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Tombaatar sabuli

Mammalia - Multituberculata - Djadochtatheriidae

Taxonomy
Tombaatar sabuli was named by Rougier et al. (1997). Its type specimen is PSS-MAE 122, a partial skull, and it is a 3D body fossil. Its type locality is Ukhaa Tolgod (AMNH), which is in a Campanian eolian sandstone/mudstone in the Djadokhta Formation of Mongolia. It is the type species of Tombaatar.

Synonymy list
YearName and author
1997Tombaatar sabuli Rougier et al. p. 4 figs. 1-5
2004Tombaatar sabuli Kielan-Jaworowska et al. pp. 254, 324

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RankNameAuthor
kingdomAnimalia()
Triploblastica
Nephrozoa
Deuterostomia
phylumChordataHaeckel 1847
subphylumVertebrata
superclassGnathostomata
Osteichthyes()
Sarcopterygii
subclassDipnotetrapodomorpha(Nelson 2006)
subclassTetrapodomorpha()
Tetrapoda()
Reptiliomorpha
Anthracosauria
Batrachosauria()
Cotylosauria()
Amniota
RankNameAuthor
Synapsida()
Therapsida()
infraorderCynodontia()
Epicynodontia
infraorderEucynodontia
Probainognathia
Mammaliamorpha
Mammaliaformes
classMammalia
Allotheria()
orderMultituberculataCope 1884
suborderCimolodonta()
superfamilyDjadochtatherioidea
familyDjadochtatheriidae
genusTombaatar
speciessabuli

If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.

Diagnosis
ReferenceDiagnosis
G. W. Rougier et al. 1997Large size multituberculate, with palate length (measured from the posterior margin of the I2) of 30.4 mm and a M1-M2 length of 9.5 mm. Palatal torus is notably strong, more than in any other presently known multituberculate, with an elaborated, stout, crescent-shaped process that projects ventrally on either palatine (as in the weaker torus of Kryptobaatar). MI cusp formula (4:5:2) and size, differentiate Tombaatar from all Mongolian multituberculates. Skull of Tombaatar is similar in dimensions to those of Djadochtatherium matthewi (Simpson, 1925) and Catopsbaatar catopsaloides (Kielan-Jaworowska, 1974, 1994; Kielan-Jaworowska and Sloan, 1979), but differs from these taxa in having the 13 alveolus formed by both the premaxilla and maxilla; a slender, posteriorly tapering process of the premaxilla on the rostrum; a high skull with a deep orbit (flat rostrum, and palate wider and shorter in Djadochtatherium and Catopsbaatar); and a proportionately short and slender anterior process of the frontal on the skull roof (long and wide in Djadochtatherium, but not preserved in Catopsbaatar). P2 absent and rounded premaxillary-maxillary suture in rostrum of Tombaatar and Catopsbaatar (P2 present and suture not rounded in Djadochtatherium; suture rounded in Nemegtbaatar, Kryptobaatar, and Chulsanbaatar, but not in Sloanbaatar, Kamptobaatar, Ectypodus, or Lambdopsalis). Tombaatar and Djadochtatherium are distinguished from Catopsbaatar in having large P1 and P3 (?plesiomorphic) and MI with only two cusps on the lingual row which form a lobe in the posterior half of the molar of Tombaatar. Four lingual cusps cover the posterior two-thirds of the labial margin of the MI in Catopsbaatar. Some additional characters are present in other multituberculates but are rendered autapomorphies of Tombaatar by their distribution on the most parsimonious trees. These are: base of zygomatic arch dorsal or anterior to P3/P4 embrasure, incisive foramina of medium size and elongated, and posterior margin of palate medial to middle of M2.