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Carusia intermedia

Reptilia - Squamata

Taxonomy
Carolina intermedia was named by Borsuk-Bialynicka (1985). Its type specimen is ZPAL Mg-R-III/34, a partial skull (Almost complete skull with mandibles. Braincase and palate damages, supratemporal arcade missing), and it is a 3D body fossil. Its type locality is Khermin Tsav (DUPLICATE), which is in a Campanian terrestrial horizon in the Barun Goyot Formation of Mongolia. It is the type species of Carusia.

It was recombined as Carusia intermedia by Borsuk-Bialynicka (1987), Gao and Hou (1996), Gao and Norell (1998) and Gao and Norell (2000).

Synonymy list
YearName and author
1985Carolina intermedia Borsuk-Bialynicka pp. 154-160 figs. 1-6, pls 1-4
1987Carusia intermedia Borsuk-Bialynicka
1996Carusia intermedia Gao and Hou p. 587
1998Carusia intermedia Gao and Norell p. 3
2000Carusia intermedia Gao and Norell p. 90

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RankNameAuthor
kingdomAnimalia()
Triploblastica
Nephrozoa
Deuterostomia
phylumChordataHaeckel 1847
subphylumVertebrata
superclassGnathostomata
Osteichthyes()
Sarcopterygii
subclassDipnotetrapodomorpha(Nelson 2006)
subclassTetrapodomorpha()
Tetrapoda()
Reptiliomorpha
Anthracosauria
Batrachosauria()
Cotylosauria()
RankNameAuthor
Amniota
Sauropsida
classReptilia
subclassEureptilia()
Romeriida
Diapsida()
Eosuchia()
Neodiapsida
SauriaGauthier 1984
Lepidosauromorpha(Benton 1983)
superorderLepidosauria()
orderSquamata
superfamilyScincoideaOppel 1811
genusCarusia
speciesintermedia()

If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.

Diagnosis
ReferenceDiagnosis
K. -Q. Gao and M. A. Norell 1998Sharing with extant xenosaurids the following derived characters: strong interorbital constriction of fused frontals; jugal-squamosal contact; dermal vermiculate rugosities on skull roof elements; postorbital branch of jugal ornamented. Distinguished from all other closely related taxa including Xenosaurus and Shinisaurus by the following autapomorphies: lacrimal absent, with anteroventral process of jugal contacting prefrontal; maxilla contributes to formation of ventral border of orbit; caudoventral process of jugal ventrally directed; fenestra exochoanalis distinctively short and elliptic; presence of midline contact of palatines anteriorly; palatine rectangular and strongly widened, suppressing suborbital fenestra into narrow and elongate opening; presence of vertical crest anteriorly on quadrate; condyle-fossa articulation for both squamosal and supratemporal with cephalic condyle of quadrate; descending process of parietal subrectangular, elongated, and laterally compressed; epipterygoid slanted strongly posterodorsally; marginal process of supraoccipital prominent and well-ossified; presence of strong adductor crest dorsolaterally on surangular; marginal teeth extremely fine, numerous, and having comblike arrangement; strong posterior extension of neural spone of axis over entire third cervical.