PBDB Taxon

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Haglidae (hump-winged cricket)

Insecta - Orthoptera - Haglidae

Taxonomy

Haglidae was named by Handlirsch (1906) [also attributed to Zeuner (no date)]. Its type is Hagla.

It was assigned to Mantoidea by Handlirsch (1906); to Grylloidea by Riek (1974) and Hong (1982); to Gryllidea by Sharov (1968) and Whalley (1985); to Orthoptera by Hong (1986); to Ensifera by Carpenter (1992); and to Hagloidea by Gorochov (1986), Gorochov (1988), Gorochov (1995), Wappler (2001), Gorochov et al. (2006), Gu et al. (2017), Gu et al. (2021) and Gorochov and Coram (2022).

Synonyms

Tshorkuphlebiidae was named by Martynov (1937). It is not extant. Its type is Tshorkuphlebia.

It was reranked as the subfamily Tshorkuphlebiinae by Sharov (1968); it was synonymized subjectively with Haglidae by Carpenter (1992) and Gorochov (1995).

It was assigned to Protorthoptera by Martynov (1937); and to Haglidae by Sharov (1968).
Aboilidae was named by Martynov (1925). It is not extant. Its type is Aboilus.

It was synonymized subjectively with Prophalangopsidae by Zeuner (1939); it was synonymized subjectively with Haglidae by Sharov (1968) and Carpenter (1992); it was reranked as the subfamily Aboilinae by Gorochov (1988), Gorochov (1995), Grant-Mackie et al. (1996), Gorochov (2003), Gorochov et al. (2006), Li et al. (2007), Fang et al. (2007), Gu et al. (2011), Wang et al. (2015), Wang et al. (2018), Wang et al. (2018), Gorochov (2020) and Gu et al. (2022).

It was assigned to Aboilodea by Martynov (1925) and Martynov (1937); and to Prophalangopsidae by Gorochov (1988), Gorochov (1995), Grant-Mackie et al. (1996), Gorochov (2003), Gorochov et al. (2006), Li et al. (2007), Fang et al. (2007), Gu et al. (2011), Wang et al. (2015), Wang et al. (2018), Wang et al. (2018), Gorochov (2020) and Gu et al. (2022).

Synonymy list

Year	Name and author
1906	Haglidae Handlirsch p. 425
1925	Aboilidae Martynov p. 581
1937	Aboilidae Martynov p. 49
1937	Tshorkuphlebiidae Martynov p. 72
1968	Haglidae Sharov
1968	Tshorkuphlebiinae Sharov
1974	Haglidae Riek p. 20
1982	Haglidae Hong p. 71
1985	Haglidae Whalley
1986	Haglidae Gorochov p. 72
1986	Haglidae Hong
1988	Haglidae Gorochov p. 51
1988	Aboilinae Gorochov p. 59
1992	Haglidae Carpenter p. 165
1995	Haglidae Gorochov p. 111
1995	Aboilinae Gorochov p. 129
1996	Aboilinae Grant-Mackie et al. p. 32
2001	Haglidae Wappler p. 89
2003	Aboilinae Gorochov p. 119
2006	Haglidae Gorochov et al. p. 646
2006	Aboilinae Gorochov et al. p. 647
2007	Aboilinae Fang et al. p. 56
2007	Aboilinae Li et al. p. 412
2011	Aboilinae Gu et al. p. 65
2015	Aboilinae Wang et al. p. 251
2017	Haglidae Gu et al. p. 152
2018	Aboilinae Wang et al. p. 177
2020	Aboilinae Gorochov p. 1255
2021	Haglidae Gu et al. p. 185
2022	Haglidae Gorochov and Coram p. 4
2022	Aboilinae Gu et al. p. 1589

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Rank	Name	Author
kingdom	Animalia	()
—	Bilateria
—	Eubilateria	Ax 1987
—	Protostomia
—	Ecdysozoa
—	Panarthropoda
phylum	Arthropoda	Latreille 1829
superclass	Hexapoda	Latreille 1825
class	Insecta
—	Dicondylia
—	Paranotalia

Rank	Name	Author
subclass	Pterygota	()
—	Neopterygota	Crampton 1924
infraclass	Neoptera	Martynov 1923
—	Polyneoptera	Martynov 1938
order	Orthoptera
suborder	Ensifera	Chopard 1920
infraorder	Tettigoniidea
superfamily	Hagloidea	Handlirsch 1906
family	Haglidae
family	Haglidae

If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.

Fm. †Haglidae Handlirsch 1906 [hump-winged cricket]

show all | hide all

G. †Afrohagla Gorochov 1990

†Afrohagla contorta Riek 1974

G. †Ahagla Gorochov 2005

†Ahagla mira Gorochov 2005

Subfm. †Angarohaglinae Gorochov 1995

G. †Angarohagla Zherikhin 1985

†Angarohagla variegata Zherikhin 1985

G. †Microvoliopus Gorochov 1990

†Microvoliopus bacharicus Gorochov 1990

G. †Archaeohagla Lin 1965

†Archaeohagla sinensis Lin 1965

G. †Archihagla Sharov 1968

†Archihagla tenuis Gorochov 1986

†Archihagla zeuneri Sharov 1968

Subfm. †Bachariinae Gorochov 1988

G. †Bacharia Gorochov 1988

†Bacharia zherichini Gorochov 1988

G. †Paratshorkuphlebia Sharov 1968

†Paratshorkuphlebia multivenosa Sharov 1968

G. †Pseudotshorkuphlebia Gorochov 1988

†Pseudotshorkuphlebia pritykinae Gorochov 1988

†Pseudotshorkuphlebia sharovi Gorochov 1988

G. †Sharovophlebia Gorochov 1988

†Sharovophlebia angusta Gorochov 1988

G. †Sogutophlebia Gorochov 1988

†Sogutophlebia issykulensis Gorochov 1988

G. †Cantohagla Gorochov 1986

†Cantohagla gracilis Gorochov 1986

Subfm. †Cyrtophyllitinae Zeuner 1935

G. †Archaboilus Martynov 1937

†Archaboilus kisylkiensis Martynov 1937

†Archaboilus martynovi Gorochov 1988

†Archaboilus musicus Gu et al. 2012

†Archaboilus polyneurus Gu et al. 2021

†Archaboilus shurabicus Martynov 1937

†Archaboilus similis Zherikhin 1985

G. †Cyrtophyllites Oppenheim 1888

†Cyrtophyllites cretaceus Gorochov et al. 2006

†Cyrtophyllites rogeri Oppenheim 1888

Invalid names: Cyrtophyllites musicus Handlirsch 1906 [synonym]

G. †Protohagla Zeuner 1962

†Protohagla langi Zeuner 1962

G. †Tasgorosailus Gorochov 1990

†Tasgorosailus orlovskajae Gorochov 1990

G. †Vitimoilus Gorochov 1996

†Vitimoilus captiosus Gorochov 1996

†Vitimoilus ovatus Gu et al. 2017

G. †Dinohagla Gorochov 1986

†Dinohagla corrugata Gorochov 1986

G. †Dolichohagla Gorochov 1986

†Dolichohagla longa Gorochov 1986

G. †Dulcihagla Gorochov 1986

†Dulcihagla beybienkoi Gorochov 1986

†Dulcihagla mistshenkoi Gorochov 1986

G. †Euhagla Gorochov 1986

†Euhagla saurensis Gorochov 1986

G. †Euspilopteron Cockerell 1915

G. †Hagla Giebel 1856

†Hagla gracilis Giebel 1856

Invalid names: Hagla anglica Handlirsch 1906 [synonym], Hagla deleta Giebel 1856 [synonym], Hagla similis Giebel 1856 [synonym]

Invalid names: Haglodes Handlirsch 1906 [synonym]

G. †Haglomorpha Gorochov 1986

†Haglomorpha martynovi Gorochov 1986

Subfm. †Haglopterinae Gorochov 1986

G. †Hagloptera Gorochov 1986

†Hagloptera intermedia Gorochov 1986

Subfm. †Isfaropterinae Martynov 1937

G. †Isfaroptera Martynov 1937

†Isfaroptera grylliformis Martynov 1937

†Isfaroptera yujiagouensis Hong 1983

G. †Laiyangohagla Wang and Liu 1996

†Laiyangohagla beipoziensis Wang and Liu 1996

G. †Lyrohagla Gorochov 1986

†Lyrohagla decipiens Gorochov 1986

†Lyrohagla pravdini Gorochov 1986

†Lyrohagla uvarovi Gorochov 1986

G. †Mesohagla Zhang 1996

†Mesohagla xinjiangensis Zhang 1996

G. †Microhagla Gorochov 1986

†Microhagla minuta Gorochov 1986

G. †Modihagla Gorochov 1986

†Modihagla ovalis Gorochov 1986

G. †Mongolohagla Zherikhin 1985

†Mongolohagla unicolor Zherikhin 1985

G. †Notopamphagopsis Cabrera 1928

†Notopamphagopsis bolivari Cabrera 1928

G. †Parahaglopsis Lin 1992

†Parahaglopsis posteria Lin 1992

G. †Permohagla Gorochov 2021

†Permohagla tomica Gorochov 2021

G. †Prohagla Riek 1954

†Prohagla imperfecta Riek 1955

†Prohagla superba Riek 1954

G. †Proisfaroptera Sharov 1968

†Proisfaroptera martynovi Sharov 1968

G. †Protshorkuphlebia Sharov 1968

†Protshorkuphlebia kirgizica Gorochov 1986

†Protshorkuphlebia similis Gorochov 1986

†Protshorkuphlebia triassica Sharov 1968

G. †Sharovohagla Gorochov 1986

†Sharovohagla plana Gorochov 1986

G. †Sinohagla Lin 1965

†Sinohagla anthoides Lin 1965

†Sinohagla pleioneura Ren 1995

G. †Sonohagla Gorochov 1986

†Sonohagla chopardi Gorochov 1986

†Sonohagla curta Gorochov 1986

†Sonohagla saussurei Gorochov 1986

G. †Tinnihagla Gorochov 1986

†Tinnihagla handlirschi Gorochov 1986

†Tinnihagla zeuneri Gorochov 1986

Subfm. †Triassaginae Gorochov and Maehr 2008

G. †Triassaga Gorochov and Maehr 2008

†Triassaga angusta Gorochov 1986

†Triassaga tshorkuphlebioides Gorochov 1986

Invalid names: Maraga Gorochov 1986 [replaced]

Invalid names: Maraginae Gorochov 1986 [replaced]

G. †Tshorkuphlebia Martynov 1937

†Tshorkuphlebia compressa Martynov 1937

†Tshorkuphlebia shurabica Sharov 1968

G. †Vocohagla Gorochov 1986

†Vocohagla clara Gorochov 1986

†Vocohagla tarbinskyi Gorochov 1986

Subfm. †Voliopinae Gorochov 1986

G. †Asiovoliopus Gorochov 1988

†Asiovoliopus kirgizicus Gorochov 1988

G. †Bacharovoliopus Gorochov 1988

†Bacharovoliopus pulcherrimus Gorochov 1988

G. †Bajanchongoria Gorochov 1988

†Bajanchongoria modesta Gorochov 1988

G. †Euvoliopus Gorochov 1986

†Euvoliopus giganteus Gorochov 1986

G. †Juravoliopus Gorochov 1988

†Juravoliopus sinuatus Martynov 1937

G. †Macrovoliopus Gorochov 1986

†Macrovoliopus declivis Gorochov 1986

G. †Melovoliopus Gorochov 1986

†Melovoliopus fasciatus Gorochov 1986

G. †Mongolovoliopus Gorochov 1988

†Mongolovoliopus sonorus Gorochov 1988

G. †Paravoliopus Gorochov 1986

†Paravoliopus dorsalis Gorochov 1986

G. †Phonovoliopus Gorochov 1986

†Phonovoliopus musicus Gorochov 1986

G. †Phyllovoliopus Gorochov 1986

†Phyllovoliopus insufficiens Gorochov 1986

G. †Platyvoliopus Gorochov 1986

†Platyvoliopus maximus Gorochov 1986

G. †Stenovoliopus Gorochov 1986

†Stenovoliopus elongatus Gorochov 1986

G. †Turkestania Sharov 1968

†Turkestania deviata Sharov 1968

G. †Voliopellus Gorochov 1986

†Voliopellus latus Gorochov 1986

G. †Voliopus Gorochov 1986

†Voliopus ancestralis Gorochov 1986

†Voliopus margueritae Wappler 2001

G. †Zavoliopus Gorochov 1986

†Zavoliopus densus Gorochov 1986

G. †Zeunerophlebia Sharov 1968

†Zeunerophlebia gigas Sharov 1968

Invalid names: Aboilidae Martynov 1925 [synonym], Chifenginae Hong 1982 [empty], Prohaglinae Riek 1954 [synonym], Tshorkuphlebiidae Martynov 1937 [synonym]

Diagnosis

No diagnoses are available

Measurements

No measurements are available

Composition:

chitinsubo

Ontogeny:

moltingsubo

Grouping:

gregarioussubo

Environment:

terrestrialsubo

Locomotion:

fast-movingsubo

Life habit:

ground dwellingsubo

Diet:

herbivoresubo

Vision:

well-developedsubo

Reproduction:

oviparoussubo

Dispersal:

direct/internalsubo

Dispersal 2:

mobilesubo

Created:

2011-07-02 09:58:43

Modified:

2011-07-01 18:58:43

Source: subo = suborder

Reference: Tillyard 1921

Age range: base of the Griesbachian to the top of the Campanian or 251.90200 to 72.10000 Ma

Collections (65 total)

Time interval	Ma	Country or state	Original ID and collection number
Griesbachian	251.902 - 250.5	Russian Federation (Kemerovo)	Permohagla tomica (104713)
Anisian	247.2 - 242.0	Australia (New South Wales)	Prohagla superba (106483)
Ladinian	242.0 - 237.0	Kyrgyzstan (Osh)	Haglomorpha martynovi, Modihagla ovalis, Dulcihagla mistshenkoi, Lyrohagla uvarovi, Sonohagla curta, Tinnihagla zeuneri, Dolichohagla longa, Voliopus ancestralis, Euvoliopus giganteus, Macrovoliopus declivis, Paravoliopus dorsalis, Voliopellus latus, Maraga angusta, Zamaraga reticulata (105007) Proisfaroptera martynovi, Protshorkuphlebia triassica, Cantohagla gracilis, Tinnihagla handlirschi, Maragella reducta (106090) Turkestania deviata, Proisfaroptera martynovi, Protshorkuphlebia triassica, Voliopinae indet., Hagloptera intermedia, Protshorkuphlebia similis, Archihagla tenuis, Dulcihagla beybienkoi, Lyrohagla pravdini, Lyrohagla decipiens, Sonohagla saussurei, Sonohagla chopardi, Microhagla minuta, Dinohagla corrugata, Platyvoliopus maximus, Stenovoliopus elongatus, Zavoliopus densus, Maraga tshorkuphlebioides, Eumaraga madygenica (106088) Zeunerophlebia gigas, Turkestania deviata, Protshorkuphlebia triassica, Archihagla zeuneri, Voliopinae indet., Ahagla mira, Protshorkuphlebia kirgizica, Sharovohagla plana, Vocohagla clara, Vocohagla tarbinskyi, Phonovoliopus musicus, Melovoliopus fasciatus, Phyllovoliopus insufficiens (110763)
Carnian	237.0 - 228.0	Lesotho	Hagla contorta (23075)
Carnian	237.0 - 228.0	Kazakhstan	Paramaraga fusca (113405)
Carnian	237.0 - 228.0	Argentina (La Rioja)	Notopamphagopsis sp. 2, Notopamphagopsis sp. 3, Hagla sp. (122987)
Carnian	237.0 - 228.0	Argentina	Notopamphagopsis bolivari (160458) Notopamphagopsis sp. 1 (123009)
Carnian	237.0 - 228.0	South Africa (Eastern Cape)	Hagla contorta, Afrohagla contorta (23142) Hagla sp. 1 (23259) Hagla sp. 2 (23133) Haglidae indet. (23263) Voliopus margueritae, Hagla contorta (23131)
Norian	228.0 - 208.5	Kazakhstan	Euhagla saurensis (105055)
Norian	228.0 - 208.5	Australia (Queensland)	Prohagla sp., Prohagla imperfecta (106484)
Rhaetian	208.5 - 201.3	United Kingdom (England)	Chauliodes sp. 10, Chauliodes sp. 6 (115223) Hagla gracilis (113961)
Rhaetian - Early/Lower Hettangian	208.5 - 196.5	United Kingdom (England)	Chauliodes sp. 8, Insecta indet. (124688)
Early/Lower Jurassic	201.4 - 174.7	China (Nei Mongol)	Archaeohagla sinensis (156316) Sinohagla anthoides (156315)
Early/Lower Hettangian	201.3 - 196.5	United Kingdom (England)	Hagla gracilis (114682 123983)
Planorbis	201.3 - 196.5	United Kingdom (England)	Hagla gracilis (123982)
Hettangian - Sinemurian	201.3 - 190.8	China (Xinjiang)	Mesohagla xinjiangensis (103480)
Hettangian - Sinemurian	201.3 - 190.8	Kyrgyzstan (Tonskiy)	Archaboilus martynovi (105084) Archaboilus sinuatus, Asiovoliopus kirgizicus (105087) Paratshorkuphlebia multivenosa (105086)
Sinemurian	199.3 - 190.8	Kyrgyzstan (Tonskiy)	Paratshorkuphlebia multivenosa (113412) Sogutophlebia issykulensis (122951)
Obtusum	196.5 - 189.6	United Kingdom (England)	Hagla gracilis (114637) Protohagla langi (114638)
Pliensbachian	190.8 - 182.7	Kyrgyzstan	Archaboilus kisylkiensis (114701)
Toarcian	182.7 - 174.1	Kyrgyzstan (Batken)	Pseudotshorkuphlebia sharovi (105124) Pseudotshorkuphlebia sharovi, Pseudotshorkuphlebia pritykinae (105003) Tshorkuphlebia shurabica (105089 110817 113411)
Toarcian	182.7 - 174.1	Russian Federation (Irkutsk)	Angarohagla variegata (104946) Archaboilus similis (113471) Haglidae indet. (104996)
Toarcian	182.7 - 174.1	Kyrgyzstan	Archaboilus shurabicus (114707) Archaboilus sinuatus (114706) Isfaroptera grylliformis (114708) Tshorkuphlebia compressa (114702)
Bajocian - Bathonian	170.3 - 166.1	China (Anhui)	Haglidae indet. (106695)
Bajocian - Bathonian	170.3 - 166.1	Mongolia (Bayan-Khongor)	Microvoliopus bacharicus, Mongolovoliopus sonorus, Bacharovoliopus pulcherrimus, Bajanchongoria modesta, Bacharia zherichini (114615)
Bathonian	168.3 - 166.1	Mongolia	Haglidae indet., Mongolohagla unicolor (119621)
Callovian - Oxfordian	166.1 - 157.3	China (Nei Mongol)	Archaboilus musicus, Archaboilus polyneurus (113365)
Callovian - Oxfordian	166.1 - 157.3	China (Liaoning)	Isfaroptera yujiagouensis (154730)
Late/Upper Jurassic - Early/Lower Cretaceous	161.5 - 100.5	Kazakhstan	Tasgorosailus orlovskajae (151964)
Early/Lower Tithonian	150.8 - 145.0	Germany (Bayern)	Cyrtophyllites musicus (123985)
Early/Lower Tithonian	150.8 - 145.0	Germany (Bavaria)	Cyrtophyllites rogeri (123984)
Early/Lower Tithonian	150.8 - 145.0	Germany	Cyrtophyllites rogeri (121197)
Early/Lower Berriasian	145.0 - 140.2	United Kingdom (England)	Cyrtophyllites cretaceus (104002)
Late/Upper Berriasian	145.0 - 140.2	United Kingdom (England)	Cyrtophyllites cretaceus (224570)
Late/Upper Hauterivian	136.4 - 130.0	China (Hebei)	Vitimoilus ovatus (113484)
Late/Upper Barremian	130.0 - 125.45	China (Hebei)	Sinohagla pleioneura (107001)
Late/Upper Barremian	130.0 - 125.45	China (Liaoning)	Vitimoilus ovatus (184476)
Aptian	125.0 - 113.0	Russian Federation (Buryatia)	Vitimoilus captiosus (122549)
Aptian	125.0 - 113.0	China (Jilin)	Parahaglopsis posteria (172387)
Aptian	125.0 - 113.0	China (Shandong)	Laiyangohagla beipoziensis (164688)
Campanian	83.6 - 72.1	South Africa (Northern Cape)	Hagla sp. (55737)