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Fulgoromorpha (planthopper)

Insecta - Hemiptera

Fulgoromorpha was named by Evans (1946). It is extant.

It was reranked as the unranked clade Fulgoromorpha by Grimaldi and Engel (2005); it was reranked as the suborder Fulgoromorpha by Szwedo (2018).

It was assigned to Auchenorrhyncha by Hong (1979); to Hemineoptera by Kukalová-Peck and Lawrence (2004); to Auchenorrhyncha by Grimaldi and Engel (2005); to Hemiptera by Szwedo (2007); and to Hemelytrata by Szwedo (2018).

Synonymy list
YearName and author
1878Cixius hesperidum Scudder pp. 772-773
1878Mnemosyne terrentula Scudder p. 773
1890Cixius hesperidum Scudder p. 287 figs. Pl 6 fig 19
1890Oliarites terrentula Scudder p. 293 figs. Pl 7 fig 17
1946Fulgoromorpha Evans
1979Fulgoromorpha Hong p. 302
2004Fulgoromorpha Kukalová-Peck and Lawrence
2005Fulgoromorpha Grimaldi and Engel p. 304
2007Fulgoromorpha Szwedo
2018Fulgoromorpha Szwedo p. 111

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EubilateriaAx 1987
phylumArthropodaLatreille 1829
superclassHexapodaLatreille 1825
NeopterygotaCrampton 1924
infraclassNeopteraMartynov 1923
HemelytrataFallen 1829
suborderFulgoromorpha(Evans 1946)
suborderFulgoromorpha(Evans 1946)

If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.

No diagnoses are available
No measurements are available
Composition: chitinc
Environment: terrestrialc
Locomotion: actively mobilep
Created: 2017-04-17 09:56:18
Modified: 2017-04-17 09:56:18
Source: c = class, p = phylum
References: Kiessling 2004, Bush and Bambach 2015

Age range

Maximum range based only on fossils: base of the Roadian to the top of the Holocene or 272.30000 to 0.00000 Ma
Minimum age of oldest fossil (stem group age): 268.8 Ma

Collections (195 total)

Time interval Ma Country or state Original ID and collection number
Roadian272.3 - 268.8Russian Federation (Kemerovo) Kaltanoscyta reticulata (122568)
Roadian272.3 - 268.8Russian Federation (Arkangelsk) Coleoscyta elytrata (116934) Coleoscyta rotundata (116933) Coleoscyta rotundata, Coleoscytodes elytrata (104993) Coleoscyta rotundata, Coleoscytodes venosa (111187) Coleoscytodes elytrata (116874)
Roadian272.3 - 268.8Russian Federation Coleoscyta occallata (96860)
Roadian272.3 - 268.8Russian Federation (Kazan) Coleoscyta martynovi, Coleoscyta kamensis (116877) Coleoscyta sp. (116803)
Wordian268.8 - 265.1Russian Federation (Kemerovo) Surijokocixius tomiensis, Reticulocicada brachyptera (122566)
Capitanian265.1 - 259.9Russian Federation (Orenburg) Scytocixius mendax (104994)
Wuchiapingian - Changhsingian259.9 - 252.17Russian Federation Scytocixius sp. (122565)
Changhsingian254.17 - 252.17Russian Federation (Vologda) Parapryg alogus (105046)
Induan252.17 - 251.2Russian Federation Boreocixius sibiricus, Boreocixius rotundatus (161389)
Bithynian247.2 - 242.0Germany (Bavaria) Heseneuma hammelburgensis (115969)
Longobardian242.0 - 235.0China (Shaanxi) Boreocixius sp. (183209)
Late/Upper Triassic237.0 - 201.3China (Shaanxi) Fulgoroidea indet. (155679)
Norian228.0 - 208.5Australia (Queensland) Tricrosbia minuta (123797)
Hettangian - Sinemurian201.3 - 190.8Kyrgyzstan (Tonskiy) Fulgoridiella raetica (105086)
Obtusum196.5 - 189.6United Kingdom (England) Mesocixiella fennahi (114638)
Pliensbachian190.8 - 182.7Kyrgyzstan Eofulgoridium kisylkiense, Eofulgoridium proximum, Fulgoropsis dubiosa (114701)
Early/Lower Toarcian183.0 - 182.0Germany (Mecklenburg-Vorpommern) Fulgoridium sp 1, Fulgoridium sp 2, Fulgoridium sp., Fulgoridulum egens (114993) Fulgoridulum egens (125507)
Early/Lower Toarcian183.0 - 182.0Germany Fulgoridium fractum (122986) Phryganidium balticum, Fulgoridulum egens, Fulgoridium pallidum, Fulgoridium venosum, Fulgoridium simplex, Fulgoridium latum, Protomyia dubia, Fulgoridium liadis, Fulgoridium lapideum, Cixiites liasinus, Fulgoridium spilographum, Fulgoridium oligoneurum, Fulgoridium regulare, Fulgoridium graphipterum, Fulgoridium inconspicuum, Fulgoridium quadrisignatum, Fulgoridium breviradiatum, Fulgoridium modestum, Fulgoridium anale, Fulgoridium marginepunctatum, Fulgoridium picturatum, Fulgoridium anomalum, Fulgoridium stigmaticum, Fulgoridium punctatum, Fulgoridium elegantulum, Fulgoridium curvipenne, Fulgoridium vicinum, Fulgoridium nebulosum, Fulgoridium clavatum, Fulgoridium ampliatum, Fulgoridium multipunctatum, Fulgoridium dilutum, Fulgoridium breve, Fulgoridium fenestratum, Fulgoridium grave, Fulgoridium obtusum, Fulgoridium nubeculum, Fulgoridium oligospilum, Fulgoridium polyneurum, Fulgoridium retractum, Fulgoridium parvispilum, Fulgoridium litorale, Fulgoridium trifurcatum, Fulgoridium remotum, Fulgoridium debile, Fulgoridium multivenosum, Fulgoridium pulchrum, Fulgoridium megapolitanum, Fulgoridium nodosum, Fulgoridium alatum, Fulgoridium mortuum, Fulgoridium defunctum, Fulgoridium plicatum, Fulgoridium exhumatum, Fulgoridium angulosum, Fulgoridium acutum, Fulgoridium rotundatum, Fulgoridium ancylla, Fulgoridium bifurcatum, Fulgoridium inaequale, Fulgoridium brachyptilum, Fulgoridium intercalatum, Fulgoridium rudimentum, Metafulgoridium spilotum, Metafulgoridium ampliatum, Metafulgoridium graptum, Margaroptilon germanicum, Fulgoridium balticum (123987)
Early/Lower Toarcian183.0 - 182.0United Kingdom (Gloucestershire) Neuroptera indet. (154307)
Early/Lower Toarcian183.0 - 182.0Germany (Niedersachsen) Fulgoridium aries, Fulgoridium praeacutum, Fulgoridium praefastigatum, Fulgoridium mancomarginatum, Fulgoridium exiguemaculatum, Fulgoridium cubitoramosum, Fulgoridium cuneiforme, Fulgoridium postredditum, Fulgoridium paulodilatatum, Fulgoridium semiperspicuum, Fulgoridium infuscatum, Fulgoridium spatulaeforme (107323) Fulgoridium balticum, Elasmocelidium boreale (117795) Fulgoridium beienrodense (107322) Fulgoridium concameratum, Fulgoridium decapitatum, Fulgoridium paenintegrum, Fulgoridium incisum, Fulgoridium verticillatum, Fulgoridium antennatum, Fulgoridium cubitofurcatum, Fulgoridium tenuimaculatum, Fulgoridium hildesheimense, Fulgoridium radioramosum, Fulgoridium hondelanum, Fulgoridium posidonicum, Fulgoridium gottingense, Fulgoridium raromaculatum, Fulgoridium basilaesum, Fulgoridium symmetricum, Fulgoridium fabri, Fulgoridium latius, Fulgoridium violatum, Fulgoridium praeobtusum, Fulgoridium incurvatum, Fulgoridium silvaticum, Margaroptilon paucisinuatum, Margaroptilon detruncatum, Margaroptilon formosum, Margaroptilon procerum, Elasmoscelidium venulosum, Tetragonidium paeneparallelum (105002) Fulgoridium hattorfense, Fulgoridium praetruncatum, Fulgoridium silvaticum, Margaroptilon cuneatum (107552) Fulgoridium incertecoloratum, Elasmocelidium boreale (107555) Fulgoridium planedorsatum, Fulgoridium bodei, Fulgoridium brunsvicense, Fulgoridium rotundatum (107556) Fulgoridium reduncum, Fulgoridium fallerslebense (108447) Fulgoridium sp., Fulgoridium obesum, Fulgoridium fronterotundum, Fulgoridium filiferum, Fulgoridium verticillatum, Margaroptilon formosum, Elasmoscelidium promotum, Elasmoscelidium rectemarginatum, Tetragonidium parallelogramma (107321) Fulgoridium spoliatum (108479)
Falciferum183.0 - 182.0United Kingdom (England) Fulgoridulum sp. (132646)
Toarcian182.7 - 174.1China (Xinjiang) Eofulgoridium tenellum (115209)
Toarcian182.7 - 174.1United Kingdom Margaroptilon woodwardi, Margaroptilon bulleni (132073)
Toarcian182.7 - 174.1Germany Fulgoridium germanicum (131767)
Toarcian - Aalenian182.7 - 170.3China (Hunan) Qiyangiricania cesta (114577)
Bajocian170.3 - 168.3China (Guangxi) Valvifulgoria tiantungensis, Valvifulgoria pingkuiensis (151540)
Callovian - Oxfordian166.1 - 157.3China (Nei Mongol) Fenghuangor imperator (113360)
Callovian - Oxfordian166.1 - 157.3Kazakhstan (Chimkent) Aulieezidium karatauense (152004) Elasmoscelidium rotundatum (114692) Fulgoridium sp. (112846)
Middle Berriasian145.0 - 140.2United Kingdom (England) Ricania fulgens, Cixius maculatus (104008)
Early/Lower Barremian130.0 - 125.45Lebanon Aafrita biladalshama (153333)
Early/Lower Barremian130.0 - 125.45Spain Chalicoridulum montsecensis (118913) Mimamontsecia cretacea (152693)
Late/Upper Barremian130.0 - 125.45Lebanon Mundopoides aptianus (123001) Neazonia imprinta (123000) Neazonia tripleta, Neazonia immatura, Aafrita biladalshama (122471)
Barremian129.4 - 125.0United Kingdom (England) Cixius petrinus (119664)
Barremian129.4 - 125.0Lebanon Aafrita biladalshama (124617)
Early/Lower Aptian125.45 - 122.46China (Liaoning) Lapicixius decorus (138436)
Early/Lower Aptian125.45 - 122.46Russian Federation (Chita) Mimarachnidae indet. (119499)
Aptian125.0 - 113.0Russian Federation (Buryatia) Mimarachne mikhailovi (125412 125413 144926) Mimarachne mikhailovi, Saltissus eskovi (122548 122549 122561) Saltissus eskovi (119498 122552 125792 144924 144925)
Aptian125.0 - 113.0Mongolia Mimarachnidae indet. (103657) Tsaganema oshanini (113355)
Aptian125.0 - 113.0Japan (Ishikawa) Fulgoridium matsuoi (152995)
Late/Upper Aptian122.46 - 112.03Brazil (Ceará) Cixiidae indet., Megaleurodes megocellata, Acixiites immodesta, Acixiites costalis, Protodelphax sp., Protodelphax miles, Protodelphax macroceps, Protodelphax rhinion, Protodelphax chamus, Ancorale sp., Ancorale flaccidum, Ancorale aschemon, Kinnarocixius sp., Kinnarocixius quassus, Lalax sp., Lalax mutabilis, Patulopes sp., Patulopes setosa, Patulopes myndoides, Carpopodus sp. A, Carpopodus sp. B, Carpopodus difficilis, Psestocixius fuscus, Psestocixius delphax, Vulcanoia sp., Vulcanoia apicalis, Vulcanoia acuceps (122352) Fennahia cretacea, Vulcanoia membranosa (125511)
Late/Upper Aptian122.46 - 112.03China (Beijing Shi) Cretocixius stigmatosus (156386) Yanducixius yihi, Yanducixius pardalinus (106965)
Albian113.0 - 100.5Jordan Fulgoroidea indet., Nablusitypus wafaai (124847)
Late/Upper Albian105.3 - 99.6Spain (Aragón) Iberofoveopsis miguelesi (91626)
Late/Upper Albian105.3 - 99.6France (Poitou-Charentes) Akmazeina santonorum (99114)
Early/Lower Cenomanian99.6 - 93.5France (Charente-Maritime) Fulgoroidea indet. (107076)
Early/Lower Cenomanian99.6 - 93.5Myanmar Burmissus raunoi, Dorytocus ornithorhynchus (190961) Dachibangus trimaculatus, Dachibangus formosus, Mimaplax ekrypsan (161726) Foveopsis heteroidea, Jaculistilus oligotrichus, Jubisentis hui, Furtivirete zhuoi (165561) Liburnia burmitina, Plecophlebus nebulosus, Cixitettiginae indet., Foveopsis fennahi (123020) Parwaina liuyei (201723) Yetkhata jiangershii (201722)
Middle Cenomanian99.6 - 93.5Argentina (Santa Cruz) Aonikenkissus zamunerae (157878)
Turonian93.9 - 89.8Kazakhstan Mimarachnidae indet. (127856)
Turonian93.9 - 89.8Russian Federation (Khabarovsk) Mimarachnidae indet. (138044)
Turonian93.9 - 89.8USA (New Jersey) Perforissus muiri (110895)
Late/Upper Santonian85.8 - 83.5Russian Federation Cretargus emeljanovi (35056) Netutela annunciator, Cixitettix yangi (122590)
Late/Upper Campanian83.5 - 70.6Myanmar Achilidae indet. (195455)
Paleocene66.0 - 56.0China (Xizang) Priscoflata subvexa, Gesaris gnapo (119466)
Paleocene66.0 - 56.0Argentina (Jujuy) Ormenis devinctus (110536)
Tiffanian61.7 - 56.8Canada (Alberta) Cixiidae indet. (119816)
Thanetian59.2 - 56.0France (Auvergne) Cintux menatensis, Mnasthaia arverniorum (123436) Hammapteryx eocenicus (113685)
Late/Upper Paleocene58.7 - 55.8Argentina Celinapterix bellissima (122881)
Clarkforkian56.8 - 55.8USA (North Dakota) Cotradechites lithinus (10975)
Ypresian56.0 - 47.8Australia (Queensland) Scolypopites bryani (125666)
Ypresian56.0 - 47.8Germany Fulgoroidea indet. (151359) Nogodinidae indet. (151361)
Ypresian56.0 - 47.8China (Yunnan) Weiwoboa meridiana (113892)
Ypresian56.0 - 47.8Denmark Cixiidae indet. (152152) Eoricania danica (123997) Hammapteryx paucistriata (156722) Lechaea primigenia (123999) Limfjordia breineri (118000) Limfjordia sp. (113025) Ormenis furcata (156747)
Ypresian56.0 - 47.8Canada (British Columbia) Cixiidae indet., Ricaniidae indet. (31677) Enchophora sp. (121046)
Wasatchian55.8 - 50.3USA (Wyoming) Fulgoroidea indet. (187759)
MP 755.8 - 48.6France (Oise) Abraracourcix curvivenatus, Stalisyne lutetiorum, Stalisyne veromanduiorum, Mnaomaia bellovaciorum, Ordralfabetix sirophatanis (123850)
Bridgerian50.3 - 46.2USA (Colorado) Aphana atava (121146) Delphax veterum (105004) Eofulgorella bradburyi (106481) Eoliarus quadristictus (140188) Hammapteryx lepidoides, Hammapteryx ceryniiformis, Lithopsis delicata, Protoliarus humatus, Dilaropsis ornatus (105005) Lithopsis simillima, Detyopsis scudderi, Detyopsis packardi, Scoparidea nebulosa, Thaumastocladius simplex, Lithopsis dubiosa, Hammapteryx tripunctata (114681) Protoliarus amabilis (151573)
Bridgerian50.3 - 46.2USA (Wyoming) Aphana rotundipennis, Lystra richardsoni, Lithopsis fimbriata (121322) Fulgora granulosa, Aphana rotundipennis, Lystra richardsoni, Cixius hesperidum, Mnemosyne terrentula, Lithopsis fimbriata (121306) Lystra richardsoni, Lithopsis elongata, Ficarasites stigmaticum (122092) Lystra richardsoni, Lithopsis fimbriata, Lystra leei, Fulgora populata, Oliarus lutensis, Diaplegma obdormitum, Hammapteryx reticulata (122090)
MP 1148.6 - 40.4Germany (Hessen) Fulgoroidea indet., Baninus thuringiorum, Amalaberga ostrogothiorum (120036) Wedelphus dichopteroides (120798)
MP 1348.6 - 40.4Germany (Eifel) Fulgoridae indet. (135640) Lycorma sp., Ricaniidae indet. 1, Ricaniidae indet. 2 (138912) Ricaniidae indet. 1 (135642 135644)
Lutetian47.8 - 41.3United Kingdom (England) Hammapteryx anglica (127644)
Lutetian47.8 - 41.3Germany (Sachsen-Anhalt) Eoricania reticulata, Eobladina antiqua, Giselia multifurcata, Giselia scalaris, Issites glaber, Eoperlites paradoxus (114060) Lithopsis major (155693)
Uintan46.2 - 40.4USA (Colorado) Lithopsis fimbriata, Emiliana alexandri (138675)
Priabonian38.0 - 33.9Denmark Emeljanovedusa gentarna, Lugeilangor elektrokleistis (123762)
Priabonian38.0 - 33.9United Kingdom (England) Hooleya indecisa, Hastites muiri (123960) Myndus wilmattae (129546)
Priabonian38.0 - 33.9Lithuania Protepiptera kaweckii, Ptychoptilum minor (113470)
Priabonian38.0 - 33.9Poland Cixius loculatus, Cixius vitreus, Pseudophana reticulata, Cixius insignis, Bothriobaltia pietrzeniukae, Cixius succineus, Cixius testudinarius, Cixius sieboldtii, Cixius fraternus, Cixius longirostris, Cixius gracilis, Poeocera nassata, Poeocera pristina (109625) Emeljanovedusa gentarna, Serafinana perperunae, Jantaritambia serafini, Perunus sudoviorum (123215) Glisachaemus jonasdamzeni, Worskaito stenexi, Patollo natangorum, Gedanotropis sontagae (110737) Positrona shcherbakovi, Kulickamia jantaris (120813) Protepiptera kaweckii (124952)
Priabonian38.0 - 33.9Russian Federation (Kaliningrad) Austris raffelis (123458) Autrimpus sambiorum (156261) Oliarus oligocenus (140082) Paratesum rasnitsyni, Protomenocria notata, Protepiptera kaweckii, Psycheona variegata, Psycheona striata, Ptychoptilum major (122591) Perunus bruziorum, Patollo aestiorum, Ptychogroehnia reducta (110735) Protomenocria notata (124951) Protomenocria notata, Protepiptera kaweckii (120610) Tritophania patruelis, Issus reticulatus (124073) Waghilde baltica, Cixidia christinae, Balticixius insignis, Angustachilus longirostris (123936)
Priabonian38.0 - 33.9Russian Federation Perunus sudoviorum (123419)
Priabonian38.0 - 33.9Ukraine Alicodoxa rasnitsyni (122997)
Chadronian37.2 - 33.9USA (Colorado) Florissantia elegans (106380) Nyctophylax uhleri, Nyctophylax vigil, Fulgora obticescens, Diaplegma haldemani, Diaplegma veterascens, Diaplegma abductum, Diaplegma venerabile, Diaplegma occultorum, Diaplegma ruinosum, Elidiptera regularis (106384)
Late/Upper Eocene37.2 - 33.9USA (Colorado) Florissantia elegans, Cixius proavus (122093)
Early/Lower Oligocene33.9 - 28.4France Cixius vitreus, Oligonila defectuosa, Cixius loculatus (121476)
Rupelian33.9 - 28.1Russian Federation Amagua fortis (122745)
MP 3028.4 - 23.03Germany Cixius sp., Hyalesthes rottensis (122010)
Chattian28.1 - 23.03France (Bouches-Du-Rhone) Asiraca sp. (122877)
Chattian28.1 - 23.03New Zealand (Otago) Fulgoromorpha indet. (191651)
Early/Lower Miocene23.03 - 15.97Mexico Oliarus sp. 1, Oliarus sp. 2, Nogodina chiapaneca, Eucanyra sp., Cenanges sp. 1, Cenanges sp. 2 (21977)
Early/Lower Miocene23.03 - 15.97Mexico (Chiapas) Copallinges chiapasensis (155387) Flatidae indet. (124810 151865) Mnemosyne sp. (140197) Oeclixius amphion (124471) Tonacatecutlius gibsoni, Cedusa baylissae (151418)
Burdigalian20.44 - 15.97China (Xizang) Cixius discretus (184254)
Burdigalian20.44 - 15.97Russian Federation Mundopoides cisthenaria (113705)
Burdigalian20.44 - 15.97China (Shandong) Fulgoropsis fusca, Oxycephala xiejiaheensis, Aphaena lithoecia (128550) Hylophylax erromena (130412) Limois shanwangensis (128552) Limois shanwangensis, Limois pardalis, Aphana lithoecia (128551) Oxycephala shanwangensis (128556) Oxycephala xiejiaheensis (128549) Ptomatosaiva endea (138946)
Burdigalian - Langhian20.44 - 13.82Dominican Republic Flatidae indet. (122982) Oeclidius browni (156229) Oliarus kulickae (155386) Oligocixia electrina (124578) Quizqueiplana alexbrowni (176579) Tainosia quisqueyae, Anticedusa dominicana, Oeclidius salaco, Quilessa stolida, Cedusa credula, Dysimia imprudens (123422) Thionia douglundbergi (124427)
Middle Miocene15.97 - 11.608Russian Federation Chloriona stavropolitana (125989) Dictyophara sp., Chanithus vishneviensis (125990) Mysidioides migdisovae (105571)
Turolian8.7 - 5.333France (Ardèche) Pterodictya ephemera (172761)
Blancan4.9 - 1.8USA (Texas) Areopodidae indet. (154909)
Holocene0.0117 - 0.0Tanzania Cixiidae indet., Derbidae indet., Issidae indet., Osaka sp. 2 (123418) Issidae indet., Pochazoides sp. 1, Pochazoides sp. 2, Pochazoides szwedoi, Acroprivesa msandarusi, Osaka sp. 1, Pocharica sp., Zoraida angolensis, Ricaniidae indet. (123991) Poeocera venulosa, Ricania multinervis (129861) Ricania equestris, Asiraca albipuncta (152160)