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Panorpida

Insecta

Taxonomy
Panorpida was named as a superorder. It is extant.

It was reranked as the unranked clade Panorpida by Engel (2022).

It was assigned to Holometabola by Beckemeyer and Hall (2007); and to Aparaglossata by Engel (2022).

Synonymy list
YearName and author
2007Panorpida Beckemeyer and Hall p. 24
2022Panorpida Engel

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RankNameAuthor
kingdomAnimalia()
—Bilateria
—EubilateriaAx 1987
—Protostomia
—Ecdysozoa
—Panarthropoda
phylumArthropodaLatreille 1829
superclassHexapodaLatreille 1825
classInsecta
RankNameAuthor
—Dicondylia
—Paranotalia
subclassPterygota()
—NeopterygotaCrampton 1924
infraclassNeopteraMartynov 1923
—Holometabola
—Aparaglossata
—Panorpida
—Panorpida

If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.

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Superor. Amphiesmenoptera Kiriakoff 1948
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G. †Aphryganoneura Tillyard 1926
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†Aphryganoneura anomala Tillyard 1926
G. †Liadoptilia Handlirsch 1939
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†Liadoptilia misera Handlirsch 1939
Unr. Metamphiesmenoptera Engel 2022
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Unr. Euamphiesmenoptera Engel 2022
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Or. †Eocoronoptera Engel 2022
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Fm. †Eocoronidae Tindale 1980
Unr. Panendymenoptera Engel 2022
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Unr. Endymenoptera Engel 2022
Or. †Necrotrichoptera Engel 2022
Invalid names: Cladochoroptera Engel 2022 [empty]
G. †Microcarbonella Prokop et al. 2022
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†Microcarbonella paradoxa Prokop et al. 2022
Or. †Permotrichoptera Martynova 1958
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Fm. †Microptysmatidae Martynova 1958
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G. †Kamopanorpa Martynov 1928
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†Kamopanorpa aristovi Sukatsheva and Sinitshenkova 2023
†Kamopanorpa grossa Novokshonov 1993
†Kamopanorpa karaungirica Novokshonov 1993
†Kamopanorpa lata Martynov 1928
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Invalid names: Microptysmodes uralicus Martynova 1958 [synonym]
†Kamopanorpa latipennata Novokshonov 1993
†Kamopanorpa maculosa Bashkuev and Sukatsheva 2021
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Invalid names: Kamopanorpa maculata Sukatsheva and Aristov 2013 [replaced]
†Kamopanorpa mongolica Sukacheva 1992
†Kamopanorpa pritykinae Sukacheva 1976
†Kamopanorpa rasnitsyni Bashkuev and Sukatsheva 2021
†Kamopanorpa rotunda Bashkuev and Sukatsheva 2021
†Kamopanorpa rotundipennis Martynov 1932
†Kamopanorpa sibirica Martynova 1958
†Kamopanorpa sivchikovi Bashkuev and Sukatsheva 2021
†Kamopanorpa sukatchevae Novokshonov 1993
†Kamopanorpa tamarae Sukatsheva 2017
†Kamopanorpa tshepanikhensis Sukatsheva and Aristov 2013
†Kamopanorpa tyulkiana Sukatsheva 2017
†Kamopanorpa uralensis Martynov 1940
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Invalid names: Kamopanorpa fasciipennis Martynova 1942 [synonym], Kamopanorpa incerta Sukacheva 1976 [synonym], Kamopanorpa pallida Sukacheva 1976 [synonym], Uraloptysma maculata Ivanov 1992 [synonym]
Invalid names: Microptysma Martynova 1958 [synonym], Microptysmodes Martynova 1958 [synonym], Uraloptysma Ivanov 1992 [synonym]
Invalid names: Uraloptysmatidae Ivanov 1992 [synonym]
Fm. †Microptysmellidae Engel 2022
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G. †Microptysmella Kukalová-Peck and Willmann 1990
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†Microptysmella moravica Kukalová-Peck and Willmann 1990
Or. †Protomeroptera Engel 2022
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Fm. †Protomeropidae Tillyard 1926
+
G. †Glanomerope Poschmann and Nel 2021
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†Glanomerope virgoferroa Poschmann and Nel 2021
G. †Permomerope Tillyard 1926
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†Permomerope australis Tillyard 1926
†Permomerope karaungirica Sukacheva 1976
†Permomerope nanus Riek 1953
†Permomerope natalensis Sukatsheva and Mostovski 2007
†Permomerope ramosa Sukacheva 1976
G. †Platychorista Tillyard 1926
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†Platychorista venosa Tillyard 1926
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Invalid names: Protomerope permiana Tillyard 1926 [synonym]
Invalid names: Protomerope Tillyard 1926 [synonym]
G. †Pseudomerope Kukalová-Peck and Willmann 1990
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†Pseudomerope gallei Kukalová-Peck and Willmann 1990
†Pseudomerope havlati Kukalová-Peck and Willmann 1990
†Pseudomerope mareki Kukalová-Peck and Willmann 1990
†Pseudomerope oborana Kukalová-Peck and Willmann 1990
G. †Pseudomeropella Kukalová-Peck and Willmann 1990
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†Pseudomeropella nekvasilovae Kukalová-Peck and Willmann 1990
G. †Stenomerope Kukalová-Peck and Willmann 1990
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†Stenomerope spinari Kukalová-Peck and Willmann 1990
G. †Westphalomerope Nel et al. 2007
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†Westphalomerope maryvonneae Nel et al. 2007
Invalid names: Marimerobiidae Willmann 1978 [synonym], Permomeropidae Riek 1953 [synonym], Platychoristidae Carpenter 1930 [synonym]
Fm. †Stereochoristidae Tillyard 1919
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G. †Stereochorista Tillyard 1919
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†Stereochorista frustrata Tillyard 1919
G. †Trichopteridium Geinitz 1880
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†Trichopteridium gracile Geinitz 1880
Invalid names: Necrotaulius handlirschi Fischer 1962 [nomen dubium], Necrotaulius regularis Handlirsch 1939 [nomen dubium], Necrotaulius vicinus Handlirsch 1939 [nomen dubium], Papilionidea Laicharting 1781 [synonym]
Superor. Antliophora Hennig 1969
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Or. Mecoptera Packard 1886 [scorpionfly]
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Fm. †Anormochoristidae Tillyard 1926
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G. †Anormochorista Tillyard 1926
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†Anormochorista oligoclada Tillyard 1926
Fm. †Belmontiidae Tillyard 1919
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G. †Belmontia Tillyard 1919
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†Belmontia mitchelli Tillyard 1919
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Invalid names: Parabelmontia permiana Tillyard 1922 [synonym]
Invalid names: Parabelmontia Tillyard 1922 [synonym]
Invalid names: Parabelmontiidae Tillyard 1922 [synonym]
Fm. Boreidae Latreille 1816
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Subfm. Boreinae Latreille 1816
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G. †Palaeoboreus Sukacheva and Rasnitsyn 1992
Fm. †Choristopsychidae Martynov 1937
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G. †Choristopsyche Martynov 1937
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†Choristopsyche asticta Qiao et al. 2013
†Choristopsyche perfecta Qiao et al. 2013
†Choristopsyche tenuinervis Martynov 1937
G. †Paristopsyche Qiao et al. 2013
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†Paristopsyche angelineae Qiao et al. 2013
Fm. †Englathaumatidae Novokshonov et al. 2016
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G. †Englathauma Novokshonov et al. 2016
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†Englathauma crabbi Novokshonov et al. 2016
†Englathauma mellishae Novokshonov et al. 2016
Subor. †Eumecoptera Tillyard 1919
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Fm. †Mesorthophlebiidae Hong 1983
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G. †Mesorthophlebia Hong 1983
Fm. †Parachoristidae Handlirsch 1937
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G. †Asiathauma Novokshonov 1996
G. †Erdosia Hong et al. 2002
G. †Jibeiorthophlebia Hong 1983
G. †Junggarochorista Hong 2009
Subfm. †Neoparachoristinae Willmann 1978
Subfm. †Parachoristinae Handlirsch 1937
G. †Potrerillopanorpa Lara and Bashkuev 2020
G. †Triasoparachorista Hong 2009
Invalid names: Choristopanorpidae Willmann 1987 [synonym], Triassochoristidae Willmann 1989 [synonym]
G. †Fortiholcorpa Wang et al. 2013
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†Fortiholcorpa paradoxa Wang et al. 2013
Fm. †Holcorpidae Willmann 1989
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G. †Conicholcorpa Li et al. 2017
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†Conicholcorpa stigmosa Li et al. 2017
Invalid names: Conicholcorpa longa Zhang et al. 2021 [nomen nudum]
G. †Holcorpa Scudder 1878
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†Holcorpa dillhoffi Archibald 2010
†Holcorpa maculosa Scudder 1878
Fm. †Kaltanidae Martynova 1958
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G. †Altajopanorpa Martynova 1948
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†Altajopanorpa brevis Martynova 1961
†Altajopanorpa cellulosa Martynova 1961
†Altajopanorpa elegans Martynova 1961
†Altajopanorpa grandis Martynova 1961
†Altajopanorpa iljinskiensis Martynova 1961
†Altajopanorpa kaltanica Martynova 1948
†Altajopanorpa karaungirica Novokshonov 1994
†Altajopanorpa nigronervosa Martynova 1961
†Altajopanorpa pilosa Martynova 1958
†Altajopanorpa postrema Novokshonov 1994
†Altajopanorpa rhombica Martynova 1961
Invalid names: Kaltana Martynova 1958 [synonym]
G. †Cyclopterina Carpenter 1986
+
†Cyclopterina autumnalis Martynova 1958
Invalid names: Cycloptera Martynova 1958 [replaced]
G. †Cyclorista Martynova 1958
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†Cyclorista convexicosta Martynova 1958
†Cyclorista simplex Novokshonov 1994
G. †Kaltanochorista Martynova 1958
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†Kaltanochorista grjasevi Martynova 1958
G. †Kamochorista Bashkuev 2008
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†Kamochorista novokshonovi Bashkuev 2008
G. †Megachorista Martynova 1958
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†Megachorista basilica Martynova 1961
†Megachorista gigantea Martynova 1961
†Megachorista khalfini Martynova 1958
G. †Moravochorista Kukalová-Peck and Willmann 1990
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†Moravochorista carolina Kukalová-Peck and Willmann 1990
G. †Pinnachorista Martynova 1958
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†Pinnachorista cubitalis Martynova 1961
†Pinnachorista fausta Martynova 1961
†Pinnachorista imitabilis Martynova 1961
†Pinnachorista indistincta Novokshonov 1994
†Pinnachorista latipennis Martynova 1961
†Pinnachorista migdisovi Martynova 1961
†Pinnachorista ovata Novokshonov 1994
†Pinnachorista problematica Novokshonov 1994
†Pinnachorista rotundipennis Martynova 1961
†Pinnachorista sarbalensis Martynova 1958
†Pinnachorista suchanovi Martynova 1961
G. †Pseudochorista Novokshonov 1994
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†Pseudochorista excusabunda Novokshonov 1997
†Pseudochorista kovbasinae Novokshonov 1994
†Pseudochorista maculata Novokshonov 1994
†Pseudochorista mongolica Novokshonov 1994
†Pseudochorista occidentalis Bashkuev 2008
G. †Tomiochorista Martynova 1958
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†Tomiochorista minuta Martynova 1961
†Tomiochorista nubila Martynova 1958
†Tomiochorista prodigialis Martynova 1961
†Tomiochorista rhombica Martynova 1961
†Tomiochorista skoki Martynova 1961
Invalid names: Cyclopteridae Martynova 1958 [synonym], Cycloristidae Martynova 1958 [synonym], Tomiochoristidae Martynova 1958 [synonym]
Fm. †Karaungiridae Novokshonov 1997
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G. †Karaungira Novokshonov and Sukacheva 1993
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†Karaungira shcherbakovi Novokshonov and Sukacheva 1993
G. †Liassopanorpa Martynov 1937
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†Liassopanorpa crassinervis Martynov 1937
G. †Mecaenigma Novokshonov and Sukacheva 2001
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†Mecaenigma suspectum Novokshonov and Sukacheva 2001
†Mecaenigma viduum Novokshonov and Sukacheva 2001
Fm. †Mendozachoristidae Brauckmann et al. 2010
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G. †Mendozachorista Brauckmann et al. 2010
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†Mendozachorista volkheimeri Brauckmann et al. 2010
Superfm. †Mesopsychoidea Tillyard 1917
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Fm. †Aneuretopsychidae Rasnitsyn and Kozlov 1990
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G. †Aneuretopsyche Rasnitsyn and Kozlov 1990
G. †Burmopsyche Zhao et al. 2020
G. †Jeholopsyche Ren et al. 2011
Fm. †Mesopsychidae Tillyard 1917
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G. †Baissopsyche Novokshonov and Sukacheva 2001
G. †Epicharmesopsyche Shih et al. 2013
G. †Ferghanopsyche Martynov 1937
G. †Lichnomesopsyche Ren et al. 2010
G. †Mesopanorpodes Tillyard 1918
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Invalid names: Mesopanorpa Tillyard 1918 [replaced]
G. †Mesopsyche Tillyard 1917
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Invalid names: Aristopsyche Tillyard 1919 [synonym], Bittacopanorpa Zalessky 1935 [synonym], Neuropsyche Tillyard 1919 [synonym], Triassopsyche Tillyard 1917 [synonym]
G. †Mesoses Riek 1976
G. †Permopsyche Bashkuev 2011
G. †Ptychopteropsis Martynov 1937
G. †Sogdopsyche Martynov 1937
G. †Tarantogus Sukatsheva 1985
G. †Tipulidites Wieland 1925
G. †Turanopsyche Martynov 1937
G. †Undisca Sukacheva 1990
G. †Vitimopsyche Novokshonov and Sukacheva 2001
G. †Xinjiangia Hong 1983
Invalid names: Mesopanorpodidae Tillyard 1918 [synonym], Mesosetidae Riek 1976 [synonym]
Fm. †Nedubroviidae Bashkuev 2011
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G. †Nedubrovia Bashkuev 2011
G. †Paranedubrovia Bashkuev 2011
Fm. †Pseudopolycentropodidae Handlirsch 1920
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G. †Dualula Lin et al. 2019
G. †Parapolycentropus Grimaldi and Rasnitsyn 2005
G. †Pseudopolycentropodes Grimaldi and Fraser 2005
G. †Pseudopolycentropus Handlirsch 1906
G. †Sinopolycentropus Shih et al. 2011
Invalid names: Dualulidae Lin et al. 2019 [synonym]
G. †Miriholcorpa Wang et al. 2013
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†Miriholcorpa forcipata Wang et al. 2013
Subor. Nannomecoptera Hinton 1981
+
Fm. Nannochoristidae Tillyard 1917
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G. †Afrochoristella Pinto and de Ornellas 1978
G. †Dahurochorista Sukatsheva 1985
G. †Dahurolarva Sukatsheva 1985
G. †Itaphlebia Sukatsheva 1985
+
Invalid names: Chrysopanorpa Ren 1995 [synonym], Netropanorpodes Sun et al. 2007 [synonym], Protochoristella Sun et al. 2007 [synonym], Stylopanorpodes Sun et al. 2007 [synonym]
G. †Jichoristella Ren 1995
G. †Namdyrus Sukacheva 1993
G. Nannochorista Tillyard 1917
Fm. †Robinjohniidae Martynova 1948
+
G. †Minusinia Novokshonov 1994
G. †Robinjohnia Martynova 1948
G. †Necropsis Hong 1983
+
†Necropsis paludis Hong 1983
Subor. †Paratrichoptera Tillyard 1919
+
Fm. †Liassophilidae Tillyard 1933
+
G. †Garazhovka Novokshonov 2002
G. †Izjumus Novokshonov 2002
G. †Laurentiptera Martynova and Willmann 1978
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Invalid names: Pseudodiptera Laurentiaux and Grauvogel 1953 [replaced]
G. †Liassophila Tillyard 1933
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Invalid names: Ijapsyche Kolosnitsyna and Martynova 1961 [synonym]
Invalid names: Laurentipteridae Martynova and Willmann 1978 [synonym], Pseudodipteridae Martynova 1961 [synonym]
Fm. †Permocentropidae Martynov 1932
+
G. †Permocentropus Martynov 1932
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Fm. †Permochoristidae Tillyard 1918
+
Subfm. †Agetopanorpinae Carpenter 1930
+
G. †Agetopanorpa Carpenter 1930
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Invalid names: Agetochorista Martynov 1932 [synonym], Agetochoristella Riek 1953 [synonym], Neoageta Riek 1953 [synonym]
G. †Argentinopanorpa Lara et al. 2015
G. †Gigantoageta Novokshonov 1993
G. †Mezena Novokshonov 1995
G. †Oochorista Martynov 1932
G. †Ovomezena Novokshonov 1995
G. †Parageta Novokshonov 1995
G. †Phipoides Riek 1953
G. †Protochorista Tillyard 1926
G. †Protopanorpa Tillyard 1926
+
Invalid names: Protopanorpoides Martynova 1962 [synonym]
G. †Seniorita Novokshonov 1993
G. †Stigmarista Martynova 1958
G. †Tshekardopanorpa Novokshonov 1993
G. †Duraznochorista Lara and Bashkuev 2020
+
†Duraznochorista zavattierii Lara and Bashkuev 2020
G. †Hongchoristites Rivaz Hernández 2018
+
Invalid names: Choristites Hong 2005 [replaced]
G. †Kalochorista Lin 1992
+
G. †Kazakhopanorpa Novokshonov 1995
+
†Kazachopanorpa oblonga Martynova 1961
†Kazakhopanorpa dubia Riek 1953
†Kazakhopanorpa jucunda Tillyard 1926
†Kazakhopanorpa terminata Novokshonov 1995
G. †Mesageta Novokshonov 1997
+
†Mesageta gigantea Novokshonov 1997
†Mesageta ignava Novokshonov 1997
†Mesageta insana Novokshonov 1997
†Mesageta pertrita Novokshonov 1997
†Mesageta rieki Novokshonov 1997
Subfm. †Permochoristinae Tillyard 1917
+
G. †Angustichorista Novokshonov 1995
G. †Archepetromantis Novokshonov 1995
G. †Asiachorista Martynova 1958
G. †Baianochorista Novokshonov 1997
G. †Callietheira Martynova 1958
G. †Echinopetromantis Novokshonov 1997
G. †Malmyzha Novokshonov 1995
G. †Mecolusor Novokshonov 2001
Tr. †Mesochoristini Tillyard 1926
G. †Neopetromantis Riek 1953
G. †Petrochorista Martynov 1931
+
Invalid names: Petrochoristella Willmann 1989 [synonym]
Tr. †Petromanteini Handlirsch 1937
G. †Shaanxichorista Hong 2005
G. †Tjulkinella Novokshonov 1995
G. †Tychtopsyche Martynova 1958
+
Invalid names: Tychtodelopterum Martynova 1958 [synonym]
G. †Xenochorista Riek 1953
Subfm. †Pseudonannochoristinae Novokshonov 1994
+
G. †Chaohuchorista Lian et al. 2022
G. †Miomeca Novokshonov 1994
G. †Nannochoristella Riek 1953
G. †Neochoristella Riek 1953
G. †Pseudonannochorista Novokshonov 1994
G. †Qingochorista Guo and Hong 2003
+
†Qingochorista conjunctiva Guo and Hong 2003
Subfm. †Sylvopanorpinae Novokshonov 1997
+
G. †Pulcheripanorpa Novokshonov 1993
G. †Sylvopanorpa Martynov 1940
G. †Uraloageta Novokshonov 1993
Invalid names: Caenoptilonidae Zalessky 1933 [synonym], Eosetidae Tindale 1945 [synonym], Idelopanorpidae Zalessky 1929 [synonym], Petrochoristidae Willmann 1989 [synonym], Protochoristinae Martynova 1961 [empty], Protopanorpidae Handlirsch 1937 [synonym], Tychtodelopteridae Ponomarenko 1977 [synonym], Tychtopsychidae Martynova 1958 [synonym], Xenochoristidae Riek 1953 [synonym]
Fm. †Permopanorpidae Tillyard 1926
+
G. †Martynopanorpa Willmann 1989
+
†Martynopanorpa angustata Martynov 1932
†Martynopanorpa zekkeli Martynova 1948
G. †Neopermopanorpa Riek 1955
+
G. †Permopanorpa Tillyard 1926
+
†Permopanorpa formosa Tillyard 1926
†Permopanorpa inaequalis Tillyard 1926
+
Invalid names: Permopanorpa dunbari Tillyard 1926 [synonym], Permopanorpa gracilis Tillyard 1926 [synonym], Permopanorpa raymondi Carpenter 1926 [synonym], Permopanorpa sellardsi Tillyard 1926 [synonym], Permopanorpa tenuis Tillyard 1926 [synonym]
†Permopanorpa martynovi Willmann 1978
+
Invalid names: Permopanorpa tenuis Martynov 1932 [replaced]
†Permopanorpa schucherti Tillyard 1926
G. †Xenopanorpa Riek 1955
+
Invalid names: Lithopanorpidae Carpenter 1930 [synonym], Martynopanorpidae Willmann 1989 [synonym]
Fm. †Permotanyderidae Riek 1953
+
Subfm. †Chimaeropterinae Bashkuev 2022
+
G. †Chimaeropteron Bashkuev 2022
Subfm. †Permotanyderinae Riek 1953
+
G. †Permotanyderites Bashkuev 2022
G. †Permotanyderus Riek 1953
+
Invalid names: Choristotanyderus Riek 1953 [synonym]
G. †Rudimentanus Bashkuev 2022
Fm. †Permotipulidae Tillyard 1929
+
G. †Permila Willmann 1989
+
†Permila borealis Martynova 1961
G. †Permotipula Tillyard 1929
+
†Permotipula patricia Tillyard 1929
Invalid names: Permilidae Willmann 1989 [synonym]
Subor. Pistillifera Willmann 1987
+
Infraor. Opisthogonopora Willmann 1987
+
Infraor. Raptipedia Willmann 1987
+
Fm. Bittacidae Handlirsch 1906 [hangingfly]
Fm. †Cimbrophlebiidae Willmann 1977 [hangingfly]
G. †Reprehensa Bode 1953
+
†Reprehensa acuminata Bode 1953
Fm. †Sibiriothaumatidae Sukatsheva and Novokshonov 1998
+
G. †Sibiriothauma Sukatsheva and Novokshonov 1998
+
†Sibiriothauma jakutensis Sukatsheva and Novokshonov 1998
Fm. †Sinoagetopanorpidae Lian et al. 2023
+
G. †Permoagetopanorpa Lian et al. 2023
+
†Permoagetopanorpa incompleta Lian et al. 2023
†Permoagetopanorpa yinpingensis Lian et al. 2023
G. †Raragetopanorpa Lian et al. 2023
+
†Raragetopanorpa zhangi Lian et al. 2023
G. †Sinoagetopanorpa Lin et al. 2010
+
†Sinoagetopanorpa elegans Lian et al. 2023
†Sinoagetopanorpa grimaldii Lian et al. 2023
†Sinoagetopanorpa lini Lian et al. 2023
†Sinoagetopanorpa magna Lian et al. 2023
†Sinoagetopanorpa minuta Lian et al. 2023
†Sinoagetopanorpa nigra Lian et al. 2023
†Sinoagetopanorpa permiana Lin et al. 2010
†Sinoagetopanorpa rotunda Lian et al. 2023
G. †Solusipanorpa Lin 1980
+
Fm. †Thaumatomeropidae Willmann 1978
+
G. †Blattomerope Rasnitsyn 1974
+
†Blattomerope polyneura Rasnitsyn 1974
G. †Thaumatomerope Rasnitsyn 1974
+
†Thaumatomerope madygenica Rasnitsyn 1974
†Thaumatomerope minuta Rasnitsyn 1974
†Thaumatomerope oligoneura Rasnitsyn 1974
†Thaumatomerope sinensis Lian and Huang 2023
†Thaumatomerope sogdiana Rasnitsyn 1974
Fm. †Volitorididae Hong 1982
+
G. †Volitoridia Hong 1982
+
†Volitoridia fulvis Hong 1982
Fm. †Worcestobiidae Soszyńska-Maj et al. 2018
+
G. †Worcestobia Soszyńska-Maj et al. 2018
+
†Worcestobia gigantea Tillyard 1933
†Worcestobia haradai Ueda 1991
Invalid names: Aneuretopsychina Rasnitsyn and Kozlov 1990 [invalid subgroup], Cyclopterinidae Carpenter 1986 [empty], Eobanksiidae Cockerell 1909 [empty], Panorpata Brauer [synonym], Paramecoptera Tillyard 1919 [empty], Protodiptera Tillyard 1937 [invalid subgroup], Protomecoptera Tillyard 1917 [empty]
+
G. †Curvilarva Novokshonov 2004
+
†Curvilarva tshekardensis Novokshonov 2004
Or. Siphonaptera Latreille 1825 [flea]
+
Fm. Ctenophthalmidae Rothschild 1915
+
Subfm. Ctenophthalminae Rothschild 1915
+
G. Palaeopsylla Wagner 1903
G. †Niwratia Jell and Duncan 1986
+
†Niwratia elongata Jell and Duncan 1986
+
G. Pulex Linnaeus 1758
+
†Pulex larimerius Lewis and Grimaldi 1997
Subfm. Spilopsyllinae Oudemans 1909
+
Tr. †Atopopsyllini Poinar 2015
Tr. Spilopsyllini Oudemans 1909
Fm. Rhopalopsyllidae Oudemans 1909
+
G. Rhopalopsyllus Baker 1905
Superfm. †Saurophthiroidea Ponomarenko 1986
+
Fm. †Pseudopulicidae Gao et al. 2012
+
G. †Hadropsylla Huang et al. 2013
G. †Pseudopulex Gao et al. 2012
G. †Tyrannopsylla Huang et al. 2013
Fm. †Saurophthiridae Ponomarenko 1986
+
G. †Saurophthirus Ponomarenko 1976
Invalid names: Saurophthiridae Gao et al. 2013 [objective synonym]
Fm. †Tarwiniidae Huang et al. 2013
+
G. †Tarwinia Jell and Duncan 1986
Invalid names: Hystrichopsyllidae Tiraboschi 1904 [empty]
Diagnosis
No diagnoses are available
Measurements
No measurements are available
Composition: chitinc
Environment: terrestrialc
Locomotion: actively mobilep
Created: 2017-04-17 09:56:18
Modified: 2017-04-17 09:56:18
Source: c = class, p = phylum
References: Bush and Bambach 2015, Kiessling 2004

Age range

Maximum range based only on fossils: base of the Westphalian A to the top of the Holocene or 318.70000 to 0.00000 Ma
Minimum age of oldest fossil (stem group age): 316.9 Ma

Collections (818 total)

Oldest occurrences

Time interval Ma Country or state Original ID and collection number
Westphalian A318.7 - 316.9France Westphalomerope maryvonneae (122506)
Moscovian315.2 - 307.0Czech Republic Phryganea arenacea, Phryganea kolbi (125662)
Westphalian C314.6 - 309.8Czech Republic Phryganea solitaria (155300)
Westphalian D309.8 - 307.5Germany (Lower Saxony) Microcarbonella paradoxa (224159)
Asselian298.9 - 295.5USA (New Mexico) Mecoptera indet. (124386)
Asselian298.9 - 295.5Czech Republic Pseudomerope gallei (114141) Stenomerope spinari, Pseudomerope havlati, Pseudomerope oborana, Pseudomerope mareki, Pseudomeropella nekvasilovae, Protomeropidae indet., Microptysmella moravica, Moravochorista carolina (114186)
Asselian298.9 - 295.5Germany Glanomerope virgoferroa (112604)
Asselian - Kungurian298.9 - 272.3Russian Federation Minusinia martynovae (126338)
Artinskian290.1 - 279.3USA (Oklahoma) Permopanorpa inaequalis (107405)
Artinskian290.1 - 279.3USA (Kansas) Permopanorpa raymondi (151412) Platychorista venosa, Permopanorpa inaequalis, Protochorista tetraclada, Agetopanorpa maculata, Protopanorpa permiana (111217) Protomerope permiana, Platychorista venosa, Permopanorpa dunbari, Permopanorpa tenuis, Permopanorpa gracilis, Permopanorpa sellardsi, Permopanorpa inaequalis, Permopanorpa formosa, Permopanorpa schucherti, Protopanorpa pusilla, Protochorista pentaclada, Protochorista tetraclada, Protopanorpa permiana, Anormochorista oligoclada (111219)
Artinskian - Kungurian290.1 - 272.3Russian Federation (Kemerovo) Mesochorista javorskyi, Asiachorista neuburgae (124614)
Kungurian279.3 - 272.3Russian Federation (Suksun) Kamopanorpa fasciipennis, Agetochorista intermedia (116652) Kamopanorpa fasciipennis, Parachorista uralensis (113379) Marimerobius sp., Kamopanorpa fasciipennis, Permochorista sylvaensis, Agetochorista tillyardi, Agetochorista permiana, Agetochorista sp., Sylvopanorpa carpenteri (116641) Marimerobius sp., Protomeropidae indet., Kamopanorpa pritykinae, Kamopanorpa incerta, Kamopanorpa pallida, Uraloptysma maculata, Kamopanorpa uralensis, Petromantis sylvaensis, Tshekardopanorpa biarmica, Phipoides ornatus, Phipoides pusillus, Agetochorista kungurica, Agetochorista tillyardi, Agetochorista punctata, Agetochorista permiana, Agetochorista sp., Agetochorista intermedia, Agetopanorpa sp., Protopanorpoides medius, Protopanorpoides minutus, Uraloageta archaica (108206) Parachorista uralensis (116642) Protomeropidae indet. (122864) Protomeropidae indet., Curvilarva tshekardensis (122753) Protomeropidae indet., Kamopanorpa fasciipennis, Parachorista uralensis (124241)
Kungurian279.3 - 272.3Russian Federation (Perm) Kamopanorpa sp. (173727) Kamopanorpa tyulkiana, Kamopanorpa tamarae, Kamopanorpa sp., Tjulkinella silantjevi, Protopanorpa similis, Protopanorpa longicubitalis, Protopanorpa minuta, Protopanorpa sp. (103639)
Kungurian279.3 - 272.3Russian Federation (Kemerovo) Asiachorista neuburgae, Asiachorista migdisovae (124615) Asiachorista tuberculata, Asiachorista neuburgae, Agetochorista sarbalensis (124613) Microptysma sibiricum, Pinnachorista sarbalensis, Tomiochorista prodigialis, Mesochorista prospera, Petromantis opinabilis, Petromantis sarbalensis, Petromantis nana, Petromantis unicolor, Callietheira sp., Asiachorista cellulosa, Callietheira major, Asiachorista gorelovae, Asiachorista lativenosa (112708) Pinnachorista imitabilis, Petromantis latimargo, Petromantis unicolor (104951)
Kungurian279.3 - 272.3Russian Federation Marimerobius sp., Tshekardopanorpa magasa, Seniorita gratiosa, Phipoides pusillus, Protopanorpoides medius, Agetopanorpidae indet. (107787) Marimerobius splendens (155819) Marimerobius sukatchevae, Marimerobius sp., Kamopanorpa uralensis, Agetochorista tillyardi, Agetopanorpa sp., Protopanorpa sp., Pulcheripanorpa aliena (114207) Petromantis sp., Protopanorpa sp. (115214) Protomeropidae indet. (102504)
Kungurian279.3 - 272.3Russian Federation (Komi) Adoptysma anomalanala (97621)
Kungurian279.3 - 272.3South Africa (West Cape) Mecoptera indet. (117279)
Kungurian279.3 - 272.3Brazil (Rio Grande do Sul) Petromantis evansi, Asiachorista beckermigdisovae (116121) Petromantis rieki (116120)
Roadian272.3 - 268.8Russian Federation (Kazan) Petrochorista minuta, Petrochorista elegantula, Petrochorista parvula, Petromantis major, Permochorista rossica (116803) Petromantis rossica (116877)
Roadian272.3 - 268.8Russian Federation (Kemerovo) Altajopanorpa kaltanica (124605) Eumecoptera indet. (124616) Kaltana pilosa, Mesochorista javorskyi (122571) Kaltana pilosa, Petromantis oreshnikovae, Asiachorista kaltanica (122572) Microptysma sibiricum, Altajopanorpa suchanovi, Pinnachorista fausta, Pinnachorista cubitalis, Pinnachorista migdisovi, Pinnachorista latipennis, Pinnachorista rotundipennis, Tomiochorista nubila, Tomiochorista rhombica, Tomiochorista minuta, Kaltanochorista grjasevi, Altajopanorpa elegans, Altajopanorpa grandis, Altajopanorpa cellulosa, Altajopanorpa rhombica, Kaltana pilosa, Altajopanorpa brevis, Cyclorista convexicosta, Petromantis grandis, Petromantis kaltanica, Petromantis sarbalensis, Petromantis oreshnikovae, Petromantis karpenkoae, Callietheira khalfini, Asiachorista permiana, Asiachorista tuberculata, Callietheira bimembris, Asiachorista albibasalis, Asiachorista neuburgae, Asiachorista migdisovae, Asiachorista kaltanica (122568) Pinnachorista gigantea, Tomiochorista skoki, Tomiochorista sp., Altajopanorpa nigronervosa, Kaltana pilosa, Mesochorista pinnigera, Petrochorista recta, Petromantis kaltanica, Petrochorista prona, Petromantis oreshnikovae (122569) Pinnachorista ovalis, Altajopanorpa postrema, Cyclorista simplex, Pseudonannochorista willmanni, Miomeca reducta, Parageta parachoristoides, Ovomezena lata (104704) Pseudonannochorista willmanni (122567)