PBDB

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Boreoeutheria

Mammalia

Taxonomy

Boreoeutheria is an unranked clade. It is extant.

It was assigned to Placentalia by Asher and Helgen (2010).

Subtaxa

Euarchontoglires (syn. Archontoglires, Supraprimates), Laurasiatheria (syn. Laurasiaplacentalia)

Sister taxa

Euarchontoglires (syn. Archontoglires, Supraprimates), Laurasiatheria (syn. Laurasiaplacentalia)

Synonyms

Boreotheria is an unranked clade. It is extant.

It was synonymized subjectively with Boreoeutheria by Asher and Helgen (2010).

It was assigned to Exafroplacentalia by Waddell et al. (2001).
Boreoplacentalia is an unranked clade. It is extant.

It was synonymized subjectively with Boreoeutheria by Asher and Helgen (2010).

Synonymy list

Year	Name and author
2001	Boreotheria Waddell et al. p. 148
2010	Boreoeutheria Asher and Helgen p. 3 figs. Figure 1

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Rank	Name	Author
kingdom	Animalia	()
—	Bilateria
—	Eubilateria	Ax 1987
—	Deuterostomia
phylum	Chordata	Haeckel 1874
subphylum	Vertebrata
superclass	Gnathostomata
—	Osteichthyes	()
subclass	Sarcopterygii	()
subclass	Dipnotetrapodomorpha	(Nelson 2006)
subclass	Tetrapodomorpha	()
—	Tetrapoda
—	Reptiliomorpha
—	Anthracosauria
subclass	Amphibiosauria	Kuhn 1967
—	Cotylosauria	()
—	Amniota
subclass	Synapsida

Rank	Name	Author
suborder	Eupelycosauria
—	Therapsida	()
infraorder	Cynodontia	()
—	Mammaliamorpha
—	Mammaliaformes
class	Mammalia
—	Theriamorpha	(Rowe 1993)
—	Theriiformes	()
—	Trechnotheria
—	Cladotheria
—	Zatheria
subclass	Tribosphenida	()
subclass	Theria
—	Eutheria	()
—	Placentalia
—	Boreoeutheria
—	Boreoeutheria

If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.

Unr. Boreoeutheria

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Unr. Euarchontoglires Murphy and Eizirik 2001

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Or. †Anagalida Szalay and McKenna 1971

Fm. †Anagalidae Simpson 1931

G. †Anagale Simpson 1931

†Anagale gobiensis Simpson 1931

G. †Anagalopsis Bohlin 1951

†Anagalopsis kansuensis Bohlin 1951

G. †Anaptogale Xu 1976

†Anaptogale wanghoensis Xu 1976

G. †Chianshania Xu 1976

†Chianshania gianghuaiensis Xu 1976

G. †Diacronus Xu 1976

†Diacronus anhuiensis Xu 1976

Invalid names: Decoredon elongetus Xu 1977 [synonym]

†Diacronus wanghuensis Xu 1976

G. †Eosigale Hu 1993

†Eosigale gujingensis Hu 1993

G. †Hsiuannania Xu 1976

†Hsiuannania maguensis Xu 1976

†Hsiuannania minor Ding and Zhang 1979

†Hsiuannania tabiensis Xu 1976

G. †Huaiyangale Xu 1976

†Huaiyangale chianshanensis Xu 1976

†Huaiyangale leura Ding and Tong 1979

G. †Interogale Huang and Zheng 1983

†Interogale datangensis Huang and Zheng 1983

G. †Linnania Chow et al. 1973

†Linnania lofoensis Chow et al. 1973

†Linnania progressus Huang 2006

†Linnania qinglingensis Xue 1986

G. †Pseudictops Matthew et al. 1929

†Pseudictops lophiodon Matthew et al. 1929

G. †Qipania Hu 1993

†Qipania yui Hu 1993

G. †Stenanagale Wang 1975

†Stenanagale xiangensis Wang 1975

G. †Wanogale Xu 1976

†Wanogale hodungensis Xu 1976

G. †Zofiagale López-Torres and Fostowicz-Frelik 2018

†Zofiagale ergilinensis López-Torres and Fostowicz-Frelik 2018

Fm. †Astigalidae Zhang and Tong 1981

G. †Astigale Zhang and Tong 1981

†Astigale nanxiogensis Zhang and Tong 1981

†Astigale wanensis Zhang and Tong 1981

G. †Yupingale Tong and Wang 2006

†Yupingale weifangensis Tong and Wang 2006

G. †Zhujegale Zhang and Tong 1981

†Zhujegale jintangensis Zhang and Tong 1981

†Zhujegale lirenensis Zhang and Tong 1981

Fm. †Pseudictopidae Sulimski 1969

G. †Allictops Qiu 1977

G. †Anictops Qiu 1977

†Anictops tabiepedis Qiu 1977

†Anictops wanghudunensis Zheng et al. 1999

G. †Cartictops Ding and Tong 1979

G. †Haltictops Ding and Tong 1979

†Haltictops meilingensis Ding and Tong 1979

†Haltictops mirabilis Ding and Tong 1979

G. †Paranictops Qiu 1977

†Paranictops majuscula Qiu 1977

G. †Suyinia Tong and Wang 2006

†Suyinia changleensis Tong and Wang 2006

G. †Wania Wang 1995

†Wania chowi Wang 1995

Invalid names: Eurymyloidea Li 1977 [empty]

Superfm. †Apatemyoidea

Fm. †Apatemyidae Matthew 1909

Subfm. †Apatemyinae Matthew 1909

Invalid names: Heterohyins Gervais 1859 [synonym]

G. †Apatemys Marsh 1872

†Apatemys bellulus Marsh 1872

Invalid names: Apatemys hurzeleri Gazin 1962 [synonym], Teilhardella whitakeri Simpson 1954 [synonym]

†Apatemys bellus Marsh 1872

†Apatemys chardini Jepsen 1930

†Apatemys downsi Gazin 1958

†Apatemys hendryi Robinson 1966

†Apatemys mutiniacus Russell et al. 1979

†Apatemys prouti Hooker 2010

†Apatemys pygmaeus Beard and Dawson 2009

†Apatemys rodens Troxell 1923

†Apatemys sigogneauae Russell et al. 1979

†Apatemys sigogneaui Russell et al. 1979

†Apatemys teilhardi Russell et al. 1979

†Apatemys uintensis Matthew 1921

Invalid names: Stehlinius Matthew 1921 [synonym]

G. †Carcinella Von Koenigswald et al. 2009

†Carcinella sigei von Koenigswald et al. 2009

G. †Eochiromys Teilhard de Chardin 1927

†Eochiromys landenensis Teilhard de Chardin 1927

G. †Frugivastodon Bajpai et al. 2005

†Frugivastodon cristatus Bajpai et al. 2005

G. †Heterohyus Gervais 1848

Subg. †Heterohyus (Gervaisyus) Sudre et al. 1990

†Heterohyus armatus Gervais 1848

†Heterohyus morinionensis Hooker 1986

†Heterohyus nanus Teihard de Chardin 1916

†Heterohyus pygmaeus Sudre et al. 1990

†Heterohyus quercyi Filhol 1890

Invalid names: Heterochiromys gracilis Stehlin 1916 [synonym]

†Heterohyus sudrei Sigé 1975

Invalid names: Amphichiromys Stehlin 1916 [synonym], Heterochiromys Stehlin 1916 [synonym], Necrosorex Filhol 1890 [synonym]

G. †Jepsenella Simpson 1940

†Jepsenella praepropera Simpson 1940

G. †Labidolemur Matthew and Granger 1921

†Labidolemur kayi Simpson 1929

†Labidolemur major West 1972

†Labidolemur serus Gingerich 1982

†Labidolemur soricoides Matthew and Granger 1921

G. †Russellmys Estravís 1990

†Russellmys denisae Estravís 1990

G. †Sinclairella Jepsen 1934

†Sinclairella dakotensis Jepsen 1934

G. †Stehlinella Matthew 1929

G. †Teilhardella Jepsen 1930

G. †Unuchinia Simpson 1937

†Unuchinia asaphes Simpson 1936

†Unuchinia diaphanes Gunnell 1988

†Unuchinia dysmathes Holtzman 1978

Invalid names: Apator Simpson 1936 [replaced]

Invalid names: Russellmyinae Estravís 1990 [empty], Unuchiniinae Van Valen and McKenna 1967 [empty]

Unr. Euarchonta

Or. Primates Linnaeus 1758 [primate]

G. †Berruvius Russell 1964

†Berruvius lasseroni Russell 1964

G. †Ceciliolemur Weigelt 1933

G. †Chasselasia Schwartz and Tattersall 1982

†Chasselasia eldredgei Schwartz and Tattersall 1982

Subor. Cheiromyoidea Wortman 1903

Invalid names: Cheiromyidae Gray 1821 [empty]

Unr. Euprimateformes Bloch et al. 2007

Unr. Euprimates Hoffstetter 1978

Superfm. †Plesiadapoidea

Infraor. Haplorhini Pocock 1918 [monkey]

Unr. Anthropoidea Mivart 1864

Invalid names: Simiiformes Hofstetter 1977 [synonym]

Infraor. Tarsiiformes Gregory 1915 [tarsier]

G. †Megatarsius Weigelt 1933

Fm. †Micromomyidae Szalay 1974

G. †Chalicomomys Beard and Houde 1989

G. †Dryomomys Bloch et al. 2007

G. †Foxomomys Chester and Bloch 2013

G. †Micromomys Szalay 1973

G. †Tinimomys Szalay 1974

Invalid names: Myrmekomomys Robinson 1994 [synonym]

Invalid names: Micromomyinae Szalay 1974 [empty]

Fm. †Microsyopidae Osborn and Wortman 1892

Subfm. †Microsyopinae Osborn and Wortman 1892

G. †Nanomomys Rose et al. 2012

G. †Navajovius Matthew and Granger 1921

Subfm. †Uintasoricinae Szalay 1969

Invalid names: Navajoviinae Gunnell 1989 [invalid subgroup]

G. †Microtarsioides Weigelt 1933

G. †Nosmips Seiffert et al. 2010

†Nosmips aenigmaticus Seiffert et al. 2010

Fm. †Paromomyidae Simpson 1940

G. †Acidomomys Bloch et al. 2002

G. †Arcius Godinot 1984

G. †Dianomomys Tong and Wang 2006

G. †Edworthia Fox et al. 2010

G. †Ignacius Matthew and Granger 1921

Subfm. †Phenacolemurinae Simpson 1955

Invalid names: Palaechthonini Szalay 1969 [empty], Paromomyinae [objective synonym], Paromomyini Simpson 1940 [empty], Phenacolemur leonardi Storer 1990 [nomen dubium], Purgatoriinae Van Valen and Sloan 1965 [empty]

Subor. †Plesiadapiformes Simons and Tattersall 1972

Superfm. †Microsyopoidea

G. †Palaechthon Gidley 1923

Invalid names: Talpohenach Kay and Cartmill 1977 [synonym]

G. †Pandemonium Van Valen 1994

Fm. †Purgatoriidae Gunnell 1989

Invalid names: Paromomyoidea Simpson 1940 [empty]

Subor. Prosimiae Milne Edwards

Invalid names: Limnotheriidae Marsh 1875 [empty]

G. †Rooneyia Wilson 1966

†Rooneyia viejaensis Wilson 1966

Subor. Strepsirrhini Geoffroy Saint-Hilaire 1812

Infraor. †Adapiformes Hoffstetter 1977

Fm. †Azibiidae Gingerich 1976

Fm. †Djebelemuridae Hartenberger and Marandat 1992

G. †Karanisia Seiffert et al. 2003

Infraor. Lemuriformes Gregory 1915 [lemur]

Infraor. Lorisiformes Gregory 1915

Invalid names: Ceciliolemuridae Weigelt 1933 [invalid subgroup], Ceciliolemuroidea Weigelt 1933 [invalid subgroup], Lemures Hubrecht [empty], Pithecus Geoffroy and Cuvier 1795 [nomen nudum], Plesitarsiiformes Gingerich 1974 [empty], Pliopithecus posthumus Schlosser 1924 [nomen dubium], Prosimii Illiger 1811 [empty], Simia Linnaeus 1758 [nomen nudum], Simia (Cercopithecus) sinicus pileatus Kerr 1792 [nomen dubium], Simia apedia Linnaeus 1758 [nomen dubium], Simiolemuriformes [empty], Tarsii [empty], Tupaioidea Dobson 1882 [empty]

Unr. Primatomorpha Beard 1991

Or. Dermoptera Illinger 1811 [colugo]

G. †Dermotherium Ducrocq et al. 1992

G. †Elpidophorus Simpson 1927

Subor. Eudermoptera Beard 1993 [true dermopteran]

G. †Plagiomene Matthew 1918

G. †Planetetherium Simpson 1928

G. †Worlandia Bown and Rose 1979

Invalid names: Galeopithecia Jaekel 1911 [synonym], Galeopithecoidea Gray 1821 [empty], Ptenopleura Van der Hoeven 1858 [synonym], Pterophorae Gray 1821 [synonym]

Unr. Sundatheria

Fm. †Plagiomenidae Matthew 1918

G. †Ellesmene Dawson et al. 1993

Subfm. †Thylacaelurinae Van Valen 1967

Invalid names: Plagiomeninae Matthew 1918 [empty]

Or. Scandentia Wagner 1855 [treeshrew]

Fm. Tupaiidae Gray 1825 [tree shrew]

Invalid names: Ptilocercidae Lyon 1913 [empty]

Invalid names: Paraprimates [synonym]

Unr. †Gliriformes Wyss and Meng 1996

Fm. †Arctostylopidae Schlosser 1923

G. †Anatolostylops Zhai 1978

†Anatolostylops dubius Zhai 1978

†Anatolostylops zhaii Wang et al. 2008

Subfm. †Arctostylopinae Tong and Wang 2006

Tr. †Arctostylopini Tong and Wang 2006

Invalid names: Palaeostylopini Tong and Wang 2006 [empty]

G. †Arctostylops Matthew 1915

†Arctostylops steini Matthew 1915

Subfm. †Asiostylopinae Zheng 1979

G. †Asiostylops Zheng 1979

Subfm. †Bothriostylopinae Tong and Wang 2006

G. †Allostylops Zheng 1979

G. †Bothriostylops Zheng and Huang 1986

G. †Enantiostylops Averianov 2020

†Enantiostylops progressus Averianov 2020

Subfm. †Kazachostylopinae Nessov 1987

G. †Kazachostylops Nessov 1987

G. †Palaeostylops Matthew and Granger 1925

†Palaeostylops iturus Matthew and Granger 1925

†Palaeostylops macrodon Matthew et al. 1929

Invalid names: Gashatostylops Cifelli et al. 1989 [synonym]

G. †Sinostylops Tang and Yan 1976

†Sinostylops promissus Tang and Yan 1976

G. †Stenostylops Huang et al. 2001

†Stenostylops xiangensis Huang et al. 2001

G. †Wanostylops Huang and Zheng 1997

†Wanostylops promissus Huang 2003

†Wanostylops youngi Huang and Zheng 1997

Unr. Glires Linnaeus 1758

Fm. Aspalacidae Gray 1825

Tr. Pseudotomina Gray 1825

Invalid names: Arctomina Gray 1825 [empty], Cricetina Gray 1825 [empty]

Unr. Duplicidentata Illiger 1811

Or. Lagomorpha Brandt 1885

Fm. †Mimotonidae Li 1977

Fm. †Eurymylidae Matthew et al. 1929

G. †Eurymylus Matthew and Granger 1925

Invalid names: Baenomys Matthew and Granger 1925 [synonym]

G. †Heomys Li 1977

G. †Matutinia Li et al. 1979

G. †Rhombomylus Zhai 1978

G. †Taizimylus Mao et al. 2017

Subfm. †Zagmyinae Dashzeveg and Russell 1988

Invalid names: Eurymylinae Dashzeveg and Russell 1988 [empty], Hypsimylinae Dashzeveg and Russell 1988 [empty], Khaychininae Dashzeveg and Russell 1988 [empty]

Fm. Ierboidae Gray 1825

Tr. Sciurina Gray 1825

Invalid names: Dipina Gray 1825 [empty], Myoxina Gray 1825 [empty], Pedestina Gray 1825 [empty]

Or. †Mixodontia Sych 1971

Fm. †Decipomyidae Dashzeveg et al. 1998

Fm. †Rhombomylidae Li et al. 1987

Unr. Simplicidentata Flower 1883

G. †Eomylus Dashzeveg and Russell 1988

Invalid names: Khaychina Dashzeveg and Russell 1988 [synonym]

G. †Nikolomylus Shevyreva and Gabunia 1986

G. †Palaeomylus Meng et al. 2005

Or. Rodentia [rodent]

Invalid names: Haplodontidae Alston 1876 [empty]

G. †Sinomylus McKenna and Meng 2001

†Sinomylus zhaii McKenna and Meng 2001

Invalid names: Lagostomyidae [empty], Ochtodontidae Waterhouse 1840 [empty], Rodentiaformes Wyss and Meng 1996 [empty]

Superfm. †Mixodectoidea Simpson 1945

Fm. †Mixodectidae Cope 1883

G. †Dracontolestes Gazin 1941

†Dracontolestes aphantus Gazin 1941

G. †Eudaemonema Simpson 1935

†Eudaemonema bohachae Scott et al. 2013

†Eudaemonema cuspidata Simpson 1935

†Eudaemonema webbi Scott 2010

G. †Mixodectes Cope 1883

†Mixodectes malaris Cope 1884

Invalid names: Oxyacodon tecumsae Van Valen 1978 [synonym]

†Mixodectes pungens Cope 1883

Invalid names: Mixodectes crassiusculus Cope 1883 [synonym], Olbodotes copei Osborn 1902 [synonym]

Invalid names: Indrodon Cope 1884 [synonym], Olbodotes Osborn 1902 [synonym]

G. †Prosinopa Trouessart 1897

Invalid names: Oldobotidae Schlosser 1907 [synonym]

Invalid names: Archonta Gregory 1910 [empty], Archontoglires [synonym], Supraprimates Waddell et al. 2001 [synonym]

Unr. Laurasiatheria Waddell et al. 1999

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Unr. Eulipotyphla

Unr. Erinaceomorpha Gregory 1910

Fm. †Chambelestidae Gheerbrant and Hartenberger 1999

G. †Chambilestes Gheerbrant and Hartenberger 1999

G. †Diacodon Cope 1875

†Diacodon alticuspis Cope 1875

†Diacodon celatus Cope 1875

Invalid names: Anaptomorphus minimus Loomis 1906 [synonym]

Superfm. Erinaceoidea Gill 1872

G. †Amphidozotherium Filhol 1877

G. †Auroralestes Holroyd et al. 2004

Invalid names: Eolestes Bown and Schankler 1982 [replaced]

G. †Dartonius Novacek et al. 1985

G. †Diacocherus Gingerich 1983

Fm. †Dimylidae Schlosser 1887

G. †Leipsanolestes Simpson 1928

G. †Ghambilentes Unknown 2000

†Ghambilentes foussanensis Gheerbrant and Hartenberger 1999

G. †Quadratodon De Bast and Smith 2017

†Quadratodon sigei De Bast and Smith 2017

G. †Seia Russell and Gingerich 1981

†Seia shahi Russell and Gingerich 1981

Fm. †Sespedectidae Novacek 1985

Subfm. †Sespedectinae Novacek 1985

G. †Talpavoides Bown and Schankler 1982

†Talpavoides dartoni Bown and Schankler 1982

G. †Talpavus Marsh 1872

†Talpavus conjunctus Storer 1995

†Talpavus duplus Krishtalka 1976

†Talpavus nitidus Marsh 1872

†Talpavus sullivani Guthrie 1971

Fm. †Vastanidae Bajpai et al. 2005

G. †Vastania Bajpai et al. 2005

Invalid names: Creotarsinae Hay 1930 [invalid subgroup]

G. †Ghamidtherium Sánchez-Villagra et al. 2007

†Ghamidtherium dimaiensis Sánchez-Villagra et al. 2007

Subor. Soricomorpha Gregory 1910

Fm. †Changlelestidae Tong and Wang 1993

G. †Changlelestes Tong and Wang 1993

G. †Ernosorex Wang and Li 1990

Subfm. †Tupaiodontinae Butler 1988

Fm. †Geolabididae McKenna 1960

G. †Batodonoides Novacek 1976

G. †Centetodon Marsh 1872

Invalid names: Embassis Cope 1873 [synonym], Geolabis Cope 1884 [synonym], Hypacodon McKenna 1960 [synonym], Protictops Peterson 1934 [synonym]

G. †Gobigeolabis Lopatin 2004

G. †Marsholestes McKenna 1992

Invalid names: Myolestes Matthew 1909 [synonym]

G. †Stilpnodon Simpson 1935

Invalid names: Centetodontinae Trouessart 1879 [synonym], Metacodontidae Butler 1948 [synonym], Protictops alticuspidens Peterson 1934 [nomen dubium]

G. †Nycticonodon Quinet 1964

Fm. †Plesiosoricidae Winge 1917

G. †Butselia Quinet and Misonne 1965

G. †Ordolestes Lopatin 2006

G. †Pakilestes Russell and Gingerich 1981

G. †Plesiosorex Pomel 1848

Invalid names: Meterix Hall 1929 [synonym]

G. †Pseudoneurogymnurus Gureev 1979

G. †Siamosorex Peigné et al. 2009

Invalid names: Butselidae Quinet and Misonne 1965 [synonym], Butseliinae Quinet and Misonne 1965 [empty], Plesiosoricinae Winge 1917 [empty]

Fm. Soricidae Fischer 1814 [shrew]

G. †Alloblarinella Storch 1995

G. †Alluvisorex Hutchison 1966

G. Amphisorex Duvernoy 1835

G. †Asoriculus Kretzoi 1959

G. †Beremendia Kormos 1930

G. †Cokia Storch 1995

G. †Cretasorex Nessov and Gureyev 1981

Subfm. Crocidosoricinae Reumer 1987

Subfm. Crocidurinae Milne Edwards 1874 [white-toothed shrew]

Subfm. †Heterosoricinae Viret and Zapfe 1951

Subfm. †Limnoecinae Repenning 1967

G. †Lunanosorex Jin and Kawamura 1996

G. †Nesiotites Bate 1945

G. †Paenepetenyia Storch 1995

G. †Parydrosorex Wilson 1968

G. †Petenyiella Kretzoi 1956

Invalid names: Allopachyura Kormos 1934 [synonym]

G. †Podihik Deraniyagala 1958

G. †Siwalikosorex Sahni and Khare 1976

Subfm. Soricinae Fischer von Waldheim 1817 [red-toothed shrew]

Subfm. †Soricolestinae Lopatin 2002

G. †Zelceina Sulimski 1962

Invalid names: Allosoricinae Fejfar 1966 [empty]

Superfm. Soricoidea Gill 1872

Fm. †Apternodontidae Matthew 1910

Fm. †Micropternodontidae

Fm. †Nyctitheriidae Simpson 1928

Fm. †Oligoryctidae Asher et al. 2002

Fm. †Parapternodontidae Asher et al. 2002

Infraor. †Soricota Kalandadze and Rautian 1992

Superfm. †Nesophontoidea Anthony 1916

Superfm. Talpoidea Fischer de Waldheim 1817

Invalid names: Micropternodontoidea Stirton and Rensberger 1964 [empty], Solenodontoidea Gill 1972 [empty]

Invalid names: Lipotyphla Haeckel 1866 [synonym]

Unr. Scrotifera Waddell et al. 1999

Or. †Acreodi Matthew 1909

G. †Andrewsarchus Osborn 1924

†Andrewsarchus crassum Ding et al. 1977

†Andrewsarchus mongoliensis Osborn 1924

Unr. †Mesonychia [mesonychian]

Fm. †Hapalodectidae Szalay and Gould 1966 [condylarth]

Fm. †Mesonychidae Cope 1875 [condylarth]

G. †Simidectes Stock 1933

Invalid names: Pleurocyon Peterson 1919 [synonym]

G. †Wyolestes Gingerich 1981

Invalid names: Andrewsarchidae Szalay and Gould 1966 [empty], Wyolestidae [empty]

G. †Mondegodon Tabuce et al. 2011

†Mondegodon eutrigonus Tabuce et al. 2011

Fm. †Triisodontidae Scott 1892

G. †Eoconodon Matthew and Granger 1921

G. †Goniacodon Scott 1892

G. †Stelocyon Gingerich 1978

G. †Triisodon Cope 1881

Invalid names: Sarcothraustes Cope 1882 [synonym]

Invalid names: Oxyclaenidae Scott 1892 [invalid subgroup]

Unr. †Bulbulodentata Archibald 1998

Fm. †Periptychidae Cope 1882 [condylarth]

G. †Alticonus Eberle and Lillegraven 1998

G. †Ampliconus Eberle and Lillegraven 1998

Subfm. †Anisonchinae Osborne and Earle 1895

G. †Anisonchus Cope 1881

Invalid names: Zetodon Cope 1883 [synonym]

G. †Auraria Middleton and Dewar 2004

G. †Beornus Atteberry and Eberle 2021

G. †Conacodon Matthew 1897

G. †Escatepos Reynolds 1936

G. †Goleroconus McKenna and Honey 2008

G. †Hemithlaeus Cope 1882

G. †Miniconus Atteberry and Eberle 2021

G. †Mithrandir Cifelli 1983

Invalid names: Gillisonchus Rigby 1981 [synonym]

G. †Oxyacodon Osborn and Earle 1895

Invalid names: Fimbrethil Van Valen 1978 [synonym]

G. †Paleoungulatum Kelly 2014

Subfm. †Periptychinae Osborn and Earle 1895

G. †Tinuviel Van Valen 1978

Invalid names: Conacodontinae Archibald et al. 1983 [empty]

Or. Chiroptera Blumenbach 1779 [bat]

Fm. †Aegyptonycteridae Simmons et al. 2016

G. †Aegyptonycteris Simmons et al. 2016

G. †Archaeopteropus Meschinelli 1903

†Archaeopteropus transiens Meschinelli 1903

G. †Australonycteris Hand et al. 1994

†Australonycteris clarkae Hand et al. 1994

Subor. Frugivora Gill 1872

Invalid names: Pteropidae Gray 1821 [empty]

Fm. †Icaronycteridae Jepsen 1966

G. †Icaronycteris Jepsen 1966

Unr. Microchiropteramorpha Simmons and Geisler 1998

Fm. †Archaeonycteridae Revilliod 1917

Fm. †Hassianycterididae Habersetzer and Storch 1987

G. †Jaegeria Bajpai et al. 2005

Subor. Microchiroptera Dobson 1875

Fm. †Palaeochiropterygidae Revilliod 1917

G. †Mixopteryx Weithofer 1887

†Mixopteryx dubia Weithofer 1887

†Mixopteryx weithoferi Maître 2008

Fm. †Nycterididae Vanderhoevan 1855

G. †Chibanycteris Sigé et al. 1994

Fm. †Onychonycteridae Simmons et al. 2008

G. †Onychonycteris Simmons et al. 2008

Fm. †Philisidae Sigé 1985

G. †Dizzya Sige 1991

G. †Philisis Sigé 1985

G. †Scotophilisis HoráČek, et al. 2006

G. †Vampyravus Schlosser 1910

Invalid names: Provampyrus Schlosser 1911 [objective synonym]

G. †Witwatia Gunnell et al. 2008

G. †Samonycteris Revilliod 1917

†Samonycteris majori Revilliod 1917

Fm. †Tanzanycterididae Gunnell et al. 2003

G. †Tanzanycteris Gunnell et al. 2003

G. †Vesperugo Cope 1873

†Vesperugo anemophilus Cope 1880

Subor. Yangochiroptera Koopman 1984

Superfm. Emballonuroidea

Fm. Myzopodidae Milne-Edwards and Grandidier 1878 [sucker-footed bat]

Superfm. Noctilionoidea Gray 1821

Superfm. Vespertilionoidea

Subor. Yinpterochiroptera Koopman 1984

Fm. Pteropodidae Gray 1821 [fruit bat]

Superfm. Rhinolophoidea

Invalid names: Animalivora Gill 1872 [empty], Eochiroptera Van Valen 1979 [empty], Fructivorae Gray 1821 [empty], Insectivorae Gray 1821 [empty], Megachiroptera Dobson 1875 [empty]

Unr. Euungulata Waddell et al. 2001

Unr. Artiodactylamorpha Spaulding et al. 2009

Unr. Artiodactyla Owen 1848 [even-toed ungulate]

Invalid names: Cetartiodactyla Montgelard et al. 1997 [synonym]

Unr. †Panperissodactyla Welker et al. 2015

Or. †Panameriungulata de Muizon and Cifelli 2000

Superor. Perissodactylamorpha Rose et al. 2020

Unr. Phenacodonta McKenna 1975

Invalid names: Eparctocyona McKenna 1975 [synonym], Ungulata Linnaeus 1766 [synonym]

Unr. Ferae Linnaeus 1758

Unr. Carnivoramorpha Wyss and Flynn 1993

Unr. †Carnivoraformes Flynn et al.

G. †Dawsonicyon Spaulding et al. 2010

G. †Oodectes Wortman 1901

Invalid names: Paeneprolimnocyon Guthrie 1967 [synonym]

G. †Tapocyon Stock 1934

Fm. †Viverravidae Wortman and Matthew 1899

Invalid names: Didymictidae Flynn and Galiano 1982 [synonym]

Or. †Creodonta Cope 1875 [creodont]

G. †Altacreodus Fox 2015

Or. †Hyaenodonta Van Valen 1967

Unr. †Oxyaenodonta Van Valen 1971

Invalid names: Ambloctonidae Cope 1877 [empty], Amblyctonidae Cope 1877 [empty], Hyaenodontida Leidy 1869 [empty], Limnocyonidae [empty], Uintacyonidae Hay 1902 [empty]

Unr. †Pholidotamorpha Gaudin et al. 2009

Subor. †Palaeanodonta Matthew 1918

Or. Pholidota Weber 1904 [pangolin]

Invalid names: Squamata Huxley 1872 [synonym]

Invalid names: Ostentoria [synonym]

Unr. Fereuungulata Waddell et al. 1999

Invalid names: Zooamata Waddell et al. 1999 [empty]

Invalid names: Variamana [synonym]

Invalid names: Laurasiaplacentalia [synonym]

Invalid names: Boreoplacentalia [synonym], Boreotheria [synonym]

Diagnosis

No diagnoses are available

Measurements

No measurements are available

Composition:

phosphaticsubp

Maximum body mass:

1000 kgg

Minimum body mass:

3 gg

Environment:

terrestrialsubc

Locomotion:

actively mobilec

Life habit:

scansorialsubc

Diet:

insectivoresubc

Reproduction:

viviparoussubc

Created:

2005-09-06 20:43:42

Modified:

2005-09-06 22:44:32

Source: g = genus, subc = subclass, c = class, subp = subphylum

References: Lillegraven 1979, Czaplewski 1997, Carroll 1988, Hendy et al. 2009, Legendre 1988, Martin 1990, Ji et al. 2002

Age range

Maximum range based only on fossils: base of the Middle Turonian to the top of the Holocene or 93.50000 to 0.00000 Ma
Minimum age of oldest fossil (stem group age): 89.3 Ma

Collections (18208 total)

Time interval	Ma	Country or state	Original ID and collection number
Oldest occurrences
Middle Turonian - Late/Upper Turonian	93.5 - 89.3	Uzbekistan (Navoi)	Cretasorex arkhangelskyi (58179) Periptychidae indet. (58168)
Judithian	83.5 - 70.6	Canada (Alberta)	Nyctitheriidae indet. (14451)
Maastrichtian	72.1 - 66.0	India (Madhya Pradesh)	Kharmerungulatum vanvaleni (75493)
Maastrichtian	72.1 - 66.0	Portugal (Coimbra)	Leptacodon sp. (49276)
Lancian	70.6 - 66.0	Canada (Alberta)	Cimolestes magnus (14630 14633 14635 14637)
Lancian	70.6 - 66.0	USA (Montana)	Cimolestes magnus (14516 14532)
Lancian	70.6 - 66.0	Canada (Saskatchewan)	Cimolestes magnus (14558)
Lancian	70.6 - 66.0	USA (Wyoming)	Cimolestes magnus (14519 14598) Protungulatum sp. (14514)
Late/Upper Maastrichtian	70.6 - 66.0	USA (North Dakota)	Cimolestes magnus (45144)
Late/Upper Maastrichtian	70.6 - 66.0	USA (Montana)	Baioconodon nordicus, Paleoungulatum hooleyi (183109) Protungulatum coombsi (110464) Protungulatum gorgun (14674)
Pu1	66.0 - 63.3	USA (Montana)	Protungulatum donnae, Mimatuta minuial, Purgatorius janisae, Purgatorius mckeeveri (14762)
Pu3	66.0 - 63.3	USA (Montana)	Eoconodon hutchisoni (143205) Eoconodon hutchisoni, Eoconodon nidhoggi, Purgatorius mckeeveri (218647) Oxyacodon ferronensis, Eoconodon hutchisoni, Eoconodon nidhoggi, Purgatorius mckeeveri, Purgatorius unio (14738)
Puercan	66.0 - 63.3	Canada (Alberta)	Pandemonium hibernalis (174625)
Puercan	66.0 - 63.3	USA (Montana)	Baioconodon nordicus (14693) Carcinodon sp., Baioconodon nordicus, Oxyacodon ferronensis, Tinuviel eurydice (14744) Carcinodon sp., Loxolophus schizophrenus, Loxolophus sp., Oxyclaenus sp., Baioconodon nordicus, Oxyacodon ferronensis, Oxyacodon apiculatus, Tinuviel eurydice, Eoconodon nidhoggi (14745) Eoconodon hutchisoni (140079 218651 218655) Eoconodon nidhoggi (218650) Mimatuta minuial (14701) Mimatuta minuial, Ragnarok engdahli (14694) Mimatuta morgoth (14675 108198 108199) Mimatuta sp. (14801 14802) Mimatuta sp., Ragnarok nordicum (14668 14669 14740 14789) Oxyacodon ferronensis (218654) Oxyacodon ferronensis, Eoconodon hutchisoni (14799) Oxyacodon ferronensis, Eoconodon hutchisoni, Eoconodon nidhoggi (218648) Oxyacodon ferronensis, Eoconodon nidhoggi (218649) Oxyclaenus pugnax, Anisonchus oligistus, Tinuviel eurydice, Leptacodon proserpinae, Purgatorius unio, Plesiadapidae indet., Protungulatum sp., Oxprimus albertensis, Tricentes sp., Eoconodon sp., Oxyacodon josephi, Eoconodon sp. (14766) Oxyprimus erikseni (14696 14702) Oxyprimus erikseni, Mimatuta sp. (14767) Oxyprimus erikseni, Mimatuta sp., Ragnarok nordicum (14773) Oxyprimus erikseni, Ragnarok engdahli, Mimatuta morgoth (14742) Oxyprimus erikseni, Ragnarok nordicum (14660) Oxyprimus sp. (14665 14673) Protungulatum donnae (14663 14667 14753) Protungulatum donnae, Mimatuta morgoth, Mimatuta sp. (14771) Protungulatum donnae, Mimatuta sp., Periptychidae indet., Mimatuta sp. (14759) Protungulatum donnae, Oxyprimus erikseni, Mimatuta minuial, Mimatuta morgoth, Mimatuta sp., Ragnarok nordicum (14777) Protungulatum donnae, Oxyprimus erikseni, Mimatuta minuial, Mimatuta sp., Purgatorius coracis (193844) Protungulatum donnae, Oxyprimus erikseni, Mimatuta minuial, Ragnarok nordicum (14786) Protungulatum donnae, Oxyprimus erikseni, Mimatuta morgoth (14671) Protungulatum donnae, Oxyprimus erikseni, Mimatuta morgoth, Mimatuta sp., Ragnarok nordicum (14751) Protungulatum donnae, Oxyprimus erikseni, Mimatuta morgoth, Ragnarok engdahli (14798) Protungulatum donnae, Oxyprimus erikseni, Mimatuta morgoth, Ragnarok nordicum (14749) Protungulatum donnae, Ragnarok nordicum (14779) Protungulatum gorgun (14755 14757) Protungulatum gorgun, Mimatuta morgoth, Mimatuta sp., Ragnarok nordicum (14763) Protungulatum gorgun, Mimatuta morgoth, Ragnarok nordicum (14782) Protungulatum gorgun, Oxyprimus erikseni (14698) Protungulatum gorgun, Oxyprimus erikseni, Mimatuta sp. (14776) Protungulatum gorgun, Protungulatum donnae, Mimatuta morgoth, Mimatuta sp., Ragnarok nordicum (14787) Protungulatum gorgun, Protungulatum donnae, Oxyprimus erikseni, Mimatuta minuial, Mimatuta morgoth, Mimatuta sp., Ragnarok nordicum (14788) Protungulatum gorgun, Protungulatum donnae, Oxyprimus erikseni, Mimatuta morgoth, Ragnarok nordicum (14699 14754) Protungulatum gorgun, Protungulatum donnae, Oxyprimus erikseni, Mimatuta sp. (14780) Protungulatum gorgun, Protungulatum donnae, Oxyprimus erikseni, Mimatuta sp., Ragnarok nordicum (14784 14785) Protungulatum gorgun, Protungulatum donnae, Oxyprimus erikseni, Oxyprimus sp., Baioconodon engdahli, Mimatuta minuial, Mimatuta morgoth, Baioconodon harbichti (14705) Protungulatum gorgun, Protungulatum donnae, Protungulatum sp., Baioconodon sp., Mimatuta morgoth (14672) Protungulatum gorgun, Protungulatum donnae, Protungulatum sp., Oxyprimus erikseni, Ragnarok harbichti, Mimatuta morgoth, Purgatorius ceratops (14741) Protungulatum gorgun, Ragnarok nordicum (14676) Protungulatum mckeeveri, Ragnarok nordicum, Mimatuta minuial, Periptychidae indet. (14761) Protungulatum sp., Carcinodon aquilonius, Loxolophus sp., Oxyclaenus simplex, Oxyprimus sp., Baioconodon denverensis, Oxyacodon sp., Anisonchus sp., Periptychidae indet., Plesiadapidae indet., Ursolestes perpetior, Purgatorius sp., Protogonodon pentacus (14809) Protungulatum sp., Mimatuta sp. (14770) Ragnarok harbichti (14743) Ragnarok nordicum (14697 14700 14752 14758 14778)
Puercan	66.0 - 63.3	USA (Colorado)	Baioconodon denverensis, Tinuviel gazini (14774) Carsioptychus coarctatus (176748 205865 205867 205895 205900 205901 205903 205907 205908 205910 205914 205928 209872 209892 209904 209921 209924 209943 209946 209957 209964 209968 209976 210033) Carsioptychus coartatus (206072 206084) Carsioptychus sp. (206071) Conacodon delphae, Periptychus coarctatus (14678) Conacodon entoconus (205919) Conacodon harbourae (88207) Ectoconus ditrigonus (14684 14687 14688 205863 205864 205868 205871 205881 205886 209900 209983 209997) Eoconodon coryphaeus (205932 205934) Loxolophus hyattianus, Anisonchus athelas, Periptychus coarctatus, Ectoconus ditrigonus (14792) Loxolophus hyattianus, Oxyclaenus cuspidatus, Periptychus coarctatus, Conacodon entoconus (14683) Loxolophus hyattianus, Periptychus coarctatus (14677) Loxolophus sp. (209874 209879 209890 209965) Maiorana sp., Protungulatum donnae, Oxyprimus sp., Baioconodon nordicus, Ampliconus browni (176863) Oxyclaenus cuspidatus, Conacodon delphae (14689) Oxyclaenus subbituminus, Oxyprimus galadrielae, Baioconodon denverensis, Arctocyonidae indet., Auraria urbana, Oxyacodon archibaldi, Conacodon matthewi, Ampliconus browni, Baioconodon cannoni, Tinuviel gazini (14658) Periptychidae indet. (205873) Periptychus coarctatus (14659 14680 14681 14682 14685 14686 14746 14750 14793) Periptychus coarctatus, Ectoconus ditrigonus (14679)
Puercan	66.0 - 63.3	USA (Utah)	Anisonchus oligistus, Ectoconus maiusculus, Periptychidae indet. (224186) Ectoconus symbolus, Ectoconus sp., Periptychus hamaxitus (14748) Loxolophus pentacus, Oxyacodon ferronensis, Anisonchus oligistus, Anisonchus athelas, Ectoconus ditrigonus, Ectoconus symbolus, Periptychus hamaxitus (14736) Loxolophus pentacus, Oxyclaenus pugnax, Oxyacodon ferronensis, Oxyacodon apiculatus, Oxyacodon sp., Periptychus coarctatus, Ectoconus symbolus (14670) Loxolophus sp., Anisonchus athelas, Ectoconus ditrigonus, Conacodon utahensis (14739) Loxolophus spiekeri, Oxyclaenus pugnax, Oxyacodon apiculatus, Anisonchus onostus, Anisonchus athelas, Periptychus coarctatus, Ectoconus ditrigonus, Ectoconus symbolus, Haploconus elachistus, Haploconus sp. (14692) Oxyclaenus simplex, Eoconodon sp. (14791) Protogonodon sp., Loxolophus sp., Oxyclaenus sp., Oxyacodon marshater, Anisonchus oligistus, Anisonchus athelas, Carsioptychus hamaxitus, Ectoconus symbolus, Goniacodon hiawathae, Mixodectidae indet., Periptychus hamaxitus, Conacodon utahensis, Haploconus sp. (14790)
Puercan	66.0 - 63.3	USA (North Dakota)	Protungulatum sp., Oxyacodon priscilla (14765)
Puercan	66.0 - 63.3	USA (New Mexico)	Arctocyon ferox (74549 125183) Hemithlaeus sp., Conacodon sp., Ectoconus sp., Anisonchus sp., Carsioptychus sp. (14661) Loxolophus hyattianus, Hemithlaeus kowalevskianus, Periptychus coarctatus, Ectoconus ditrigonus, Conacodon entoconus, Eoconodon gaudrianus, Erinaceomorpha indet., Gillisonchus gillianus (14794) Loxolophus kimbetovius, Loxolophus hyattianus, Oxyclaenus simplex, Oxyclaenus cuspidatus, Hemithlaeus kowalevskianus, Conacodon kohlbergeri, Periptychus coarctatus, Ectoconus ditrigonus, Conacodon entoconus, Dissacus navajovius, Eoconodon coryphaeus, Eoconodon gaudrianus, Gillisonchus gillianus (14703) Loxolophus kimbetovius, Oxyclaenus simplex, Oxyclaenus cuspidatus, Oxyacodon apiculatus, Oxyacodon priscilla, Escatepos campi, Hemithlaeus kowalevskianus, Conacodon cophater, Conacodon kohlbergeri, Periptychus coarctatus, Ectoconus ditrigonus, Conacodon entoconus, Eoconodon ginibitohia, Eoconodon coryphaeus, Eoconodon gaudrianus, Eoconodon sp., Erinaceomorpha indet., Mimotricentes mirielae, Fimbrethil ambaronae, Gillisonchus gillianus (14760) Loxolophus priscus, Loxolophus pentacus, Loxolophus hyattianus, Oxyclaenus simplex, Oxyclaenus cuspidatus, Oxyacodon apiculatus, Eoconodon coryphaeus, Chriacus antiquus, Protogonodon grangeri, Conacodon sp., Ectoconus sp., Gillisonchus gillianus, Plagioptychus matthewi (14662) Loxolophus priscus, Loxolophus pentacus, Loxolophus hyattianus, Periptychus coarctatus, Gillisonchus gillianus (14795)
Puercan	66.0 - 63.3	USA (Wyoming)	Arctocyonidae indet. (14715 14735) Arctocyonidae indet., Conacodon cophater, Periptychus coarctatus, Ectoconus ditrigonus (14716) Arctocyonidae indet., Triisodontinae indet. (14805) Baioconodon sp. (14712) Conacodon cophater (14727) Ectoconus ditrigonus (14717) Loxolophus faulkneri, Loxolophus sp., Mithrandir gillianus, Ampliconus browni (14713) Loxolophus faulkneri, Mithrandir gillianus (14707) Loxolophus faulkneri, Oxyacodon priscilla, Mithrandir gillianus, Periptychus coarctatus, Periptychidae indet. (14706) Loxolophus hyattianus, Arctocyonidae indet., Oxyacodon agapetillus, Mithrandir gillianus, Eoconodon sp. (14731) Loxolophus hyattianus, Mithrandir gillianus (14728) Loxolophus priscus, Loxolophus hyattianus, Baioconodon middletoni, Oxyacodon priscilla, Oxyacodon agapetillus, Mithrandir gillianus, Alticonus gazini, Ectoconus cavigellii, Ampliconus antoni, Periptychidae indet. (14797) Maiorana ferrisensis, Protungulatum donnae, Mimatuta sp. (14711) Maiorana noctiluca, Oxyprimus galadrielae, Oxyprimus putorius, Mimatuta minuial, Loxolophus nordicus, Eoconodon sp., Oxyacodon josephi (14808) Maiorana noctiluca, Protungulatum donnae, Oxyprimus sp., Sigynorum magnadivisus, Mimatuta sp., Miniconus jeanninae, Beornus honeyi, Conacodon hettingeri, Conacodon harbourae, Ampliconus antoni (174233) Mimatuta makpialutae, Ragnarok wovokae (14807) Mimatuta sp. (14725) Mithrandir gillianus (14714 14718 14724 14726) Oxyacodon apiculatus (14734) Oxyclaenus cuspidatus, Arctocyonidae indet., Mithrandir gillianus, Conacodon cophater (14723) Oxyclaenus cuspidatus, Oxyclaenus sp., Baioconodon denverensis (14729) Oxyclaenus simplex (14803) Oxyprimus galadrielae, Arctocyonidae indet. (14804) Oxyprimus sp., Eoconodon sp. (14710) Periptychus coarctatus (14721 14733) Periptychus coarctatus, Periptychidae indet. (14719) Protungulatum donnae (14708 14709) Protungulatum donnae, Loxolophus priscus, Oxyacodon priscilla, Periptychus coarctatus (14720) Protungulatum gorgun, Protungulatum donnae, Arctocyonidae indet., Oxyacodon agapetillus, Mithrandir gillianus, Conacodon delphae, Ectoconus sp., Periptychidae indet., Conacodon harbourae (14722) Protungulatum sloani, Protungulatum donnae, Oxyprimus galadrielae, Oxyprimus sp., Mimatuta sp. (14796)
Puercan	66.0 - 63.3	Canada (Saskatchewan)	Loxolophus schizophrenus, Oxyclaenus sp., Baioconodon denverensis, Arctocyonidae indet. (14691) Protungulatum donnae, Oxyprimus erikseni, Mimatuta sp., Ragnarok sp. (14756) Protungulatum donnae, Periptychidae indet. (14737) Ravenictis krausei, Carnivora indet., Carcinodon aquilonius, Loxolophus schizophrenus, Oxyclaenus corax, Baioconodon denverensis, Oxyacodon agapetillus, Anisonchus oligistus, Eoconodon sp., Purgatorius sp. (14768)