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Aegyptocetus tarfa

Mammalia - Cetacea - Protocetidae

Taxonomy
Aegyptocetus tarfa was named by Bianucci and Gingerich (2011). Its type specimen is MSNTUP I-15459, a partial skeleton (skull and partial skeleton), and it is a 3D body fossil. Its type locality is Khashm el-Raqaba, which is in a Bartonian carbonate limestone in the Gebel Hof Formation of Egypt. It is the type species of Aegyptocetus.

Synonymy list
YearName and author
2011Aegyptocetus tarfa Bianucci and Gingerich p. 1175

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RankNameAuthor
kingdomAnimalia()
Bilateria
EubilateriaAx 1987
Deuterostomia
phylumChordataHaeckel 1874
subphylumVertebrata
superclassGnathostomata
Osteichthyes()
subclassSarcopterygii()
subclassDipnotetrapodomorpha(Nelson 2006)
subclassTetrapodomorpha()
Tetrapoda
Reptiliomorpha
Anthracosauria
subclassAmphibiosauriaKuhn 1967
Cotylosauria()
Amniota
subclassSynapsida
Therapsida()
infraorderCynodontia()
Mammaliamorpha
Mammaliaformes
RankNameAuthor
classMammalia
Theriamorpha(Rowe 1993)
Theriiformes()
Trechnotheria
Cladotheria
Zatheria
subclassTribosphenida()
subclassTheria
Eutheria()
Placentalia
Boreoeutheria
Laurasiatheria
Scrotifera
Euungulata
Artiodactylamorpha
Artiodactyla()
Whippomorpha
orderCetacea
familyProtocetidae
genusAegyptocetus
speciestarfa

If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.

Aegyptocetus tarfa Bianucci and Gingerich 2011
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Diagnosis
ReferenceDiagnosis
G. Bianucci and P. D. Gingerich 2011Aegyptocetus tarfa differs from all protocetids except possibly Babiacetus indicus in having a cranium with a downwardly deflected frontal shield and rostrum of the skull (clinorhynchy; see below). Aegyptocetus differs in having a dentary with the mandibular condyle positioned well below the level of the lower cheek teeth, an ascending ramus deflected dorsally and medially to wrap around the braincase, and lower molars with the principle cusps angled backward (features related to cranial clinorhynchy). The protocetid Babiacetus has a cranium with a downwardly deflected frontal and rostrum, suggesting clinorhynchy like that documented here for Aegyptocetus, but deformation during burial means that clinorhynchy in Babiacetus cannot be quantified reliably. Aegyptocetus resembles Babiacetus in having a fused mandibular symphysis, but differs in having transversely broader upper posterior premolars (P3–4) and molars (M1–3). Upper premolars lack the well-developed posterobasal cusp present on P3 and the large metacone present on P4 in Babiacetus. There is no distinct protocone cusp on upper posterior premolars or molars of either Aegyptocetus or Babiacetus, but Aegyptocetus retains a prominent protocone lobe on the crowns of P3–M3, supported by a separate root. This lobe is medial, or slightly posterior of medial, relative to the paracone on P3–M3 of Aegyptocetus, whereas it is medial, or slightly posterior of medial relative to the posterior edge of the crown in Babiacetus. Cheek tooth occlusion in Aegyptocetus is more complex, like that of primitive protocetids, whereas cheek tooth occlusion in Babiacetus is simpler, more sectorial, and more derived.