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Holmiella
Taxonomy
Holmiella was named by Fritz (1972) [Sepkoski's age data: Cm Atda-u Sepkoski's reference number: 475,885]. Its type is Holmiella preancora.
It was assigned to Olenellida by Sepkoski (2002); to Holmiinae by Palmer and Repina (1993), Lieberman (1999) and Ebbestad et al. (2003); and to Holmiidae by Whittington et al. (1997) and Hollingsworth (2006).
It was assigned to Olenellida by Sepkoski (2002); to Holmiinae by Palmer and Repina (1993), Lieberman (1999) and Ebbestad et al. (2003); and to Holmiidae by Whittington et al. (1997) and Hollingsworth (2006).
Species
Synonymy list
| Year | Name and author |
|---|---|
| 1972 | Holmiella Fritz |
| 1993 | Holmiella Palmer and Repina p. 26 fig. 6.4.6 |
| 1997 | Holmiella Whittington et al. p. 415 |
| 1999 | Holmiella Lieberman p. 86 |
| 2002 | Holmiella Sepkoski |
| 2003 | Holmiella Ebbestad et al. |
| 2006 | Holmiella Hollingsworth p. 328 |
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If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.
G. †Holmiella Fritz 1972
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†Holmiella domackae Gapp and Lieberman 2014
†Holmiella falcuta Fritz 1972
†Holmiella falx Hollingsworth 2006
†Holmiella millerensis Hollingsworth 2006
†Holmiella preancora Fritz 1972
†Holmiella taurus Gapp and Lieberman 2014
Diagnosis
| Reference | Diagnosis | |
|---|---|---|
| A. R. Palmer and L. N. Repina 1993 | Holmiinae with outline of cephalon subpentagonal; genal spine prominent, originating oppo- site or anterior to S3; short intergenal spine or node lo- cated at or on axial side of intergenal angle. Glabella expanded anteriorly. Posterior tips of ocular lobes opposite or posterior to Li. Pygidium large, wider than long, of at least 3 segments; anterior two segments extended into short border spines. | |
| B. S. Lieberman 1999 | Anterior cephalic border developed as flattened plane, prominently separated from extraocular area by furrow; frontal lobe contacts anterior border furrow, lateral mar- gins distal to lateral margins of L0, expands prominently dorsally; anterolateral margins of glabella prominently separated from extraocular area by furrow; length (sag.) of frontal lobe 1 to 1.1 times length (sag.) of L0 and L1; dorsal notch-like truncation in ocular lobes where they contact frontal lobe of the glabella; anterodistal margins of L3 formed by axial furrows; distal margins of L3 straight; S3 evenly convex, medial and distal tips as far for- ward; lateral margins of glabella opposite L2 subparallel; S1, S2 and S3 not conjoined me- dially; medial and distal tips of S2 as far forward; line from posterior edge of ocular lobe to junction of lobe with glabella forms roughly 10 to 15 degree angle with sagittal line; poste- rior margin of ocular lobe opposite distal tip of S0; lateral lobes present on L0; node or spine on L0 posterior of midline; intergenal angle with small spine directly behind distal tip of ocular lobe; intergenal angle forms 80 to 90 degree angle with transverse line; extraocu- lar region prominently vaulted, very narrow, width (tr.) at L1 10% to 15% width of glabella at L1; genal spine angle opposite distal tip of L3 or L4; lateral margin of genal spine near spine angle forms roughly 60 to 70 degree angle relative to sagittal line; pygidium twice as wide (tr.) as long (sag.), posterior margin notched. | |
| J. S. Hollingsworth 2006 | Holmiid with advanced genal spines ini- tially directed outward to slightly forward from anterolateral mar- gin of cephalon, posterolateral margin rounded, intergenal spines or node a short distance in from posterolateral corner, or may be absent. Glabellar sides concave, LA expanded and front broadly curved against anterior border furrow. Ocular lobes long. Extra- ocular area ranges from wider than interocular area to much less than interocular area width. Small occipital spine present on some species. Thorax of 17 segments, pleural regions slightly narrower than axis. Pygidium large for a holmiid, wide, one axial ring clearly defined and a second defined by pits and a shallow furrow. Posterior margin transverse, bearing short spines. |