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Insecta -

Synonymy list
YearName and author
2004Coleopterida Kukalová-Peck and Lawrence
2006Coleopterida Beutel and Pohl
2014Coleopterida Misof et al.

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EubilateriaAx 1987
phylumArthropodaLatreille 1829
superclassHexapodaLatreille 1825
NeopterygotaCrampton 1924
infraclassNeopteraMartynov 1923

If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.

No diagnoses are available
No measurements are available
Composition: chitinc
Environment: terrestrialc
Locomotion: actively mobilep
Created: 2017-04-17 09:56:18
Modified: 2017-04-17 09:56:18
Source: c = class, p = phylum
References: Kiessling 2004, Bush and Bambach 2015

Age range

Maximum range based only on fossils: base of the Gzhelian to the top of the Holocene or 303.70000 to 0.00000 Ma
Minimum age of oldest fossil (stem group age): 303.4 Ma

Collections (1839 total)

Oldest occurrences

Time interval Ma Country or state Original ID and collection number
Stephanian B304.8 - 303.4France Blanzelytron vetteri (172760)
Gzhelian303.7 - 298.9France Stephanastus polinae (121979)
Stephanian C303.4 - 301.2Germany (Thuringia) Blattocoleus tillyardi (152999)
Asselian298.9 - 295.5Germany (Rhineland-Palatinate) Coleopsis archaica (156248)
Sakmarian295.5 - 290.1Czech Republic Tshekardocoleidae indet., Blattelytridae indet., Moravocoleus fractus, Moravocoleus neglegens, Moravocoleus perditus, Apachelytron transversum, Moravocoleus permianus, Eocoleus scaber, Umoricoleus perplex, Boscoleus blandus, Votocoleus submissus, Prosperocoleus prosperus, Retelytron conopeum, Oborocoleus rohdendorfi, Liberocoleus intactus, Ortelytron europaeum, Ortelytron artum, Crasselytron convexum, Rugelytron fuscum, Venelytron tuberculatum, Planelytron planum, Glabelytron lativenosum (114186)
Sakmarian295.5 - 290.1Germany (Thüringer Wald) Pachypleurites nodosus (128485)
Artinskian290.1 - 279.3South Africa (Western Cape) Afrocupes firmae (156986)
Artinskian290.1 - 279.3USA (Kansas) Elytroneura permiana (111217) Parablattelytron rectum (21979) Parablattelytron subincisum (114450) Protelytron permianum, Parablattelytron rectum, Protelytron furcatum, Parablattelytron elongatum (114152) Protelytron permianum, Parablattelytron subincisum, Parablattelytron rectum, Permelytron schucherti, Protelytron furcatum (114151) Protelytron permianum, Protelytron angustum, Permelytropsis cubitalis, Archelytron superbum, Acosmelytron elongatum, Parablattelytron subincisum, Parablattelytron rectum (111319) Protelytron permianum, Protelytron tillyardi, Acosmelytron elongatum, Parablattelytron subincisum, Parablattelytron latum, Parablattelytron rectum (111320) Protelytron sp., Protelytron permianum, Acosmelytron sp., Acosmelytron elongatum, Acosmelytron acutum, Acosmelytron delicatum, Parablattelytron sp., Parablattelytron subincisum, Parablattelytron rectum, Protelytron dunbari, Protelytropsis grandis, Permelytron schucherti, Megelytron robustum, Blattelytron permianum (111219)
Artinskian290.1 - 279.3USA (Oklahoma) Permocoleus wellingtonensis (104816 104826)
Kungurian279.3 - 272.3Russian Federation (Suksun) Sylvacoleodes admirandus, Uralelytron martynovi, Sylvacoleus richteri, Tshekardocoleus minor, Sylvacoleus sharovi, Uralelytron insignis (108206) Uralelytron martynovi, Tshekardocoleus magnus (116652)
Kungurian279.3 - 272.3Brazil (Rio Grande do Sul) Kaltanicupes ponomarenkoi, Protocupoides rohdendorfi (116121)
Kungurian279.3 - 272.3Russian Federation Maricoleus valentinae (115214) Tshekardocoleus magnus, Sylvacoleus richteri (107787) Uralocoleus splendens, Bardocoleus insignis (155819)
Roadian272.3 - 268.8Russian Federation (Kemerovo) Cytocupoides elongatus, Archicupes jacobsoni, Protocupes martynovi, Eocupes lukjanovitshi, Rossocoleus avus (122567) Ichthyocupes kuznetskiensis (122572) Kaltanicupes richteri (122568) Kaltanocoleus pospelovi, Protocupes martynovi (122569) Palaeocupes kaltanicus, Eocupes cellulosus, Kaltanocoleus pospelovi, Archicupes reichardti, Stegocupes efremovi, Carinicupes beckermigdisovae, Asiocoleus novojilovi, Cytocupes angustus, Protocupoides plavilstshikovi, Tomiocupes sharovi, Schizotaldycupes ananjevi, Ichthyocupes tyzhnovi, Uskatocoleus kaltanicus (104704) Schizocoleus kuznetskiensis (124604)
Roadian272.3 - 268.8Russian Federation (Kirov) Kaltanicupes major, Rossocoleus altus, Kaltanicupes kitjakensis, Kaltanicupes acutus (105058)
Roadian272.3 - 268.8Russian Federation (Kazan) Permocupes semenovi, Permocupoides distinctus (116877)
Roadian272.3 - 268.8Russian Federation (Arkangelsk) Arctocoleus ivensis (111187) Arctocoleus ivensis, Permocrossos elongatus (104993) Permocupes semenovi (104716) Permocupes sojanensis, Permocupes grandis (116933) Sojanocoleus reticulatus (116874) Uralocoleus ultimus, Permocupes sojanensis (104992)
Roadian272.3 - 268.8Russian Federation Elateridopsis permiensis (156163)
Wordian268.8 - 265.1Russian Federation (Kemerovo) Palademosyne martynovae (124612) Protocupes ogloblini, Taldycupes chachlovi, Tychticupes tchihatcheffi, Tecticupes indistinctus, Stegocupes fedotovi, Tecticupes heckeri, Uskatocoleus avitus, Uskatocoleus iljinskiensis, Uskatocoleus surijokovicus, Karakanocoleus janischewskyi, Rossocoleus surijokovensis, Rossocoleus grandis, Rossocoleus parvus, Rossocoleus latus, Rossocoleus latior (122566) Schizocupes obrutshevi (124600) Uskatocoleus convexus, Rossocoleus verrucosus (124603)
Wordian - Capitanian268.8 - 259.9Russian Federation Uskatocoleus kushejakovicus (124610)
Urzhumian268.0 - 265.0Russian Federation (Udmurtia) Coleoptera hindwing indet., Schizophoridae indet., Permocupes latus, Protocupoides elongatus, Rossocoleus angustus, Rossocoleus novojilovi (103502) Schizotaldycupes pubescens, Rhombocoleites uralensis (104750)