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Bathyurellinae

Trilobita - Proetida - Bathyuridae

Synonymy list
YearName and author
1953Bathyurellinae Hupe
1953Bathyurellinae Hupé
2013Bathyurellinae Fortey and Bruton

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RankNameAuthor
kingdomAnimalia()
—Bilateria
—EubilateriaAx 1987
—Protostomia
—Ecdysozoa
—Panarthropoda
phylumArthropodaLatreille 1829
RankNameAuthor
subphylumArtiopoda(Hou and Bergstrom)
classTrilobitaWalch 1771
orderProetidaFortey and Owens 1975
familyBathyuridaeWalcott 1886
subfamilyBathyurellinaeHupe 1953
subfamilyBathyurellinaeHupe 1953

If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.

Subfm. †Bathyurellinae Hupe 1953
show all | hide all
G. †Bathyurellus Billings 1865
+
†Bathyurellus abruptus Billings 1865
†Bathyurellus diclementsae Fortey and Bruton 2013
†Bathyurellus expansus Billings 1865
†Bathyurellus formosus Billings 1865
†Bathyurellus fraternus Billings 1865
†Bathyurellus platypus Fortey 1979
+
Invalid names: Bathyurellus pogonopensis Hintze 1951 [replaced]
†Bathyurellus teretus Young 1973
G. †Benthamaspis Poulsen 1946
+
†Benthamaspis conica Fortey 1979
+
Invalid names: Benthamaspis hintzei Boyce 1989 [synonym], Oculomagnus obreptus Lochman 1966 [synonym]
†Benthamaspis diminutiva Hintze 1953
†Benthamaspis gibberula Billings 1865
+
Invalid names: Benthamaspis distinctus Young 1973 [synonym]
†Benthamaspis mediacrista Cullison 1944
†Benthamaspis onomeris Loch 2007
†Benthamaspis problematica Poulsen 1946
†Benthamaspis sera Fortey and Droser 1999
†Benthamaspis striata Whitfield 1897
Invalid names: Oculomagnus Lochman 1966 [synonym]
G. †Ceratopeltis Poulsen 1937
+
†Ceratopeltis bachensis Adrain and Westrop 2005
†Ceratopeltis forteyi Adrain and Westrop 2005
†Ceratopeltis latilimbata Poulsen 1937
G. †Chapmania Loch 2007
+
†Chapmania carterensis Loch 2007
†Chapmania taylori Loch 2007
G. †Chapmanopyge Fortey and Bruton 2013
+
†Chapmanopyge knudseni McCobb et al. 2014
†Chapmanopyge sanddoelaensis Adrain and Westrop 2005
G. †Grinnellaspis Poulson 1946
+
†Grinnellaspis feildeni Poulsen 1946
G. †Harlandaspis Fortey and Bruton 2013
+
†Harlandaspis elongata Fortey and Bruton 2013
†Harlandaspis inflata Loch 2007
G. †Licnocephala Ross 1951
+
†Licnocephala brevicauda Poulsen 1937
†Licnocephala cavigladius Hintze 1953
†Licnocephala ovata Ross 1953
†Licnocephala sanddoelaensis Adrain and Westrop 2005
†Licnocephala sminue Fortey and Peel 1990
G. †Lutesvillia Cullison 1944
+
†Lutesvillia bispinosa Cullison 1944
G. †Punka Fortey 1979
+
†Punka adamsi McCobb et al. 2014
†Punka akoura Loch 2007
†Punka caecata Fortey 1980
†Punka flabelliformis Fortey 1979
†Punka latissima Fortey and Bruton 2013
†Punka nitidus Billings 1865
†Punka teicherti Poulsen 1937
†Punka verecunda Loch 2007
G. †Uromystrum Whittington 1953
+
†Uromystrum affine Poulsen 1937
†Uromystrum brevispinum Raymond 1905
†Uromystrum drepanon Fortey and Bruton 2013
†Uromystrum marginiatus Billings 1865
†Uromystrum minor Raymond 1905
†Uromystrum patulum Whittington 1963
†Uromystrum validus Billings 1865
Diagnosis
No diagnoses are available
Measurements
No measurements are available
Composition: low Mg calcitec
Composition 2: chitinc
Ontogeny: molting, addition of partsc
Environment: marinec
Locomotion: fast-movingc
Life habit: low-level epifaunalc
Diet: deposit feedero
Vision: well-developedf
Created: 2009-12-19 02:31:34
Modified: 2009-12-19 04:31:34
Source: f = family, o = order, c = class
References: Aberhan et al. 2004, Whittington et al. 1997, Fortey and Owens 1999

Age range: base of the Tulean to the top of the Llandeilo or 480.80000 to 458.40000 Ma

Collections (316 total)


Time interval Ma Country or state Original ID and collection number
Skullrockian - Tulean487.5 - 474.9Canada (Newfoundland) Grinnellaspis newfoundlandensis (195781)
Ibexian487.5 - 470.0USA (Nevada) Licnocephala sp. (104563)
Ibexian487.5 - 470.0USA (New York) Benthamaspis striata (64618)
Ibexian487.5 - 470.0Canada (Quebec) Peltabellia amplimarginata (111446)
Tremadoc486.5 - 477.7Canada (Alberta) Benthamaspis canadensis (10930 10934) Benthamaspis diminutiva (10903) Benthamaspis obrepta (10946)
Stairsian485.4 - 478.6Canada (Newfoundland) Parahystricurus mediacrista (196438)
Stairsian485.4 - 478.6USA (Utah) Licnocephala sp. (104110)
Stairsian485.4 - 478.6USA (Idaho) Benthamaspis sp. 1, Benthamaspis sp. 2 (228283)
Tremadocian485.4 - 477.7Greenland Licnocephala sminue (181518)
Early/Lower Ordovician485.4 - 470.0USA (Texas) Bathyurellus sp. (293)
Jeffersonian485.4 - 470.0USA (Missouri) Jeffersonia mediacrista (56505)
Jeffersonian485.4 - 470.0USA (Oklahoma) Bathyurellus arbucklensis (106615) Bathyurellus arbucklensis, Benthamaspis rhochmotis, Lutesvillia bispinosa (106876) Bathyurellus arbucklensis, Benthamaspis rhochmotis, Lutesvillia bispinosa, Punka akoura, Punka verecunda (106624) Bathyurellus arbucklensis, Benthamaspis rhochmotis, Punka akoura, Punka verecunda (106614) Bathyurellus arbucklensis, Punka verecunda (106928) Bathyurellus inflatus (106733) Bathyurellus inflatus, Benthamaspis rhochmotis (106734 106935) Bathyurellus inflatus, Benthamaspis rhochmotis, Lutesvillia bispinosa (106737) Bathyurellus sp. 1, Punka verecunda (106740) Benthamaspis mediacrista (106577 106860) Benthamaspis rhochmotis (106620 106625 106925 106926) Benthamaspis rhochmotis, Lutesvillia bispinosa, Punka akoura (106612) Benthamaspis rhochmotis, Punka akoura (106610 106611 106617) Benthamaspis rhochmotis, Punka akoura, Punka verecunda (106613) Benthamaspis rhochmotis, Punka verecunda (106927) Chapmania carterensis (106871) Chapmania carterensis, Lutesvillia bispinosa (106579 106580 106601 106602) Chapmania oklahomensis (106582 106587 106589 106591 106595 106866) Chapmania oklahomensis, Chapmania carterensis (106592 106597) Chapmania oklahomensis, Chapmania carterensis, Lutesvillia bispinosa (106588) Chapmania oklahomensis, Lutesvillia bispinosa (106590 106598 106870) Chapmania oklahomensis, Lutesvillia bispinosa, Punka akoura (106593) Chapmania oklahomensis, Punka akoura (106596) Chapmania sp. 1, Chapmania oklahomensis (106863) Chapmania taylori (106572 106573 106838 106839 106840 106841 106842 106843 106844 106849 106854 106855 106857 106858) Lutesvillia bispinosa (106578 106585 106586 106594 106603 106604 106607 106608 106627 106630 106631 106632 106633 106736 106865 106868 106934) Lutesvillia bispinosa, Punka akoura (106599 106731) Lutesvillia bispinosa, Punka akoura, Punka verecunda (106932 106933) Punka akoura (106600 106605 106616 106634 106872 106875 106922) Punka sp. 1, Punka akoura (106874) Punka verecunda (106628 106629 106742 106930)
Jeffersonian485.4 - 470.0Canada (Newfoundland) Benthamaspis hintzei, Uromystrum affine (195785) Benthamaspis sp., Bathyurellus abruptus (195774) Grinnellaspis sp., Benthamaspis conica, Benthamaspis hintzei, Grinnellaspis newfoundlandensis, Uromystrum affine (195756)
Jeffersonian485.4 - 470.0Canada (Newfoundland and Labrador) Benthamaspis hintzei (106229) Benthamaspis hintzei, Grinnellaspis newfoundlandensis (106236) Benthamaspis hintzei, Grinnellaspis newfoundlandensis, Uromystrum affine (133147) Grinnellaspis newfoundlandensis (106224 106230 106234 106235)
Cassinian482.3 - 470.0USA (Oklahoma) Bathyurellus inflatus (106750 106937) Bathyurellus inflatus, Punka verecunda (106954) Bathyurellus sp. 1, Punka verecunda (106941) Benthamaspis onomeris (106789 106791 106799 106800 106803 106825 106826) Benthamaspis onomeris, Punka verecunda (106790 106794) Lutesvillia sp. 1, Punka verecunda (106955) Punka akoura (106745) Punka verecunda (106751 106753 106755 106766 106768 106769 106770 106777 106778 106787 106792 106805 106943 106944 106945 106949 106957)
Cassinian482.3 - 470.0United Kingdom (Scotland) Punka aetholiciocorus (75589)
Cassinian482.3 - 470.0Canada (Newfoundland) Bathyurellus abruptus, Grinnellaspis flabelliformis (195784) Bathyurellus platypus, Benthamaspis gibberula, Bathyurellus abruptus (195782) Bathyurellus sp., Uromystrum sp., Bathyurellus platypus, Benthamaspis gibberula, Bathyurellus abruptus, Grinnellaspis marginiatus, Grinnellaspis flabelliformis (195758) Benthamaspis conica, Bathyurellus abruptus, Grinnellaspis flabelliformis (195783) Uromystrum sp., Bathyurellus platypus, Benthamaspis conica, Bathyurellus abruptus, Grinnellaspis flabelliformis (195757)
Cassinian482.3 - 470.0Canada (Newfoundland and Labrador) Bathyurellus abruptus (133128) Bathyurellus abruptus, Uromystrum marginatus (133127) Bathyurellus platypus, Benthamaspis gibberula, Bathyurellus abruptus, Uromystrum marginatus, Grinnellaspis flabelliformis (133140) Benthamaspis conica, Uromystrum affine, Bathyurellus abruptus (106239 106241) Benthamaspis hintzei, Grinnellaspis newfoundlandensis, Uromystrum affine (106238) Benthamaspis sp. (69196) Punka sp., Bathyurellus platypus, Punka flabelliformis, Benthamaspis conica, Uromystrum affine, Benthamaspis gibberula, Bathyurellus abruptus (103540)
Cassinian482.3 - 470.0Greenland Bathyurellus teicherti, Niobe brevicauda, Ceratopeltis latilimbata (122083)
Cassinian482.3 - 470.0USA (Missouri) Lutesvillia bispinosa (210295)
Tulean480.8 - 474.9USA (Utah) Benthamaspis obrepta (104101 104102) Benthamaspis sp. 3 (228284) Benthamaspis sp. 6 (228306) Benthamaspis sp. 7, Benthamaspis sp. 8, Benthamaspis sp. 9 (228309) Licnocephala bicornuta (104083) Licnocephala cavigladius (104099 104103 104104 104105 104115)
Tulean480.8 - 474.9Canada (Newfoundland) Benthamaspis conica (195769)
Tulean480.8 - 474.9Greenland Chapmanopyge knudseni (223777 223792 223793 223794) Chapmanopyge knudseni, Punka adamsi (223795) Punka adamsi (223796)
Tulean480.8 - 474.9Canada (Nunavut) Licnocephala n. sp. A, Licnocephala sanddoelaensis (173915) Licnocephala sanddoelaensis (173917) Licnocephala sanddoelaensis, Ceratopeltis bachensis (173916)
Tulean480.8 - 474.9USA (Idaho) Benthamaspis sp. 4, Licnocephala sp. 3, Licnocephala bicornuta (228285) Benthamaspis sp. B, Licnocephala sp. 5 (228300) Benthamaspis sp., Licnocephala sp. (228268) Licnocephala bicornuta (103803) Licnocephala cavigladius, Licnocephala ovata (105094) Licnocephala ovata (105091) Licnocephala sp. (228266) Licnocephala sp. 1 (228282) Licnocephala sp. 3, Licnocephala bicornuta (228298) Licnocephala sp. A (228299) Licnocephala sp., Licnocephala bicornuta (103805 103820) Trilobita undet. gen. and sp. C (103806)
Tulean480.8 - 474.9Norway (Spitsbergen) Chapmanopyge amplimarginiata (189834) Licnocephala sp. (189832) Licnocephala sp., Uromystrum affine, Bathyurellus diclementsae, Chapmanopyge sp., Chapmanopyge sanddoelaensis, Punka latissima, Ceratopeltis batchensis (189836) Licnocephala sp., Uromystrum affine, Bathyurellus diclementsae, Chapmanopyge sp., Harlandaspis elongata, Licnocephala brevicauda, Punka latissima, Uromystrum drepanon (189837)
Tulean480.8 - 474.9USA (Montana) Lichnocephala sp. (287) Licnocephala sp. A (174361)
Tulean480.8 - 474.9Canada (Newfoundland and Labrador) Benthamaspis gibberula (195775)
Tulean - Blackhillsian480.8 - 470.0Canada (Newfoundland) Benthamaspis conica, Bathyurellus abruptus (195779) Benthamaspis sp., Grinnellaspis sp., Uromystrum sp., Bathyurellus platypus, Punka flabelliformis, Benthamaspis conica, Uromystrum affine, Benthamaspis gibberula, Bathyurellus abruptus (196380) Uromystrum sp., Benthamaspis conica, Uromystrum affine, Bathyurellus abruptus (195770) Uromystrum sp., Uromystrum affine (196385)
Tulean - Blackhillsian480.8 - 470.0Canada (Newfoundland and Labrador) Benthamaspis gibberula (196504) Benthamaspis sp., Grinnellaspis sp., Benthamaspis conica (196503) Benthamaspis sp., Punka sp., Uromystrum affine, Bathyurellus abruptus (196383) Punka flabelliformis, Benthamaspis conica, Uromystrum affine, Benthamaspis gibberula, Bathyurellus abruptus (196381) Punka flabelliformis, Benthamaspis gibberula, Bathyurellus abruptus (195772) Uromystrum forteyi, Punka sp., Bathyurellus platypus, Punka flabelliformis, Benthamaspis conica, Benthamaspis gibberula, Bathyurellus abruptus (195776)
Arenigian478.6 - 468.1USA (Utah) Bathyurellus sp. (64496 64502 64508 64516)
Arenigian478.6 - 468.1USA (Nevada) Bathyurellus sp. (64522)
Arenig478.6 - 466.0USA (North Dakota) Oculomagnus sp., Lichnocephala sp. (429)
Arenig478.6 - 466.0Greenland Bathyurellus abruptus (221131) Bathyurellus sp., Bathyurellus teicherti (122088) Bathyurellus teicherti (122080 122082 122116) Niobe brevicauda (122081) Punka flabelliformis, Benthamaspis conica, Bathyurellus abruptus (221130)
Arenig478.6 - 466.0USA (Vermont) Grinnellaspis marginiata (55241)
Arenig478.6 - 466.0Argentina Uromystrum sp. (8721)
Arenig478.6 - 466.0USA (New York) Bathyurellus platypus, Grinnellaspis marginiata (55243)
Arenig478.6 - 466.0Canada (Newfoundland and Labrador) Bathyurellus sp., Benthamaspis sp. (78095) Benthamaspis sp. (78092 78093)
Floian477.7 - 470.0Canada (Newfoundland) Bathyurellus sp., Benthamaspis sp., Punka sp., Punka flabelliformis, Benthamaspis conica, Uromystrum affine, Benthamaspis gibberula, Bathyurellus abruptus, Uromystrum marginatus (195748) Benthamaspis sp., Bathyurellus abruptus (195746)
Floian477.7 - 470.0Russian Federation (Siberia) Bathyurellus sp. (205383)
Floian - Darriwilian477.7 - 458.4Russian Federation (Siberia) Bathyurellus nonnulus (205664)
Blackhillsian474.9 - 470.0Canada (Newfoundland) Benthamaspis sp. (195777)
Blackhillsian474.9 - 470.0USA (Utah) Bathyurellus sp. (104130) Bathyurellus teretus, Benthamaspis distinctus (69261) Benthamaspis diminutiva (63846 63848)
Blackhillsian474.9 - 470.0Canada (Alberta) Benthamaspis diminutiva (10899)
Blackhillsian474.9 - 470.0Norway (Spitsbergen) Benthamaspis conica, Benthamaspis gibberula (189839) Punka flabelliformis, Benthamaspis conica, Grinnellaspis newfoundlandensis, Uromystrum affine, Benthamaspis gibberula, Bathyurellus abruptus, Licnocephala brevicauda, Uromystrum drepanon (189838)
Blackhillsian474.9 - 470.0USA (Nevada) Licnocephala cavigladius (104568) Licnocephala sp. (104354)
Rangerian470.0 - 470.3USA (Nevada) Bathyurellus sp. (104430) Bathyurellus sp. 3 (80059) Benthamaspis sera (104653) Benthamaspis sp. (104655) Uromystrum validum (37485)
Rangerian470.0 - 470.3USA (Utah) Bathyurellus sp., Bathyurellus nov. sp. (104061) Punka nitida (37454 37458 37473 37474) Uromystrum sp. (37452) Uromystrum validum, Punka nitida (37462)
Castlemainian - Yapeenian470.0 - 467.3Ireland Benthamaspis diminutiva (153922)
Dapingian470.0 - 467.3USA (Utah) Bathyurellus pogonopensis (104199 104214 104215 104216 104217 104218 104220 104221 104222 104223 104226 104227)
Chazyan470.0 - 457.3USA (New York) Bathyurellus minor (172321 172348)
Chazyan470.0 - 457.3Canada (Newfoundland and Labrador) Uromystrum validum (104870)
Whiterockian470.0 - 457.3USA (Nevada) Bathyurellus sp. (80081) Bathyurellus sp. 3 (80061) Bathyurellus sp. 4 (80054) Uromystrum sp. (104376 104378) Uromystrum validum (37486 37488 37490)
Whiterockian470.0 - 457.3USA (Oklahoma) Bathyurellus sp. (105616 105625 105641 105645)
Whiterockian470.0 - 457.3Norway (Spitsbergen) Punka caecata (106480)
Whiterockian470.0 - 457.3Canada (Newfoundland and Labrador) Uromystrum fraternum, Uromystrum formosum, Uromystrum patulum, Bathyurellus nitidus (104752) Uromystrum validum (104863 104875 104878 104881 104882 104883 104884 104885 104886 104887)
Whiterockian470.0 - 457.3Canada (Quebec) Uromystrum minor (111333 111361 111363 111431 111432)
Whiterockian470.0 - 457.3USA (New York) Uromystrum brevispinum (103280) Uromystrum minor (102995 103004 103311)
Whiterockian470.0 - 457.3USA (Utah) Bathyurellus pogonipensis (298) Bathyurellus pogonopensis (104244) Uromystrum sp. (104064)
Llanvirn468.0 - 458.4Canada (Newfoundland and Labrador) Benthamaspis sp. (78097) Uromystrum sp., Bathyurellus nitidus (78099)
Darriwilian467.3 - 458.4Russian Federation (Gorny Altai) Bathyurellus nonnulus (186015 194972)
Darriwilian467.3 - 458.4Russian Federation (Siberia) Bathyurellus nonnulus (206165 206166)
Llandeilo461.1 - 458.4Greenland Bathyurellus sp. (122117)