PBDB Taxon

Basic info	Taxonomic history	Classification	Included Taxa
Morphology	Ecology and taphonomy	External Literature Search	Age range and collections

Colymbiculus udovichenkoi

Reptilia - Gaviidae

Taxonomy

Colymbiculus udovichenkoi was named by Mayr and Zvonok (2011). Its type specimen is SMF Av 545, a limb element (left humerus), and it is a 3D body fossil. Its type locality is Ikovo, which is in a Lutetian marine sandstone in Ukraine. It is the type species of Colymbiculus.

Synonymy list

Year	Name and author
2011	Colymbiculus udovichenkoi Mayr and Zvonok p. 1354

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Rank	Name	Author
kingdom	Animalia	()
—	Bilateria
—	Eubilateria	Ax 1987
—	Deuterostomia
phylum	Chordata	Haeckel 1874
subphylum	Vertebrata
superclass	Gnathostomata
—	Osteichthyes	()
subclass	Sarcopterygii	()
subclass	Dipnotetrapodomorpha	(Nelson 2006)
subclass	Tetrapodomorpha	()
—	Tetrapoda
—	Reptiliomorpha
—	Anthracosauria
subclass	Amphibiosauria	Kuhn 1967
—	Cotylosauria	()
—	Amniota
—	Sauropsida
class	Reptilia
subclass	Eureptilia	()
—	Romeriida
—	Diapsida	()
—	Archosauromorpha	(Huene 1946)

Rank	Name	Author
—	Crocopoda
—	Archosauriformes	Gauthier 1986
—	Eucrocopoda
—	Archosauria	()
informal	Avemetatarsalia
—	Ornithodira
—	Dinosauromorpha
—	Dinosauriformes
—	Dinosauria	()
—	Theropoda	()
—	Neotheropoda
—	Averostra	Paul 2002
—	Tetanurae
—	Coelurosauria	()
—	Maniraptora
—	Paraves
class	Aves
subclass	Neornithes	Gadow 1893
—	Aequornithes
—	Gaviiformes	(Wetmore and Miller 1926)
family	Gaviidae	Allen 1897
genus	Colymbiculus
species	udovichenkoi

If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.

†Colymbiculus udovichenkoi Mayr and Zvonok 2011

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Diagnosis

Reference	Diagnosis
G. Mayr and E. A. Zvonok 2011	Small loon, the size of Colymboides minutus and ?C. metzleri. Distinguished from Colymboides in the following plesiomorphic features: (1) distal end of humerus with ventral portion more ventrally protruding (Text-fig. 5F, H); (2) ulna with smaller tuberculum carpale (Text-fig. 5M, P).