PBDB Taxon

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Sciophilinae

Insecta - Diptera - Mycetophilidae

Taxonomy

Sciophilinae was named by Winnertz (1863). It is extant.

It was reranked as the family Sciophilidae by Hong (1992).

It was assigned to Diptera by Hong (1992); and to Mycetophilidae by Johannsen (1909), Enderlein (1911), Theobald (1937), Edwards (1940), Matile (1989), Blagoderov (2007), Borkent and Wheeler (2013), Blagoderov (2019) and Oliveira and Amorim (2021).

Synonyms

Fushunosciophila was named by Hong (2002).

It was considered a nomen nudum by Hong (2002) and Hong (2002).
Fushunosciophila furvis was named by Hong (2002). It is an inclusion in amber. Its type locality is Xilutian coal mine, Fushun amber (post-1980 collection), which is in a Ypresian terrestrial amber in China.

It was considered a nomen nudum by Hong (2002) and Hong (2002).

Synonymy list

Year	Name and author
1863	Sciophilinae Winnertz p. 658
1909	Sciophilinae Johannsen p. 30
1911	Sciophilinae Enderlein p. 154
1937	Sciophilinae Theobald p. 323
1940	Sciophilinae Edwards p. 120
1989	Sciophilinae Matile p. 136
1992	Sciophilidae Hong p. 4
2002	Fushunosciophila Hong p. 326
2002	Fushunosciophila furvis Hong p. 327 figs. 2.7.541-544
2007	Sciophilinae Blagoderov
2013	Sciophilinae Borkent and Wheeler p. 417
2019	Sciophilinae Blagoderov p. 535
2021	Sciophilinae Oliveira and Amorim

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Rank	Name	Author
kingdom	Animalia	()
—	Bilateria
—	Eubilateria	Ax 1987
—	Protostomia
—	Ecdysozoa
—	Panarthropoda
phylum	Arthropoda	Latreille 1829
superclass	Hexapoda	Latreille 1825

Rank	Name	Author
class	Insecta
order	Diptera
—	Neodiptera	Michelsen 1994
infraorder	Bibionomorpha
superfamily	Sciaroidea
family	Mycetophilidae
subfamily	Sciophilinae
subfamily	Sciophilinae

If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.

Subfm. Sciophilinae Winnertz 1863

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G. Acnemia Winnertz 1863

†Acnemia bolsiusi Meunier 1904

†Acnemia cyclosoma Cockerell 1925

†Acnemia simplex Cockerell 1921

G. Allocotocera Mik 1886

†Allocotocera burmitica Blagoderov and Grimaldi 2004

†Allocotocera xavieri Blagoderov and Arillo 2002

G. Anaclileia Meunier 1904

†Anaclileia anacliniformis Meunier 1904

†Anaclileia dissimilis Meunier 1904

†Anaclileia gazagnairei Meunier 1904

†Anaclileia sylvatica Meunier 1904

G. Aneura Marshall 1896

†Aneura apicalis Riek 1954

G. Azana Walker 1856

†Azana akarenos Kerr and Greenwalt 2022

†Azana cockerelli Blagoderov 2019

†Azana rarissima Meunier 1904

G. †Baisodicrana Blagoderov 1995

†Baisodicrana incompleta Blagoderov 1995

†Baisodicrana mongolica Blagoderov 1998

†Baisodicrana secunda Blagoderov 1995

G. †Ekhiritus Blagoderov 1995

†Ekhiritus disannuus Blagoderov 1995

G. Leptomorphus Curtis 1831

†Leptomorphus africanus Meunier 1907

†Leptomorphus palaeospilus Cockerell 1921

†Leptomorphus sepultus Meunier 1917

Invalid names: Diomonus Walker 1848 [synonym]

G. †Necromyza Scudder 1895

†Necromyza pedata Scudder 1895

G. Neoaphelomera Miller 1945

G. Neuratelia Rondani 1856

†Neuratelia gibbosa Meunier 1904

†Neuratelia giebeli Meunier 1904

†Neuratelia maimecha Blagoderov and Grimaldi 2004

†Neurotelia landrocki Statz 1944

Invalid names: Anaclinia Winnertz 1863 [synonym], Odontopoda Aldrich 1897 [synonym], Proanaclinia Meunier 1904 [synonym]

G. Phthinia Winnertz 1863

†Phthinia longipoda Statz 1944

G. †Pollicitator Blagoderov 1995

†Pollicitator baisae Blagoderov 1995

†Pollicitator bontsaganicus Blagoderov 1998

†Pollicitator pollicitator Blagoderov 2000

G. Polylepta Winnertz 1863

Subg. †Polylepta (Lyptolape) Blagoderov 1998

†Polylepta (Lyptolape) lyptolape Blagoderov 1998

†Polylepta (Lyptolape) olenguiensis Blagoderov 2000

†Polylepta filipes Meunier 1904

G. †Prospeolepta Blagoderov 1995

†Prospeolepta brevicubita Blagoderov 2000

†Prospeolepta parallelimedia Blagoderov 1998

†Prospeolepta simplex Blagoderov 1995

†Prospeolepta trapezia Blagoderov 1995

G. †Protasmanina Riek 1954

†Protasmanina nana Riek 1954

G. †Pseudomanota Blagoderov and Grimaldi 2004

†Pseudomanota perplexa Blagoderov and Grimaldi 2004

G. Sciophila Meigen 1818

†Sciophila armipes Meunier 1899

†Sciophila atra Giebel 1856

†Sciophila carbonaria Meunier 1899

†Sciophila curvipetiolata Meunier 1904

†Sciophila dilatata Loew 1850

†Sciophila hyatti Scudder 1890

†Sciophila inermis Meunier 1899

†Sciophila loewi Giebel 1856

†Sciophila micropora Meunier 1899

†Sciophila minutissima Meunier 1915

†Sciophila mirandula Cockerell 1909

†Sciophila socialis Giebel 1856

†Sciophila takoyensis Blagoderov 2007

Invalid names: Lasiosoma Winnertz 1863 [synonym], Sciophila atropos Meunier 1899 [nomen nudum], Sciophila blotho Meunier 1899 [nomen nudum], Sciophila cognata Meunier 1899 [nomen nudum], Sciophila disjuncta Meunier 1899 [nomen nudum], Sciophila lachesis Meunier 1899 [nomen nudum], Sciophila obscura Meunier 1899 [nomen nudum], Sciophila spinipes Meunier 1899 [nomen nudum], Sciophila trapezoides Meunier 1899 [nomen nudum]

Tr. Sciophilini Winnertz 1863

G. Neuroteliadup Rondani 1856

G. Syntemna Winnertz 1863

†Syntemna compressa Meunier 1904

†Syntemna dama Meunier 1917

†Syntemna elongata Meunier 1904

†Syntemna falcata Blagoderov 2007

†Syntemna fissurata Blagoderov and Grimaldi 2004

†Syntemna johannseni Meunier 1922

†Syntemna lundstromi Meunier 1922

†Syntemna mesozoica Blagoderov 1995

†Syntemna minuta Meunier 1917

†Syntemna oblita Meunier 1917

†Syntemna pinites Meunier 1904

†Syntemna prolongata Meunier 1904

†Syntemna sciophiliformis Meunier 1904

†Syntemna subcylindrica Meunier 1904

†Syntemna subquadrata Meunier 1904

†Syntemna tele Blagoderov 1998

†Syntemna zherikhini Blagoderov 2000

†Syntemna zhuzhan Blagoderov 1998

G. †Zazicia Blagoderov 1995

†Zazicia innuba Blagoderov 1995

Invalid names: Fushunosciophila Hong 2002 [nomen nudum]

Diagnosis

No diagnoses are available

Measurements

No measurements are available

Composition:

chitinc

Environment:

terrestrialc

Locomotion:

actively mobilep

Created:

2017-04-17 09:56:18

Modified:

2017-04-17 09:56:18

Source: c = class, p = phylum

References: Bush and Bambach 2015, Kiessling 2004

Age range

Maximum range based only on fossils: base of the Hauterivian to the top of the Holocene or 132.90000 to 0.00000 Ma
Minimum age of oldest fossil (stem group age): 125.0 Ma

Collections (44 total)

Time interval	Ma	Country or state	Original ID and collection number
Hauterivian - Barremian	132.9 - 125.0	Mongolia (Bayankhongor)	Syntemna tele (113357)
Aptian	125.0 - 113.0	Russian Federation (Buryatia)	Baisodicrana incompleta (125797) Baisodicrana secunda, Prospeolepta trapezia, Pollicitator baisae (125791) Ekhiritus disannuus, Prospeolepta trapezia (125413) Zazicia innuba (106103 122552) Zazicia innuba, Baisodicrana incompleta, Prospeolepta trapezia (119498) Zazicia innuba, Prospeolepta trapezia, Pollicitator baisae (119441) Zazicia innuba, Syntemna mesozoica, Ekhiritus disannuus, Prospeolepta trapezia, Prospeolepta simplex, Pollicitator baisae (122549)
Aptian	125.0 - 113.0	Mongolia	Prospeolepta brevicubita (103273) Prospeolepta parallelimedia (121711) Syntemna sp., Syntemna tele (129998) Syntemna sp., Syntemna tele, Syntemna zhuzhan (121710) Syntemna sp., Syntemna tele, Syntemna zhuzhan, Polylepta sp., Baisodicrana mongolica, Polylepta lyptolape (113359) Syntemna sp., Syntemna tele, Syntemna zhuzhan, Polylepta sp., Baisodicrana mongolica, Polylepta lyptolape, Pollicitator bontsaganicus, Zazicia innuba (113356) Syntemna tele (129999) Syntemna tele, Polylepta sp., Polylepta lyptolape (130000) Syntemna tele, Syntemna zhuzhan (113355 125194) Syntemna tele, Syntemna zhuzhan, Zazicia innuba (121714)
Early/Lower Aptian	121.4 - 122.46	Russian Federation	Polylepta olenguiensis (123615)
Late/Upper Albian	105.3 - 99.6	Spain (Álava)	Allocotocera xavieri (108769)
Early/Lower Cenomanian	99.6 - 93.5	Myanmar	Allocotocera burmitica, Pseudomanota perplexa (111688)
Late/Upper Santonian	85.8 - 83.5	Russian Federation	Neuratelia maimecha (35056)
Early/Lower Campanian	83.5 - 70.6	Russian Federation (Tenkinskii)	Pollicitator pollicitator, Syntemna zherikhini, Syntemna tele, Syntemna zhuzhan (105375)
Middle Campanian	83.5 - 70.6	Canada (Alberta)	Syntemna fissurata (34896)
Ypresian	56.0 - 47.8	China (Liaoning)	Fushunosciophila furvis (124820)
Ypresian	56.0 - 47.8	Australia (Queensland)	Protasmanina nana, Aneura apicalis (125666)
Bridgerian	50.3 - 46.2	USA (Colorado)	Acnemia cyclosoma (122746) Diomonus palaeospilus (121193)
Bridgerian	50.3 - 46.2	USA (Wyoming)	Anaclinia sp., Sciophila hyattii (122092)
Lutetian	47.8 - 41.3	USA (Montana)	Azana akarenos (117614)
Lutetian	47.8 - 41.3	Russian Federation	Sciophila takoyensis, Syntemna falcata (107340)
Priabonian	38.0 - 33.9	Poland	Sciophila dilatata (109625)
Priabonian	38.0 - 33.9	United Kingdom (England)	Acnemia simplex, Azana cockerelli (123959)
Priabonian	38.0 - 33.9	Russian Federation (Kaliningrad)	Polylepta filipes, Lasiosoma curvipetiolata, Syntemna elongata, Syntemna pinites, Syntemna compressa, Syntemna subcylindrica, Syntemna subquadrata, Syntemna sciophiliformis, Proanaclinia giebeli, Proanaclinia gibbosa, Anaclileia anacliniformis, Anaclileia sylvatica, Anaclileia gazagnairei, Anaclileia dissimilis, Azana rarissima, Acnemia bolsiusi, Syntemna dama, Syntemna oblita, Syntemna minuta, Diomonus sepultus, Sciophila armipes, Sciophila carbonaria, Sciophila inermis, Sciophila micropora, Syntemna johannseni, Syntemna lundstromi, Syntemna prolongata (123927) Sciophila loewi (147348) Sciophila socialis, Sciophila atra (147344)
Chadronian	37.2 - 33.9	USA (Colorado)	Lasiosoma mirandula (106381)
MP 30	28.4 - 23.03	Germany (Nordrhein-Westfalen)	Lasiosoma minutissima (138696)
Chattian	28.1 - 23.03	Germany (North Rhine-Westphalia)	Neurotelia landrocki, Phthinia longipoda (122010)
Burdigalian	20.44 - 15.97	New Zealand (Otago)	Neoaphelomera sp. (191650)
Sarmatian	12.7 - 11.608	Germany (Baden-Württemberg)	Necromyza pedata (113830)
Holocene	0.0117 - 0.0	Madagascar	Leptomorphus africanus (152187)