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Tupandactylus imperator

Reptilia -

Taxonomy
Tapejara imperator was named by Campos and Kellner (1997). Its type specimen is MCT 1622-R, a skull, and it is a compression fossil. Its type locality is Nova Olinda quarries, which is in an Aptian lacustrine - large lime mudstone/mudstone in the Crato Formation of Brazil.

It was recombined as Ingridia imperator by Unwin and Martill (2007); it was recombined as Tupandactylus imperator by Kellner and Campos (2007), Pinheiro et al. (2011), Andres et al. (2014) and Headden and Campos (2014).

Synonymy list
YearName and author
1997Tapejara imperator Campos and Kellner p. 83 figs. 1-2
2003Tapejara imperator Unwin
2007Tupandactylus imperator Kellner and Campos p. 2
2007Ingridia imperator Unwin and Martill
2011Tupandactylus imperator Pinheiro et al.
2014Tupandactylus imperator Andres et al.
2014Tupandactylus imperator Headden and Campos

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RankNameAuthor
kingdomAnimalia()
Triploblastica
Nephrozoa
Deuterostomia
phylumChordataHaeckel 1847
subphylumVertebrata
superclassGnathostomata
Osteichthyes()
Sarcopterygii
subclassDipnotetrapodomorpha(Nelson 2006)
subclassTetrapodomorpha()
Tetrapoda()
Reptiliomorpha
Anthracosauria
Batrachosauria()
Cotylosauria()
Amniota
Sauropsida
classReptilia
subclassEureptilia()
Romeriida
Diapsida()
RankNameAuthor
Eosuchia()
Neodiapsida
SauriaGauthier 1984
Archosauromorpha(Huene 1946)
Crocopoda
ArchosauriformesGauthier 1986
Eucrocopoda
Archosauria()
informalAvemetatarsalia
Ornithodira
PterosauromorphaPadian 1997
Pterosauria(Kaup 1834)
suborderPterodactyloideaPlieninger 1901
Lophocratia
superfamilyOrnithocheiroideaSeeley 1870
superfamilyAzhdarchoideaUnwin 1992
Neoazhdarchia
superfamilyTapejaroideaKellner 1989
Tapejaridae()
genusTupandactylus
speciesimperator()

If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.

Diagnosis
ReferenceDiagnosis
F. L. Pinheiro et al. 2011Tapejarid pterosaur with an occipital process that reaches about the length of the rest of the skull (measured from the tip of the premaxillae to the squamosals); an extremely large soft-tissue median cranial crest supported anteriorly by a spine-like, caudally inclined suprapremaxillary process; soft-tissue component of the cranial crest composed of parallel fibers curving in caudal direction; an anteriorly projecting convex blade on the premaxillae and a lower jaw bearing a very deep rounded median mandibular crest with a steep rostral margin forming an angle of approximately 60 degrees with the mandible.