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Itemirus medullaris

Reptilia - Avetheropoda - Dromaeosauridae

Taxonomy
Itemirus medullaris was named by Kurzanov (1976). Its type specimen is PIN 327/699. Its type locality is Dzharakuduk II (general), which is in a Turonian/Turonian delta plain sandstone in the Bissekty Formation of Uzbekistan. It was considered monophyletic by Currie (2000).

Synonymy list
YearName and author
1976Itemirus medullaris Kurzanov p. 363 figs. 1-3
1981Itemirus medullaris Nessov p. 91
1991Itemirus medullaris Molnar p. 167
1997Itemirus medullaris Ryan p. 443
1998Itemirus medullaris Makovicky and Norell p. 13
2000Itemirus medullaris Currie p. 450
2004Itemirus medullaris Holtz, Jr. p. 114
2009Itemirus medularis Bever and Norell p. 22
2009Itemirus medullaris Longrich and Currie p. supplementary
2014Itemirus medullaris Sues and Averianov p. 227

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RankNameAuthor
kingdomAnimalia()
Triploblastica
Nephrozoa
Deuterostomia
phylumChordataHaeckel 1847
OlfactoresJefferies 1991
subphylumVertebrata
Gnathostomata()
Osteichthyes()
Sarcopterygii
subclassDipnotetrapodomorpha(Nelson 2006)
subclassTetrapodomorpha()
Tetrapoda()
Reptiliomorpha
Anthracosauria
Batrachosauria()
Cotylosauria()
Amniota
Sauropsida
classReptilia
subclassEureptilia()
Romeriida
Diapsida()
Eosuchia()
RankNameAuthor
Neodiapsida
SauriaGauthier 1984
Archosauromorpha(Huene 1946)
Crocopoda
ArchosauriformesGauthier 1986
Eucrocopoda
Archosauria()
informalAvemetatarsalia
Ornithodira
Dinosauromorpha
Dinosauriformes
Dinosauria()
Saurischia()
Theropoda()
Neotheropoda
AverostraPaul 2002
Tetanurae
orderAvetheropoda
suborderCoelurosauria
Maniraptoriformes
familyDromaeosauridae
subfamilyDromaeosaurinae
genusItemirus
speciesmedullaris

If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.

Diagnosis
ReferenceDiagnosis
H.-D. Sues and A. Averianov 2014None of the features in the original diagnosis offered by Kurzanov (1976) has diagnostic value, and two characters cited by this author (position of foramen for endolymphatic duct and presence of “recessus interacusticus”) are based on incorrect anatomical interpretations. Itemirus is referable to Dromaeosauridae based on the following synapomorphies for this clade hypothesized by Turner et al. (2012): 17(0), accessory tympanic recess dorsal to crista interfenestralis absent; 96(0), anterior cervical centra level with or shorter than posterior extent of neural arch; 103(1), parapophyses of posterior trunk vertebrae distinctly projected on pedicels. Within Dromaeosauridae Itemirus medullaris is referable to Dromaeosaurinae on the basis of the following synapomorphies: 91(1), premaxillary tooth crowns asymmetrical (D-shaped) in cross-section, with flat lingual surface; 238(1), jugal process of maxilla ventral to external antorbital fenestra dorsoventrally wide. Among Dromaeosaurinae, Itemirus medullaris differs from Dromaeosaurus albertensis in characters 15(1), basipterygoid recesses on dorsolateral surfaces of basipterygoid processes present; 247(1), anterior denticles of teeth significantly smaller than posterior denticles. Itemirus differs from Utahraptor ostrommaysorum in character 265(1), pleurocoels present in anterior dorsals. Itemirus medullaris differs from both Utahraptor ostrommaysorum and Achillobator giganticus in character 318(1), lateral surfaces of centra of thoracic vertebrae have deep, emarginated fossae.