightipsocus minimus Azar 2014

Infraor. Caeciliusetae Pearman 1936

Fm. Amphipsocidae Pearman 1936 [hairy-winged barklouse]

Subfm. Amphipsocinae Pearman 1936

Fm. Caeciliusidae Kolbe 1880 [lizard barklouse]

Subfm. Caeciliusinae Kolbe 1880

G. †Ptenolasia Enderlein 1911

Invalid names: Caeciliusoidea Kolbe 1880 [invalid subgroup]

G. †Cretapsocus Vishnyakova 1975

†Cretapsocus capillatus Vishnyakova 1975

Infraor. Epipsocetae Pearman 1936

Fm. Epipsocidae Karny 1930 [elliptical bark louse]

G. Epipsocus Hagen 1866

G. Mesepipsocus Badonnell 1969

Fm. Ptiloneuridae Enderlein 1910

G. †Brisacia Azar et al. 2009

G. †Eotriplocania Zhang et al. 2016

G. Euplocania Enderlein 1910

Infraor. Homilopsocidea Pearman 1936

G. †Burmesopsocus Yoshizawa 2021

†Burmesopsocus lienhardi Yoshizawa 2021

Fm. Ectopsocidae Roesler 1940 [outer barklouse]

G. Ectopsocus McLachlan 1899

Fm. Elipsocidae Kolbe 1882 [damp barklouse]

Subfm. Elipsocinae Kolbe 1882

Fm. Lachesillidae Karny 1930 [fateful barklouse]

G. †Archaelachesis Vishnyakova 1975

Subfm. Eolachesillinae Mockford and Sullivan 1986

Fm. Mesopsocidae Enderlein 1903

G. †Mesopsocoides Azar et al. 2014

G. Mesopsocus Kolbe 1880

Invalid names: Elipsocoidea Kolbe 1882 [invalid subgroup], Lachesilloidea Karny 1930 [invalid subgroup], Peripsocoidea Pearman 1936 [invalid subgroup]

Infraor. Philotarsetae Yoshizawa and Johnson 2014

Fm. Philotarsidae Pearman 1936 [loving barklouse]

Subfm. Philotarsinae Pearman 1936

Fm. Pseudocaeciliidae Pearman 1936 [false lizard barklouse]

Subfm. †Electropsocinae Roesler 1940

Fm. Trichopsocidae Pearman 1936 [lash-faced psocid]

G. †Palaeopsocus Kolbe 1883

G. Trichopsocus Kolbe 1882

Invalid names: Pseudocaecilioidea Pearman 1936 [invalid subgroup]

Infraor. Psocetae Pearman 1936

Fm. Myopsocidae Enderlein 1901 [mouse like barklouse]

G. Myopsocus Hagen 1866

Fm. Psocidae Leach 1815 [common bark louse]

Subfm. Psocinae Leach 1815

G. †Wightimicropsocus Azar 2014

†Wightimicropsocus inexpectatus Azar 2014

G. †Wightipsocus Azar 2014

†Wightipsocus acourti Cockerell 1921

†Wightipsocus intasi Azar 2014

†Wightipsocus rossi Azar 2014

Invalid names: Hemipsocetae Yoshizawa 2002 [empty]

Subor. Troctomorpha Roesler 1940 [book louse]

Infraor. Amphientometae Pearman 1936

Superfm. Amphientomoidea Kolbe 1883

Fm. Amphientomidae Kolbe 1883 [tropical barklouse]

Invalid names: Arcantipsocidae Azar et al. 2009 [synonym]

Superfm. Electrentomoidea Enderlein 1911

Fm. Compsocidae Mockford 1967

Fm. †Electrentomidae Enderlein 1911

Invalid names: Paramesopsocidae Azar et al. 2008 [synonym]

G. †Libanomphientomum Choufani et al. 2011

Fm. Troctopsocidae Mockford 1967

G. †Scocompus Azar et al. 2014

†Scocompus atelisus Azar et al. 2014

Fm. Manicapsocidae Mockford 1967

G. †Azarpsocus Maheu and Nel 2019

†Azarpsocus perreaui Maheu and Nel 2019

G. †Eomanicapsocus Nel et al. 2005

†Eomanicapsocus melaniae Nel et al. 2005

G. †Eoprotroctopsocus Nel et al. 2005

†Eoprotroctopsocus celinea Nel et al. 2005

G. †Palaeomanicapsocus Azar et al. 2016

†Palaeomanicapsocus fouadi Azar et al. 2016

†Palaeomanicapsocus margoae Azar et al. 2016

G. †Paramanicapsocus Hakim et al. 2019

†Paramanicapsocus longiantennatus Hakim et al. 2019

Infraor. Nanopsocetae Pearman 1936

Fm. Liposcelididae Enderlein 1911 [booklouse]

G. †Cretoscelis Grimaldi and Engel 2006

Subfm. Embidopsocinae Karny 1930

Subfm. Liposcelidinae Enderlein 1911

Fm. Pachytroctidae Enderlein 1911 [thick barklouse]

Subfm. †Libaneuphorinae Azar et al. 2015

Subfm. Tapinellinae Enderlein 1908

Unr. Phthiraptera Haeckel 1896 [parasitic louse]

Superfm. Amblycera Kellogg 1896

Invalid names: Anoplura Leach 1815 [empty], Ischnocera Kellogg 1896 [empty]

Fm. Sphaeropsocidae Kolbe 1883

G. †Asphaeropsocites Azar et al. 2010

G. †Sphaeropsocites Grimaldi and Engel 2006

G. †Sphaeropsocoides Grimaldi and Engel 2006

G. Sphaeropsocus Hagen 1882

Invalid names: Palaeotroctes Enderlein 1911 [synonym]

Subor. Trogiomorpha Roesler 1940

Infraor. Atropetae Pearman 1936

Fm. †Empheriidae Kolbe 1883

G. †Archaeatropos Baz and Ortuño 2000

G. †Bcharreglaris Azar and Nel 2004

G. †Burmempheria Li et al. 2020

G. †Empheria Hagen 1856

G. †Empherium Hakim et al. 2021

G. †Empheropsocus Baz and Ortuño 2001

G. †Eoempheria Nel et al. 2005

G. †Heliadesdakruon Cumming and Le Tirant 2021

G. †Jerseyempheria Azar et al. 2010

G. †Latempheria Li et al. 2022

G. †Libanoglaris Azar et al. 2003

G. †Preempheria Baz and Ortuño 2001

G. †Proprionoglaris Perrichot et al. 2003

G. †Prospeleketor Perrichot et al. 2003

G. †Setoglaris Azar and Nel 2004

G. †Trichempheria Enderlein 1911

Invalid names: Archaeatropidae Baz and Ortuño 2000 [synonym]

Fm. Lepidopsocidae Enderlein 1903 [scaly-winged bark louse]

Subfm. Perientominae Kolbe 1882

Subfm. Thylacinae Enderlein 1903

Invalid names: Thylacellinae Roesler 1944 [synonym]

Fm. Psoquillidae Kolbe 1882

G. †Eorhyopsocus Nel et al. 2005

Fm. Trogiidae Enderlein 1911 [granary booklouse]

G. †Cretolepinotus Cockx et al. 2020

G. †Eolepinotus Vishnyakova 1975

G. †Paralepinotus Azar et al. 2018

G. †Brachyantennum Liang and Liu 2022

†Brachyantennum spinosum Liang and Liu 2022

Fm. †Cormopsocidae Yoshizawa and Lienhard 2020

G. †Cormopsocus Yoshizawa and Lienhard 2020

†Cormopsocus baleoi Hakim et al. 2021

†Cormopsocus groehni Yoshizawa and Lienhard 2020

†Cormopsocus neli Hakim et al. 2021

†Cormopsocus perantiqua Cockerell 1919

G. †Longiglabellus Wang et al. 2021

†Longiglabellus edentatus Wang et al. 2021

†Longiglabellus pedhyalinus Wang et al. 2021

G. †Stimulopsocus Liang and Liu 2021

†Stimulopsocus jiewenae Liang and Liu 2021

G. †Empheriopsis Vishnyakova 1975

†Empheriopsis vulnerata Vishnyakova 1975

G. †Parapsyllipsocus Perrichot et al. 2003

†Parapsyllipsocus vergereaui Perrichot et al. 2003

Infraor. Prionoglaridetae Smithers 1972

Fm. Prionoglarididae Karny 1930

Subfm. Prionoglaridinae Karny 1930

Infraor. Psyllipsocetae Smithers 1972

Fm. Psyllipsocidae Kolbe 1884 [cave barklouse]

G. †Annulipsyllipsocus Hakim et al. 2017

G. †Concavapsocus Wang et al. 2019

G. †Globopsocus Azar and Engel 2008

G. †Khatangia Vishnyakova 1975

G. †Libanopsyllipsocus Azar and Nel 2011

G. Psyllipsocus Selys-Longchamps 1872

G. †Sinopsyllipsocus Zhang et al. 2016

Invalid names: Psocatropetae Pearman 1936 [invalid subgroup]

Invalid names: Psocoptera [replaced]

Fm. †Saurodectidae Rasnitsyn and Zherikhin 1999

G. †Saurodectes Rasnitsyn and Zherikhin 1999

†Saurodectes vrsanskyi Rasnitsyn and Zherikhin 1999

G. †Sibosmylina Ponomarenko 1985

†Sibosmylina libelluloides Ponomarenko 1985

Fm. †Sinembiidae Huang and Nel 2009

G. †Juraembia Huang and Nel 2009

†Juraembia ningchengensis Huang and Nel 2009

G. †Sinembia Huang and Nel 2009

†Sinembia rossi Huang and Nel 2009

G. †Sphalmatoblattina Handlirsch 1906

†Sphalmatoblattina latinervis Heer 1864

G. †Sycopteron Bolton 1917

†Sycopteron symmetricum Bolton 1917

Fm. †Synomaloptilidae Martynov 1938

G. †Synomaloptila Martynov 1938

†Synomaloptila longipennis Martynov 1938

G. †Tillyardites Handlirsch 1937

†Tillyardites bairdae Tillyard 1922

Fm. †Trachopterygidae Carpenter 1976

G. †Trachopteryx Carpenter 1976

†Trachopteryx martynovi Carpenter 1976

G. †Triadochorista Geyer and Kleber 1987

†Triadochorista schleeriethensis Geyer and Kleber 1987

G. †Trichorthophlebia Handlirsch 1939

†Trichorthophlebia pilifera Handlirsch 1939

Fm. †Uninervidae Becker-Migdisova 1951

G. †Mononeura Becker-Migdisova 1951

†Mononeura angustipennis Becker-Migdisova 1951

G. †Redactineura Riek 1973

†Redactineura acuminata Riek 1973

G. †Uninervus Becker-Migdisova 1951

†Uninervus zorapteroides Becker-Migdisova 1951

Invalid names: Blattoneoptera Kukalová-Peck 1991 [empty], Dictyophara scudderi Piton 1940 [nomen dubium], Endoneoptera [empty], Hemineoptera [empty], Naucoris crassus Piton 1935 [nomen dubium], Orthoneoptera [empty], Paraneoptera [invalid subgroup], Sycopteridae Tillyard 1918 [invalid subgroup]

Diagnosis

No diagnoses are available

Measurements

No measurements are available

Composition:

chitinc

Environment:

terrestrialc

Locomotion:

actively mobilep

Created:

2017-04-17 09:56:18

Modified:

2017-04-17 09:56:18

Source: c = class, p = phylum

References: Kiessling 2004, Bush and Bambach 2015

Age range

Maximum range based only on fossils: base of the Morrowan to the top of the Holocene or 318.10000 to 0.00000 Ma
Minimum age of oldest fossil (stem group age): 315.2 Ma

Collections (4909 total)

Time interval	Ma	Country or state	Original ID and collection number
Oldest occurrences
Bashkirian	323.2 - 315.2	USA (Illinois)	Mylacris anthracophila (122575)
Bashkirian - Moscovian	323.2 - 307.0	Argentina (Chubut)	Anthracoblattina archangelskyi, Archangelskyblatta vishniakovae (162148)
Pennsylvanian	323.2 - 298.9	USA (New Mexico)	Phyloblattidae indet. (217076)
Pennsylvanian	323.2 - 298.9	Russian Federation	Archimylacridae indet. (117186)
Alportian - Kinderscoutian	318.1 - 314.6	Netherlands	Ampeliptera limburgica (137302)
Kinderscoutian	318.1 - 314.6	Poland	Stygne roemeri (123954)
Marsdenian	318.1 - 314.6	Germany (Ruhr)	Baryshnyala occulta (152121) Heterologopsis ruhrensis (155261)
Marsdenian	318.1 - 314.6	Netherlands	Limburgina antiqua (157405)
Marsdenian	318.1 - 314.6	Belgium	Protoprosbole straeleni (164479)
Morrowan	318.1 - 314.6	USA (Arkansas)	Archimastax americanus (126550)
Westphalian A	318.1 - 314.6	Poland	Owadpteron dareki, Parapalaeomastax dariuszi (107805)
Westphalian A	318.1 - 314.6	France (Nord-Pas-de-Calais)	Climaconeura reumauxi (151687) Omalia macroptera (152785)
Westphalian A	318.1 - 314.6	Canada (New Brunswick)	Geroneura wilsoni (114468)
Westphalian A	318.1 - 314.6	Germany	Manoblatta oberstebrinki (157047) Micropalentomum minusculum (157048)
Westphalian A	318.1 - 314.6	United Kingdom (England)	Genentomum subacutum (127291) Plesioidischia sp. (122420)
Westphalian A	318.1 - 314.6	United Kingdom (Wales)	Blattodea indet. (151394)
Westphalian A	318.1 - 314.6	Czech Republic	Katerinka hilaris (124366)
Westphalian A	318.1 - 314.6	France	Westphalomerope maryvonneae (122506)
Westphalian A	318.1 - 314.6	Belgium	Omalia macroptera (123821) Protopachytylopsis leckwycki (156161)
Westphalian A	318.1 - 314.6	Canada (Nova Scotia)	Caloneurodea indet. (216802) Mixotermitidae indet. (216607)