PBDB Taxon

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Scutosaurus karpinskii

Reptilia - Pareiasauridae

Taxonomy

Pareiosaurus karpinskii was named by Amalitzky (1922). Its type specimen is PIN 2005/1532, a skeleton, and it is a 3D body fossil. Its type locality is Sokolki locality, PIN 2005, which is in a Severodvinian/Vyatkian terrestrial horizon in the Salarevo Formation of the Russian Federation. It is the type species of Scutosaurus.

It was recombined as Scutosaurus karpinsky by Hartmann-Weinberg (1930); it was recombined as Scutosaurus karpinskii by Lee (1997), Lee (2000) and Ivakhnenko (2008).

Synonyms

Proelginia permiana was named by Hartmann-Weinberg (1937). Its type specimen is PIN 156/2, a skull (without lower jaw), and it is a 3D body fossil. Its type locality is Semin Ovrag, PIN 156, which is in a Severodvinian terrestrial sandstone in the Russian Federation.

It was recombined as Scutosaurus permianus by Efremov (1940); it was synonymized subjectively with Scutosaurus karpinskii by Lee (1997) and Lee (2000).
Pareiosaurus elegans was named by Amalitzky (1922). Its type specimen is PIN 2005/1538, a skeleton, and it is a 3D body fossil. Its type locality is Sokolki locality, PIN 2005, which is in a Severodvinian/Vyatkian terrestrial horizon in the Salarevo Formation of the Russian Federation.

It was synonymized subjectively with Scutosaurus karpinskii by Lee (1997), Lee (2000) and Ivakhnenko (2008).
Pareiosaurus horridus was named by Amalitzky (1922). Its type specimen is PIN 2005/1535, a skeleton, and it is a 3D body fossil. Its type locality is Sokolki locality, PIN 2005, which is in a Severodvinian/Vyatkian terrestrial horizon in the Salarevo Formation of the Russian Federation.

It was synonymized subjectively with Scutosaurus karpinskii by Lee (1997), Lee (2000) and Ivakhnenko (2008).
Pareiosaurus tuberculatus was named by Amalitzky (1922). Its type specimen is PIN 2005/1533, a skeleton, and it is a 3D body fossil. Its type locality is Sokolki locality, PIN 2005, which is in a Severodvinian/Vyatkian terrestrial horizon in the Salarevo Formation of the Russian Federation.

It was synonymized subjectively with Scutosaurus karpinskii by Lee (1997) and Lee (2000); it was recombined as Scutosaurus tuberculatus by Ivakhnenko (2008).
Scutosaurus itelensis was named by Ivakhnenko (1987).

It was synonymized subjectively with Scutosaurus karpinskii by Lee (1997) and Lee (2000).

Synonymy list

Year	Name and author
1922	Pareiosaurus karpinskii Amalitzky p. 334 fig. 8
1922	Pareiosaurus elegans Amalitzky p. 335
1922	Pareiosaurus horridus Amalitzky p. 335
1922	Pareiosaurus tuberculatus Amalitzky p. 335 fig. 9
1930	Scutosaurus karpinsky Hartmann-Weinberg p. 59 fig. 1
1937	Proelginia permiana Hartmann-Weinberg pp. 691, 709 figs. Pls. X-XIII
1940	Scutosaurus permianus Efremov p. 137
1987	Scutosaurus itelensis Ivakhnenko
1997	Scutosaurus karpinskii Lee p. 271
2000	Scutosaurus karpinskii Lee p. 77 figs. 5.4-5.7
2008	Proelginia permiana Ivakhnenko p. 78 fig. 34
2008	Scutosaurus karpinskii Ivakhnenko p. 81 fig. 35
2008	Scutosaurus tuberculatus Ivakhnenko p. 82 fig. 36

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Rank	Name	Author
kingdom	Animalia	()
—	Bilateria
—	Eubilateria	Ax 1987
—	Deuterostomia
phylum	Chordata	Haeckel 1874
subphylum	Vertebrata
superclass	Gnathostomata
—	Osteichthyes	()
subclass	Sarcopterygii	()
subclass	Dipnotetrapodomorpha	(Nelson 2006)
subclass	Tetrapodomorpha	()
—	Tetrapoda
—	Reptiliomorpha
—	Anthracosauria

Rank	Name	Author
subclass	Amphibiosauria	Kuhn 1967
—	Cotylosauria	()
—	Amniota
—	Sauropsida
class	Reptilia
subclass	Parareptilia	Lawrence 1868
—	Procolophonomorpha	()
—	Procolophonia	()
—	Hallucicrania
superfamily	Pareiasauroidea	Nopsca 1928
—	Pareiasauria	()
family	Pareiasauridae	Lydekker 1889
genus	Scutosaurus
species	karpinskii	()

If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.

†Scutosaurus karpinskii Amalitzky 1922

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Invalid names: Pareiosaurus elegans Amalitzky 1922 [synonym], Pareiosaurus horridus Amalitzky 1922 [synonym], Pareiosaurus tuberculatus Amalitzky 1922 [synonym], Proelginia permiana Hartmann-Weinberg 1937 [synonym], Scutosaurus itelensis Ivakhnenko 1987 [synonym]

Diagnosis

Reference	Diagnosis
M. S. Y. Lee 2000	Large pareiasaur, 2.5 m in body length, with extremely rugose dermal sculpture, large cheek flanges with long pointed bosses, and osteoderms covering the entire body and limbs. Autapomorphies: small median tubercle on basioccipital between basal tubera; tips of teeth on upper jaw point slightly outwards; radiating ridges covering skull very coarse.

Measurements

No measurements are available

Composition:

phosphaticsubp

Environment:

terrestrialuc

Locomotion:

actively mobilec

Diet:

herbivoresubo

Created:

2009-01-03 20:32:59

Modified:

2009-01-03 22:32:59

Source: subo = suborder, c = class, subp = subphylum, uc = unranked clade

References: Benton 1983, Hendy et al. 2009, Carroll 1988

Age range: Late/Upper Severodvinian or 264.28000 to 259.51000 Ma

Collections (6 total)

Time interval	Ma	Country or state	Original ID and collection number
Late/Upper Severodvinian	264.28 - 259.51	Russian Federation (Orenburg)	Proelginia permiana (189501 189505)
Late/Upper Severodvinian	264.28 - 259.51	Russian Federation (Tatarstan)	Proelginia permiana (80712)
Late/Upper Severodvinian - Vyatkian	264.28 - 257.4	Russian Federation (Vologda)	Scutosaurus tuberculatus (187939)
Late/Upper Severodvinian - Vyatkian	264.28 - 257.4	Russian Federation (Arkhangelsk)	Pareiosaurus karpinskii, Pareiosaurus elegans, Pareiosaurus horridus, Pareiosaurus tuberculatus, Scutosaurus karpinskii, Scutosaurus tuberculatus (type locality: 28065) Scutosaurus karpinskii (27952)