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Blattula

Insecta - Blattodea - Blattulidae

Taxonomy
Blattula was named by Handlirsch (1906). It is the type genus of Blattulidae.

It was assigned to Mesoblattinidae by Handlirsch (1906), Martynov (1937), Bode (1953) and Hong (1986); and to Blattulidae by Vishnyakova (1982), Lin (1986), Vršanský (2004), Vršanský and Ansorge (2007), Vršanský (2008), Barna (2014) and Vršanský (2020).

Species
B. aberrans, B. anuniversala, B. apicifurca, B. bacharensis, B. brevicaudata, B. choutinensis, B. colorata, B. ctinoida, B. cubitalis, B. curvula, B. delicatula, B. discors, B. disjuncta, B. dubia, B. elongata, B. extensa, B. flamma, B. geinitzi, B. hymena, B. iensis, B. incompleta, B. intercalata, B. isotoma, B. kellos, B. kiensis, B. lanceolata, B. langfeldti (syn. Metablattula lipomena, Blattina (Mesoblattina) dobbertinensis, B. pusillima, Peloblattula oligoneura, Englyptoblatta elegantula, B. debilis, Chiloblattula subcostalis, Parablattula simplicissima, Mesoblattula ala, B. riparia, Chiloblattula longipennis, B. vicina, Ectinoblattula medialis, B. brunneri, Blattina nana, B. hattorfensis, Blattina incerta, Mesoblattula geinitziana, Mesoblattula dobbertiniana, B. ancilla, Chiloblattula simplex, B. acutipennis, Dipluroblattina scudderi, Parablattula reticulata), B. liaoningensis, B. mikro, B. mini, B. mirta, B. mongolica, B. pachohymena, B. piveteaui, B. platypa, B. prestwichii, B. rudis, B. shurabica, B. sibirica, B. similis, B. sincera, B. tchakrasensis, B. tuodianensis, B. turanica, B. udensis, B. universala, B. velika, B. vidlickai, B. vulgara, B. willmotti, B. zaoshangensis, Mesoblattula chengdeensis

Synonyms
Synonymy list
YearName and author
1906Mesoblattula Handlirsch p. 430
1906Blattula Handlirsch p. 431
1937Mesoblattula Martynov p. 203
1937Blattula Martynov p. 209
1939Parablattula Handlirsch p. 60
1939Peloblattula Handlirsch p. 60
1939Chiloblattula Handlirsch p. 61
1939Metablattula Handlirsch p. 61
1939Ectinoblattula Handlirsch p. 62
1953Blattula Bode
1953Englyptoblatta Bode
1982Ectinoblattula Vishnyakova p. 67
1982Metablattula Vishnyakova p. 67
1982Peloblattula Vishnyakova p. 67
1982Blattula Vishnyakova p. 69
1982Mesoblattula Vishnyakova p. 69
1982Parablattula Vishnyakova p. 74
1986Blattula Hong
1986Mesoblattula Lin p. 48
1986Blattula Lin p. 71
1995Parablattula Ren p. 54
2003Mesoblattula Zhang and Hong p. 453
2004Blattula Vršanský p. 460
2007Blattula Vršanský and Ansorge p. 114
2008Blattula Vršanský p. 40
2014Blattula Barna p. 666
2020Blattula Vršanský p. 50

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RankNameAuthor
kingdomAnimalia()
—Bilateria
—EubilateriaAx 1987
—Protostomia
—Ecdysozoa
—Panarthropoda
phylumArthropodaLatreille 1829
superclassHexapodaLatreille 1825
classInsecta
—Dicondylia
—Paranotalia
RankNameAuthor
subclassPterygota()
—NeopterygotaCrampton 1924
infraclassNeopteraMartynov 1923
—PolyneopteraMartynov 1938
—Holopandictyoptera
superorderDictyopteraLatreille 1829
orderBlattodea(Latreille 1810)
—Solumblattodea
superfamilyCorydioidea
familyBlattulidae
genusBlattula

If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.

G. †Blattula Handlirsch 1906
show all | hide all
†Blattula aberrans Vishnyakova 1982
†Blattula anuniversala Vršanský 2020
†Blattula apicifurca Lin 1986
†Blattula bacharensis Vršanský 2020
†Blattula brevicaudata Vishnyakova 1968
†Blattula choutinensis Vršanský 2008
†Blattula colorata Vishnyakova 1982
†Blattula ctinoida Lin 1986
†Blattula cubitalis Vishnyakova 1982
†Blattula curvula Ren 1995
†Blattula delicatula Ren 1995
†Blattula discors Barna 2014
†Blattula disjuncta Scudder 1886
†Blattula dubia Handlirsch 1939
†Blattula elongata Martynov 1937
†Blattula extensa Vishnyakova 1982
†Blattula flamma Vršanský 2020
†Blattula geinitzi Handlirsch 1906
†Blattula hymena Lin 1986
†Blattula iensis Vishnyakova 1982
†Blattula incompleta Giebel 1856
†Blattula intercalata Handlirsch 1939
†Blattula isotoma Vishnyakova 1982
†Blattula kellos Zhang 1986
†Blattula kiensis Martynov 1937
†Blattula lanceolata Vishnyakova 1982
†Blattula langfeldti Geinitz 1880
+
Invalid names: Blattina (Mesoblattina) dobbertinensis Geinitz 1884 [synonym], Blattina incerta Geinitz 1884 [synonym], Blattina nana Geinitz 1883 [synonym], Blattula acutipennis Handlirsch 1939 [synonym], Blattula ancilla Handlirsch 1906 [synonym], Blattula brunneri Handlirsch 1939 [synonym], Blattula debilis Handlirsch 1906 [synonym], Blattula hattorfensis Bode 1953 [synonym], Blattula pusillima Handlirsch 1906 [synonym], Blattula riparia Handlirsch 1939 [synonym], Blattula vicina Handlirsch 1939 [synonym], Chiloblattula longipennis Handlirsch 1939 [synonym], Chiloblattula simplex Handlirsch 1939 [synonym], Chiloblattula subcostalis Handlirsch 1939 [synonym], Dipluroblattina scudderi Geinitz 1888 [synonym], Ectinoblattula medialis Handlirsch 1939 [synonym], Englyptoblatta elegantula Bode 1953 [synonym], Mesoblattula ala Handlirsch 1939 [synonym], Mesoblattula dobbertiniana Handlirsch 1906 [synonym], Mesoblattula geinitziana Handlirsch 1906 [synonym], Metablattula lipomena Handlirsch 1939 [synonym], Parablattula reticulata Handlirsch 1939 [synonym], Parablattula simplicissima Handlirsch 1939 [synonym], Peloblattula oligoneura Handlirsch 1939 [synonym]
†Blattula mikro Vršanský 2020
†Blattula mini Vršanský 2020
†Blattula mirta Zhang 1986
†Blattula mongolica Vršanský 2004
†Blattula piveteaui Laurentiaux 1946
†Blattula platypa Ren 1995
†Blattula prestwichii Scudder 1886
†Blattula rudis Ren 1995
†Blattula shurabica Martynov 1937
†Blattula sibirica Vishnyakova 1985
†Blattula similis Vishnyakova 1982
†Blattula sincera Lin 1985
†Blattula tchakrasensis Laurentiaux 1946
†Blattula tuodianensis Zhang and Hong 2003
†Blattula turanica Martynov 1937
†Blattula udensis Vishnyakova 1982
†Blattula universala Vršanský 2020
†Blattula velika Vršanský 2020
†Blattula vidlickai Vršanský 2004
†Blattula vulgara Vršanský 2020
†Blattula willmotti Martin 2010
Invalid names: Chiloblattula Handlirsch 1939 [synonym], Ectinoblattula Handlirsch 1939 [synonym], Englyptoblatta Bode 1953 [synonym], Mesoblattula Handlirsch 1906 [synonym], Metablattula Handlirsch 1939 [synonym], Parablattula Handlirsch 1939 [synonym], Peloblattula Handlirsch 1939 [synonym]
Diagnosis
No diagnoses are available
Measurements
No measurements are available
Composition: chitinc
Environment: terrestrialc
Locomotion: actively mobilep
Created: 2017-04-17 09:56:18
Modified: 2017-04-17 09:56:18
Source: c = class, p = phylum
References: Kiessling 2004, Bush and Bambach 2015

Age range: base of the Rhaetian to the top of the Albian or 208.50000 to 100.50000 Ma

Collections (55 total)


Time interval Ma Country or state Original ID and collection number
Rhaetian208.5 - 201.3Turkey B. tchakrasensis, Mesoblattula piveteaui (152783)
Rhaetian208.5 - 201.3United Kingdom (England) Insecta indet. (113961)
Hettangian201.3 - 199.3China (Hunan) Mesoblattula apicifurca (155338)
Sinemurian199.3 - 190.8China (Hunan) B. zaoshangensis (155353 155356 155358) Mesoblattula kiensis (155357)
Pliensbachian190.8 - 182.7Kyrgyzstan B. sp., Mesoblattula kiensis, B. turanica, B. elongata (114701)
Early/Lower Toarcian183.0 - 182.0Germany (Mecklenburg-Vorpommern) B. langfeldti, B. dubia (114993)
Early/Lower Toarcian183.0 - 182.0Germany (Niedersachsen) B. hattorfensis, Englyptoblatta elegantula (107552) B. langfeldti (117795)
Early/Lower Toarcian183.0 - 182.0Germany Blattina langfeldti, Blattina nana, Blattina dobbertinensis, Blattina incerta, Dipluroblattina scudderi, B. ancilla, B. geinitzi, B. debilis, B. pusillima, Parablattula reticulata, Mesoblattula ala, B. riparia, B. intercalata, B. brunneri, B. vicina, Peloblattula oligoneura, Parablattula simplicissima, Metablattula lipomena, Chiloblattula simplex, Chiloblattula subcostalis, Chiloblattula longipennis, Ectinoblattula medialis, Chiloblattula dubia, Mesoblattina dobbertinensis, Mesoblattina dobbertinensis (123987) B. acutipennis (122986) B. langfeldti, B. dubia (114991)
Falciferum183.0 - 182.0Germany B. langfeldti (151355)
Toarcian182.7 - 174.1Australia (Western Australia) B. willmotti (114239)
Toarcian182.7 - 174.1China (Guangxi) B. pachohymena, B. hymena, Mesoblattula sincera (151540)
Toarcian182.7 - 174.1Kyrgyzstan Mesoblattula shurabica (114706)
Toarcian182.7 - 174.1Russian Federation (Irkutsk) B. extensa, B. iensis, Parablattula cubitalis (104946) B. similis, Parablattula isotoma (104996) Mesoblattula colorata (104995)
Toarcian - Aalenian182.7 - 170.3China (Hunan) B. ctinoida (155351)
Bajocian - Bathonian170.3 - 166.1Russian Federation (Krasnoyarsk) B. aberrans (104947) Chiloblattula sibirica (118010)
Bajocian - Bathonian170.3 - 166.1China (Anhui) B. pachohymena, Mesoblattula kiensis (106695)
Bajocian - Bathonian170.3 - 166.1Mongolia (Bayan-Khongor) B. flamma (215956) B. flamma, B. universala (121709) B. mini (215951) B. velika, B. vulgara, B. mini, B. mikro, B. anuniversala (124340)
Callovian - Oxfordian166.1 - 157.3China (Hebei) Mesoblattula chengdeensis (154837) Mesoblattula mirta, B. kellos (157564) Parablattula curvula (107000)
Callovian - Oxfordian166.1 - 157.3China (Liaoning) B. liaoningensis (110855)
Callovian - Oxfordian166.1 - 157.3Kazakhstan (Chimkent) B. brevicaudata (106101)
Oxfordian163.5 - 157.3Russian Federation (Buryat) B. lanceolata (104949)
Oxfordian - Kimmeridgian163.5 - 152.1China (Yunnan) Mesoblattula sp., Mesoblattula tuodianensis (164858)
Oxfordian - Kimmeridgian163.5 - 152.1Russian Federation (Buryat) Parablattula udensis (104950)
Tithonian152.1 - 145.0Mongolia (Dund-Gov’ Aymag) B. choutinensis (117697 117698)
Tithonian152.1 - 145.0Mongolia (Bayan-Khongor) B. bacharensis (122322)
Tithonian152.1 - 145.0Mongolia B. mongolica, B. vidlickai (104748 121393 121394 125981) B. vidlickai (118966 152133 165034)
Late/Upper Berriasian145.0 - 140.2United Kingdom (England) Nannoblattina prestwichii (133535)
Berriasian145.0 - 139.8United Kingdom (England) Rithma disjuncta (133536)
Late/Upper Barremian130.0 - 125.45China (Hebei) B. platypa, B. delicatula (107001)
Barremian129.4 - 125.0Russian Federation (Chita) B. discors (105968)
Late/Upper Aptian122.46 - 113.0China (Beijing Shi) B. rudis (106965)
Albian113.0 - 100.5South Korea B. hymena (184784) B. sp. (184785) Mesoblattula sp. (184788)