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Trigonoptera

Insecta - Odonata

Taxonomy
Trigonoptera was named by Bechly (1996). It is extant.

It was assigned to Odonata by Fleck et al. (2003); and to Anisopteromorpha by Bechly (2007).

Subtaxa
Pananisoptera, Stenophlebioptera
Sister taxa
Pananisoptera, Stenophlebioptera

Synonymy list
YearName and author
1996Trigonoptera Bechly
2003Trigonoptera Fleck et al.
2007Trigonoptera Bechly

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RankNameAuthor
kingdomAnimalia()
—Bilateria
—EubilateriaAx 1987
—Protostomia
—EcdysozoaAguinaldo et al. 1997
—Panarthropoda
phylumArthropodaLatreille 1829
—Deuteropoda
—Mandibulata
—PancrustaceaZrzavý and Štys 1997
—AltocrustaceaRegier et al. 2010
superclassAllotriocarida
—Athalassocarida
—LabiocaridaSchwentner et al. 2017
superclassHexapodaLatreille 1825
—Cercofilata
classInsecta
—Dicondylia
—Paranotalia
subclassPterygota()
infraclassHydropalaeopteraRohdendorf 1969
RankNameAuthor
—Euhydropalaeoptera
—OdonatopteraMartynov 1932
—Palaeodonatoptera
—Plesiodonatoptera
—Apodonatoptera
—Paneodonatoptera
—Neodonatoptera
—Euodonatoptera
—OdonatocladaBechly 1999
—Nodialata
—Discoidalia
—Stigmoptera
—Lobodonata
—Panodonata
orderOdonata
—Epiproctophora
—EuepiproctophoraBechly 1998
—Anisopteromorpha
—Trigonoptera
—Trigonoptera

If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.

Unr. Trigonoptera Bechly 1996
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Unr. Pananisoptera Bechly 1996
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Fm. †Liassogomphidae Tillyard 1935
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G. †Chrysogomphus Ren 1994
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G. †Elattogomphus Bode 1953
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†Elattogomphus latus Bode 1953
G. †Heterothemis Handlirsch 1906
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†Heterothemis brodiei Brodie 1845
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Invalid names: Gomphites (Palaeogomphus) paeneparallelus Bode 1953 [synonym], Heterophlebia propinqua Bode 1905 [synonym], Heterothemis germanica Handlirsch 1906 [synonym]
Invalid names: Gomphites Handlirsch 1920 [synonym], Liassogomphus Cowley 1934 [synonym], Palaeogomphus Handlirsch 1939 [synonym]
G. †Necygomphus Cowley 1942
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†Necygomphus brunswigiae Handlirsch 1939
Invalid names: Necrogomphus Handlirsch 1939 [replaced]
G. †Phthitogomphus Cowley 1942
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†Phthitogomphus angulatus Handlirsch 1939
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Invalid names: Gomphites (Palaeogomphus) aranea Bode 1953 [synonym]
Invalid names: Paragomphus Handlirsch 1939 [replaced]
G. †Proinogomphus Handlirsch 1939
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†Proinogomphus bodei Handlirsch 1939
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Invalid names: Proinogomphus aequalicellatus Bode 1953 [synonym], Proinogomphus bicellatus Bode 1953 [synonym]
Invalid names: Proinogomphus Cowley 1942 [objective synonym]
Invalid names: Gomphitidae Tillyard 1925 [replaced]
Unr. Neoanisoptera Bechly 1998
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Fm. †Aeschnidiidae Handlirsch 1906 [small darner]
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G. †Aegyptidium Schlüter and Hartung 1982
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†Aegyptidium aburasiensis Schlüter and Hartung 1982
G. †Aeschnidiella Zalessky 1953
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†Aeschnidiella kabanovi Zalessky 1953
G. †Aeschnidiopsis Tillyard 1918
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G. †Aeschnidium Westwood 1854
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†Aeschnidium antiquum Brodie 1845
†Aeschnidium bubas Westwood 1854
Invalid names: Estemoa Giebel 1856 [objective synonym], Neimenggolgomphus Hong 1982 [synonym]
G. †Angloaeschnidium Fleck and Nel 2003
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†Angloaeschnidium lacaui Fleck and Nel 2003
†Angloaeschnidium montreuili Fleck and Nel 2003
†Angloaeschnidium toyei Fleck and Nel 2003
G. †Bergeriaeschnidia Nel et al. 1996
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†Bergeriaeschnidia inexpectata Nel et al. 1996
G. †Brunetaeschnidium Fleck and Nel 2003
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G. †Brunneaeschnidia Hong 1982
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G. †Cooperaeschnidium Fleck and Nel 2003
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†Cooperaeschnidium durandi Fleck and Nel 2003
G. †Coramaeschnidium Fleck and Nel 2003
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†Coramaeschnidium minimum Fleck and Nel 2003
G. †Cratoaeschnidium Nel et al. 2022
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†Cratoaeschnidium martinsnetoi Nel et al. 2022
G. †Dakotaeschnidium Nel 2021
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G. †Delclosaeschidium Fleck and Nel 2003
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†Delclosaeschidium magnum Fleck and Nel 2003
G. †Diastatopaeschnidium Fleck and Nel 2003
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†Diastatopaeschnidium reneeheiko Fleck and Nel 2003
G. †Dracontaeschnidium Zhang and Zhang 2001
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†Dracontaeschnidium orientale Zhang and Zhang 2001
G. †Gansuaeschnidia Zhang et al. 2022
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†Gansuaeschnidia hongi Zhang et al. 2022
G. †Gigantoaeschnidium Nel and Martínez-Delclòs 1993
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†Gigantoaeschnidium ibericus Nel and Martínez-Delclòs 1993
G. †Iberoaeschnidium Nel and Martínez-Delclòs 1993
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†Iberoaeschnidium conquensis Nel and Martínez-Delclòs 1993
G. †Jarzembowskiaeschnidium Fleck and Nel 2003
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†Jarzembowskiaeschnidium polandi Fleck and Nel 2003
G. †Kesseleraeschnidium Fleck and Nel 2003
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†Kesseleraeschnidium simonae Fleck and Nel 2003
G. †Kimmeridgebrachypteraeschnidium Fleck and Nel 2003
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G. †Leptaeschnidium Pritykina 1977
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†Leptaeschnidium latum Pritykina 1977
G. †Linaeschnidium Huang et al. 2009
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†Linaeschnidium sinensis Huang et al. 2009
G. †Lithoaeschnidium Nel and Martínez-Delclòs 1993
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†Lithoaeschnidium viohli Nel and Martínez-Delclòs 1993
G. †Lleidoaeschnidium Nel and Martínez-Delclòs 1993
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†Lleidoaeschnidium maculatum Fleck and Nel 2003
†Lleidoaeschnidium valloryi Nel and Martínez-Delclòs 1993
G. †Malmaeschnidium Nel and Martínez-Delclòs 1993
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†Malmaeschnidium mayeri Nel and Martínez-Delclòs 1993
G. †Misofaeschnidium Fleck and Nel 2003
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G. †Nannoaeschnidium Nel and Martínez-Delclòs 1993
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†Nannoaeschnidium pumilio Nel and Martínez-Delclòs 1993
G. †Rossaeschnidium Fleck and Nel 2003
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†Rossaeschnidium patriciae Fleck and Nel 2003
G. †Santanoptera Martill and Nel 1996
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†Santanoptera gabbotti Martill and Nel 1996
G. †Sinaeschnidia Hong 1965
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†Sinaeschnidia huzhouensis Zhou and Wei 1980
†Sinaeschnidia martinezdelclosi Fleck and Nel 2003
G. †Sinostenophlebia Hong 1984
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G. †Solnhofenia Bechly 2000
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†Solnhofenia stoebeneri Bechly 2000
G. †Stylaeschnidium Zhang and Zhang 2001
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†Stylaeschnidium rarum Zhang and Zhang 2001
G. †Tauropteryx Pritykina 1993
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†Tauropteryx krassilovi Pritykina 1993
G. †Urogomphus Handlirsch 1906
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†Urogomphus giganteus Germar 1839
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Invalid names: Petalura eximia Hagen 1862 [synonym], Petalura latialata Hagen 1862 [objective synonym]
G. †Wightonia Carle 1990
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†Wightonia araripina Carle and Wighton 1990
Invalid names: Hebeiaeschnidia Hong 1982 [nomen dubium], Libellula abscissa Hagen 1862 [nomen dubium], Neimengogomphus Hong 1982 [nomen dubium], Neimengogomphus dongwugaiensis Hong 1982 [nomen dubium], Pseudosamarura Lin 1976 [nomen dubium], Pseudosamarura largina Lin 1976 [nomen dubium], Sonidae Pritykina 1986 [synonym], Yixiangomphus Lin 1986 [nomen dubium], Yixiangomphus labius Lin 1976 [nomen dubium]
Subor. Anisoptera Selys-Longchamps and Hagen 1854 [true dragonfly]
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Fm. †Cratopetaliidae Bechly 2010
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G. †Cratopetalia Bechly 2010
Fm. Epigomphidae Fraser 1934
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G. Macrogomphus Selys-Longchamps 1857
Unr. Euanisoptera Bechly 1996
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Unr. Aeshnoptera Bechly 1996
Unr. Exophytica Bechly 1996
Fm. Hageniidae Tillyard and Fraser 1940
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G. †Cratohagenius Bechly 2010
Fm. †Henrotayiidae Fleck et al. 2003
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G. †Henrotayia Fleck et al. 2003
G. †Ixtahua Feldmann et al. 1998
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†Ixtahua benjamini Feldmann et al. 1998
Fm. †Juragomphidae Nel et al. 2001
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G. †Juragomphus Nel et al. 2001
G. †Oxfordgomphus Huang and Nel 2023
G. †Kazakhophlebiella Nel et al. 1998
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†Kazakhophlebiella aktassica Pritykina 1968
Fm. †Libanoaeshnidae Azar et al. 2024
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G. †Libanoaeshna Azar et al. 2024
Superfm. Libelluloidea
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G. †Condalia Whalley and Jarzembowski 1985
Fm. Macromiidae Tillyard 1917
Invalid names: Libellula cassandra Hagen 1863 [nomen dubium], Libellula ceres Hagen 1863 [nomen dubium]
Fm. †Magnathemidae Bechly 2010
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G. †Magnathemis Bechly 2010
Fm. †Megaphlebiidae Bechly 2010
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G. †Megaphlebia Bechly 2010
G. †Necrogomphus Campion 1923
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Invalid names: Mesogomphus Handlirsch 1906 [replaced]
Fm. †Nothomacromiidae Carle 1995
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G. †Nothomacromia Carle 1995
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Invalid names: Conan Martins-Neto 1998 [synonym], Pseudomacromia Carle and Wighton 1990 [replaced]
Invalid names: Pseudomacromiidae Carle and Wighton 1990 [replaced]
Unr. Petalurida Bechly 1996
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Fm. Petaluridae Needham 1903 [grayback dragonfly]
Unr. Petalurodea Bechly 1996
Fm. †Protolindeniidae Handlirsch 1906
G. †Pritykiniella Nel et al. 1998
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†Pritykiniella deichmuelleri Pritykina 1968
Fm. †Proterogomphidae Bechly et al. 1998
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Subfm. †Cordulagomphinae Carle and Wighton 1990
G. †Lingomphus Vernoux et al. 2010
G. †Proterogomphus Bechly et al. 1998
Invalid names: Aeschna eudore Heer 1849 [nomen dubium], Aeschna parkinsoni Selys-Longchamps 1850 [nomen dubium], Aeschnidioidea Carle and Wighton 1990 [empty], Cercus Hong 1982 [nomen dubium], Cercus clavus Hong 1982 [nomen dubium], Cordulia scheuchzeri Massalongo 1856 [nomen dubium], Cymatophlebia mongolica Cockerell 1924 [nomen dubium], Dissurus Hong 1982 [nomen dubium], Dissurus liauyuanensis Hong 1982 [nomen dubium], Dissurus qinquanensis Hong 1982 [nomen dubium], Ictinogomphus fur Hagen 1863 [nomen dubium], Libellula calypso Heer 1849 [nomen dubium], Libellula doris Heer 1849 [nomen dubium], Libellula eurynome Heer 1849 [nomen dubium], Libellula melobasis Heer 1849 [nomen dubium], Libellula perse Heer 1849 [nomen dubium], Libellula thetis Heer 1849 [nomen dubium], Libellula thoe Heer 1849 [nomen dubium], Miopetalura orientalis Hong 1985 [nomen dubium], Stenogomphus scudderi Cockerell 1921 [nomen dubium]
Unr. †Stenophlebioptera Bechly 1996
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Fm. †Gondvanogomphidae Bechly 1996
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G. †Gondvanogomphus Schlüter and Hartung 1982
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†Gondvanogomphus bartheli Schlüter and Hartung 1982
Fm. †Juraheterophlebiidae Fleck et al. 2003
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G. †Juraheterophlebia Fleck et al. 2003
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†Juraheterophlebia cancellosa Huang and Nel 2017
†Juraheterophlebia kazakhstanensis Fleck et al. 2003
†Juraheterophlebia sinica Nel et al. 2014
Fm. †Liassostenophlebiidae Fleck et al. 2003
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G. †Liassostenophlebia Fleck et al. 2003
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†Liassostenophlebia germanica Fleck et al. 2003
Superfm. †Stenophlebioidea Needham 1903
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Fm. †Prostenophlebiidae Fleck et al. 2003
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G. †Prostenophlebia Nel and Martínez-Delclòs 1993
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†Prostenophlebia jurassica Nel and Martínez-Delclòs 1993
Fm. †Stenophlebiidae Needham 1903
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G. †Burmastenophlebia Huang et al. 2018
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†Burmastenophlebia flecki Huang et al. 2018
G. †Cratostenophlebia Bechly 2007
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G. †Cretastenophlebia Fleck et al. 2003
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†Cretastenophlebia jiuquanensis Zheng et al. 2018
†Cretastenophlebia mongolica Fleck et al. 2003
G. †Gallostenophlebia Nel et al. 2015
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†Gallostenophlebia incompleta Nel et al. 2015
G. †Hispanostenophlebia Fleck et al. 2003
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†Hispanostenophlebia barremiana Fleck et al. 2003
G. †Liaostenophlebia Zheng et al. 2016
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†Liaostenophlebia yixianensis Zheng et al. 2016
G. †Mesostenophlebia Fleck et al. 2003
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†Mesostenophlebia anglicana Fleck et al. 2003
G. †Reschiostenophlebia Bechly 2019
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G. †Stenophlebia Hagen 1866
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†Stenophlebia corami Nel and Jarzembowski 1996
†Stenophlebia eichstaettensis Nel et al. 1993
†Stenophlebia karatavica Pritykina 1968
†Stenophlebia latreillei Germar 1839
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Invalid names: Heterophlebia aequalis Hagen 1862 [synonym]
†Stenophlebia liaoningensis Zheng et al. 2016
†Stenophlebia phryne Hagen 1862
†Stenophlebia rolfhuggeri Bechly et al. 2003
†Stenophlebia ryonsangensis Won et al. 2022
G. †Yixianstenophlebia Nel and Huang 2015
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†Yixianstenophlebia magnifica Nel and Huang 2015
Invalid names: Stenophlebia lithographica Giebel 1857 [nomen dubium]
Diagnosis
No diagnoses are available
Measurements
No measurements are available
Composition: chitino
Form: roller-shapedo
Environment: freshwater, terrestrialo
Locomotion: fast-movingo
Life habit: volanto
Diet: carnivoreo
Diet 2: insectivoreo
Vision: well-developedo
Dispersal: watero
Created: 2011-03-07 03:41:10
Modified: 2011-03-06 10:41:10
Source: o = order
Reference: Ren and Guo 1996

Age range

Maximum range based only on fossils: base of the Falciferum to the top of the Holocene or 184.20000 to 0.00000 Ma
Minimum age of oldest fossil (stem group age): 182.9 Ma

Collections (404 total)

Oldest occurrences

Time interval Ma Country or state Original ID and collection number
Early/Lower Toarcian184.2 - 182.9Luxembourg Phthitogomphus angulatus (106698) Proinogomphus kreuzerorum (195801)
Early/Lower Toarcian184.2 - 182.9Germany (Mecklenburg-Vorpommern) Liassogomphidae indet. 1, Liassogomphidae indet. 2 (114993)
Early/Lower Toarcian184.2 - 182.9Germany (Baden-Würtemberg) Liassogomphus brodiei (117796)
Early/Lower Toarcian184.2 - 182.9Germany (Niedersachsen) Gomphites brodiei, Gomphites aranea, Proinogomphus bodei, Proinogomphus bicellatus, Elattogomphus latus (105002) Gomphites brodiei, Paragomphus angulatus (107321) Gomphites paeneparallelus (107556) Gomphites propinquus (107343 107344) Necrogomphus brunswigae (107341) Proinogomphus aequalicellatus (107553)
Early/Lower Toarcian184.2 - 182.9United Kingdom (Gloucestershire) Aeschna brodiei (154308)
Early/Lower Toarcian184.2 - 182.9Germany Heterophlebia buckmanni (123987)
Falciferum184.2 - 182.9United Kingdom (England) Gomphites brodiei (154307) Heterothemis brodiei (234541 234546) Heterothemis sp. (234544) Heterothemis sp., Heterothemis brodiei (234543) Liassogomphidae indet. (132646)
Falciferum184.2 - 182.9Luxembourg Henrotayia marci, Phthitogomphus angulatus (105427)
Falciferum184.2 - 182.9Germany Heterothemis brodiei (151353) Liassostenophlebia germanica (105428)
Toarcian184.2 - 174.7Switzerland Liassogomphus brodiei (97601)
Bathonian168.2 - 165.3United Kingdom (Oxfordshire) Libellula westwoodii (140495)
Callovian165.3 - 161.5China (Henan) Oxfordgomphus trescellulae (202211)
Callovian - Oxfordian165.3 - 154.8China (Nei Mongol) Daohugoulibellula lini, Juralibellula ningchengensis, Cymatophlebia daohugouensis, Sinacymatophlebia mongolica, Linqibinia panae, Sinomesuropetala daohugensis, Propecymatophlebia magnifica (113371) Juraheterophlebia cancellosa (189820) Juraheterophlebia sinica (201985) Sinocymatophlebiella hasticercus (113366)
Callovian - Oxfordian165.3 - 154.8Kazakhstan (Chimkent) Juraheterophlebia kazakhstanensis, Juragomphus karatauensis, Paracymatophlebia splendida (105426) Protolindenia aktassica, Cymatophlebiella euryptera (105970) Stenophlebia karatavica, Aktassia magna, Cymatophlebiella euryptera (105971) Stenophlebia karatavica, Protolindenia aktassica, Aktassia magna (106102) Stenophlebia karatavica, Protolindenia aktassica, Protolindenia deichmuelleri, Mesuropetala costalis, Mesuropetala auliensis (106101)
Early/Lower Kimmeridgian154.8 - 152.21Germany (Baden-Wurttemberg) Cymatophlebia herrienae (105965) Cymatophlebia suevica (105964)
Late/Upper Kimmeridgian152.21 - 149.2United Kingdom (England) Kimmeridgebrachypteraeschnidium etchesi (106418)
Late/Upper Kimmeridgian152.21 - 149.2Germany (Swabian Alb) Libellulium longialatum, Misofaeschnidium densum, Urogomphus giganteus (106402) Stenophlebia rolfhuggeri (105835)
Early/Lower Tithonian149.2 - 145.06Germany (Bavaria) Aeschnogomphus kuempeli (117532) Aeschnogomphus sp. (117533) Calopteryx lithographica, Aeschna multicellulosa (152880) Cymatophlebia longialata (152884) Protolindenia viohli, Cymatophlebia longialata, Malmaeschnidium mayeri (152603) Protolindenia wittei, Aeschnogomphus sp., Aeschnogomphus intermedius, Cymatophlebia longialata, Eumorbaeschna jurassica, Mesuropetala muensteri, Mesuropetala koehleri, Urogomphus giganteus (123994) Stenophlebia amphitrite, Heterophlebia phryne, Stenophlebia latreillei, Stenophlebia aequalis, Prostenophlebia jurassica, Protolindenia wittei, Anax buchi, Aeschnogomphus intermedius, Juracordulia schiemenzi, Nannogomphus bavaricus, Cymatophlebia longialata, Cymatophlebia pumilio, Cymatophlebia kuempeli, Eumorbaeschna jurassica, Aeschnopsis perkinsi, Mesuropetala muensteri, Protolindenia koehleri, Urogomphus giganteus, Morbaeschna muensteri (122586) Stenophlebia latreillei, Protolindenia wittei, Aeschnogomphus intermedius, Nannogomphus bavaricus, Cymatophlebia longialata, Cymatophlebia jurassica, Urogomphus giganteus (139876)
Early/Lower Tithonian149.2 - 145.06Germany Cymatophlebia longialata, Eumorbaeschna jurassica (105799) Nannogomphus buergeri, Cymatophlebia longialata, Urogomphus giganteus, Urogomphus eximius (122880) Urogomphus giganteus (151861 151862)
Early/Lower Tithonian149.2 - 145.06Germany (Bayern) Aeschna grandis (154305) Aeschna wittei (153703) Aeshna antiqua (154208) Cymatophlebia densa (152881) Cymatophlebia longialata (152882 152901) Cymatophlebia longialata, Eumorbaeschna jurassica, Bergeriaeschnidia inexpectata, Urogomphus giganteus (122730) Cymatophlebia longialata, Mesuropetala muensteri (105797) Cymatophlebia longialata, Mesuropetala muensteri, Urogomphus giganteus (131959) Cymatophlebia longialata, Urogomphus giganteus (105798) Cymatophlebia pumilio (117534) Eumorbaeschna jurassica (139547) Heterophlebia amphitrite, Stenophlebia phryne, Agrion latreillei, Heterophlebia aequalis, Protolindenia wittei, Aeschnogomphus buchi, Aeschnogomphus intermedius, Agrion vetustum, Agrion gracile, Libellula longialata, Eumorbaeschna jurassica, Aeschna muensteri, Mesuropetala koehleri, Aeschna gigantea, Petalura eximia, Libellula densa, Brunetaeschnidium sp., Lithoaeschnidium viohli (123985) Insecta indet. (154306) Mesuropetalidae indet. (131851) Protolindenia wittei, Cymatophlebia longialata, Bergeriaeschnidia inexpectata (139550) Protolindenia wittei, Cymatophlebia pumilio, Aeschnopsis tischlingeri (105541) Reschiostenophlebia koschnyi (227142) Reschiostenophlebia koschnyi, Prohemeroscopus kuehnapfeli (227117) Solnhofenia stoebeneri (152902) Stenophlebia amphitrite, Stenophlebia eichstaettensis, Stenophlebia latreillei, Prostenophlebia sp., Prostenophlebia jurassica, Protolindenia wittei, Protolindenia viohli, Aeschnogomphus buchi, Proterogomphus renateae, Nannogomphus buergeri, Cymatophlebia longialata, Cymatophlebia pumilio, Cymatophlebia kuempeli, Eumorbaeschna jurassica, Mesuropetala koehleri, Urogomphus giganteus (139551) Stenophlebia amphitrite, Stenophlebia latreillei, Aeschnogomphus intermedius, Nannogomphus bavaricus, Cymatophlebia longialata, Mesuropetala koehleri, Aeschnidium densum (123144) Stenophlebia latreillei, Prohemeroscopus kuehnapfeli, Prohemeroscopus jurassicus, Cymatophlebia longialata, Eumorbaeschna jurassica (152883) Stenophlebia phryne, Cymatophlebia longialata, Mesuropetala muensteri (139548) Stenophlebia sp., Protolindenia wittei, Aeschna charpentieri, Cordulegaster intermedius, Cymatophlebia longialata, Uropetala koehleri, Estemoa densa (133578) Stenophlebia sp., Stenophlebia aequalis, Cordulegaster intermedius, Agrion vetustum, Cymatophlebia longialata, Uropetala koehleri (123906) Urogomphus eximius (117531)