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Ceratopogonini

Insecta - Diptera - Ceratopogonidae

Synonyms
Synonymy list
YearName and author
1834Ceratopogonini Newman
1952Stilobezziini Wirth
1976Ceratopogonini Remm p. 345
1988Ceratopogonini Szadziewski p. 49
1988Stilobezziini Szadziewski p. 109
1996Ceratopogonini Szadziewski p. 63
1997Ceratopogonini Borkent and Wirth p. 86
2011Ceratopogonini Sontag and Szadziewski p. 784
2013Ceratopogonini Borkent p. 105
2016Ceratopogonini Stebner et al. p. 2
2017Ceratopogonini Stebner and Szadziewski p. 5
2019Ceratopogonini Szadziewski p. 526
2019Ceratopogonini Szadziewski and Sontag p. 660
2020Ceratopogonini Borkent and Dominiak p. 143
2020Ceratopogonini Szadziewski et al. p. 286
2021Ceratopogonini Peñalver et al. p. 426
2022Ceratopogonini Szadziewski et al. p. 2

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RankNameAuthor
kingdomAnimalia()
—Bilateria
—EubilateriaAx 1987
—Protostomia
—Ecdysozoa
—Panarthropoda
phylumArthropodaLatreille 1829
superclassHexapodaLatreille 1825
RankNameAuthor
classInsecta
orderDiptera
infraorderCulicomorpha
superfamilyChironomoideaMacquart 1838
familyCeratopogonidae
subfamilyCeratopogoninae
tribeCeratopogonini
tribeCeratopogonini

If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.

Tr. Ceratopogonini Newman 1834
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G. Alluaudomyia Kieffer 1913
+
†Alluaudomyia succinea Szadziewski 1988
G. Atyphohelea Borkent 1998
+
†Atyphohelea cothurnata Meunier 1904
+
Invalid names: Ceratopogon flagellus Meunier 1904 [synonym]
G. Austrohelea Wirth and Grogan 1988
G. Baeodasymyia Clastrier and Raccurt 1979
+
†Baeodasymyia dominicana Szadziewski and Grogan 1994
G. Baeohelea Wirth and Blanton 1970
G. †Brachycretacea Szadziewski 1996
+
G. Brachypogon Kieffer 1899
+
Subg. Brachypogon (Brachypogon) Kieffer 1899
+
†Brachypogon (Brachypogon) americanus Szadziewski and Grogan 1998
Subg. Brachypogon (Isohelea) Kieffer 1917
+
†Brachypogon (Isohelea) dominicanus Szadziewski and Grogan 1998
†Brachypogon (Isohelea) prominuloides Szadziewski and Grogan 1998
†Brachypogon balticus Szadziewski 1988
G. Camptopterohelea Wirth and Hubert 1960
+
†Camptopterohelea odora Stebner et al. 2017
G. Ceratoculicoides Wirth and Ratanaworabhan 1971
+
†Ceratoculicoides danicus Szadziewski 1988
G. †Ceratopalpomyia Szadziewski 1988
+
†Ceratopalpomyia eocenica Szadziewski 1988
G. Ceratopogon Meigen 1803
+
†Ceratopogon bitterfeldi Szadziewski 1993
†Ceratopogon ceranowiczi Szadziewski 1988
†Ceratopogon crypticus Szadziewski 1988
†Ceratopogon eminens Meunier 1904
†Ceratopogon forcipiformis Meunier 1904
+
Invalid names: Ceratopogon defectus Meunier 1904 [synonym]
†Ceratopogon gedanicus Szadziewski 1988
†Ceratopogon grogani Szadziewski 1988
†Ceratopogon hennigi Szadziewski 1988
†Ceratopogon kotejai Szadziewski 1993
†Ceratopogon margaritae Szadziewski 1988
†Ceratopogon miocaenicus Szadziewski 1993
†Ceratopogon nanalobus Borkent and Grogan 1995
†Ceratopogon paraeminens Borkent and Grogan 1995
†Ceratopogon piotrowskii Szadziewski 1988
†Ceratopogon pisinnus Borkent and Grogan 1995
†Ceratopogon remmicola Szadziewski 1988
†Ceratopogon remmicolus Szadziewski 1988
†Ceratopogon ritzkowskii Szadziewski 1988
†Ceratopogon spiniger Loew 1850
†Ceratopogon subeminens Szadziewski 1993
†Ceratopogon succinicola Szadziewski 1993
†Ceratopogon tertiaricus Szadziewski 1988
Invalid names: Ceratopogon (Ceratopogon) Meigen 1803 [invalid subgroup]
G. †Eohelea Petrunkevitch 1957
+
†Eohelea fossicola Szadziewski 1993
†Eohelea gedanica Szadziewski 1988
†Eohelea grogani Szadziewski 1988
†Eohelea indica Stebner and Szadziewski 2017
†Eohelea miocaenea Szadziewski 1993
†Eohelea petrunkevitchi Szadziewski 1985
†Eohelea sakhalinica Szadziewski 1990
†Eohelea sinuosa Meunier 1904
+
Invalid names: Eohelea stridulans Petrunkevitch 1957 [synonym]
G. Fanthamia de Meillon 1939
G. †Fossihelea Szadziewski 1988
+
†Fossihelea gracilitarsis Meunier 1904
†Fossihelea miocaenica Szadziewski 1993
G. †Gedanohelea Szadziewski 1988
+
†Gedanohelea fushunensis Stebner et al. 2016
†Gedanohelea gerdesorum Stebner and Szadziewski 2017
†Gedanohelea liaoningensis Stebner et al. 2016
†Gedanohelea loewi Szadziewski 1988
†Gedanohelea succinea Szadziewski 1988
†Gedanohelea wirthi Szadziewski 1988
G. †Mantohelea Szadziewski 1988
+
†Mantohelea gedanica Szadziewski 1988
†Mantohelea laca Meunier 1904
†Mantohelea sinica Stebner et al. 2016
G. Meunierohelea Szadziewski 1988
+
†Meunierohelea anglesensis Peñalver et al. 2021
†Meunierohelea borkenti Stebner and Szadziewski 2017
†Meunierohelea cambayana Stebner and Szadziewski 2017
†Meunierohelea fudalai Szadziewski et al. 2020
†Meunierohelea gedanicola Szadziewski 1988
†Meunierohelea miocaenica Szadziewski 1993
†Meunierohelea nielseni Szadziewski 1988
†Meunierohelea orientalis Stebner and Szadziewski 2017
†Meunierohelea wirthi Szadziewski 1988
Invalid names: Chimaerohelea Debenham 1988 [synonym]
G. †Monogedania Szadziewski et al. 2022
+
†Monogedania clunipes Loew 1850
G. Monohelea Kieffer 1917
G. Nannohelea Grogan and Wirth 1980
+
†Nannohelea eocenica Szadziewski 1988
†Nannohelea grogani Szadziewski 1988
G. †Nelohelea Szadziewski and Sontag 2019
+
†Nelohelea neli Szadziewski and Sontag 2019
G. †Palaeobrachypogon Borkent 1995
+
†Palaeobrachypogon remmi Borkent 1995
†Palaeobrachypogon taimyricus Szadziewski 1996
†Palaeobrachypogon vetus Borkent 1995
G. †Peronehelea Borkent 1995
+
†Peronehelea chrimikalydia Borkent 1995
†Peronehelea frigida Remm 1976
†Peronehelea sibirica Szadziewski 1996
G. Schizohelea Kieffer 1917
+
†Schizohelea baltica Szadziewski 1988
G. Serromyia Meigen 1818
+
†Serromyia alphea von Heyden 1870
+
Invalid names: Serromyia austera Statz 1944 [synonym], Serromyia colorata Statz 1944 [synonym], Serromyia spinofemorata Statz 1944 [synonym]
†Serromyia errata Szadziewski 2018
†Serromyia polonica Szadziewski 1988
†Serromyia ryszardi Borkent 1990
†Serromyia sinuosa Borkent 1990
†Serromyia spinigera Loew 1850
+
Invalid names: Ceratopogon elongatus Meunier 1904 [synonym]
†Serromyia succinea Szadziewski 1988
G. Stilobezzia Kieffer 1911
+
Subg. Stilobezzia (Acanthohelea) Kieffer 1917
+
†Stilobezzia (Acanthohelea) dominicana Szadziewski and Grogan 1998
+
Invalid names: Ceratopogon spinosus Meunier 1904 [synonym]
+
Invalid names: Stilobezzia goetghebueri Statz 1944 [synonym]
†Stilobezzia (Acanthohelea) wirthicola Szadziewski and Grogan 1998
+
Invalid names: Stilobezzia (Acanthohelea) succinea Szadziewski 1993 [replaced]
Subg. Stilobezzia (Stilobezzia) Kieffer 1911
+
†Stilobezzia (Stilobezzia) antilleana Szadziewski and Grogan 1998
†Stilobezzia kurthi Borkent 2000
†Stilobezzia pikei Borkent 2012
†Stilobezzia roggeroi Choufani and Nel 2013
G. †Wirthohelea Szadziewski 1988
+
†Wirthohelea trifida Szadziewski 1988
Invalid names: Stilobezziini Wirth 1952 [synonym]
Diagnosis
No diagnoses are available
Measurements
No measurements are available
Composition: chitinc
Environment: terrestrialc
Locomotion: actively mobilep
Created: 2017-04-17 09:56:18
Modified: 2017-04-17 09:56:18
Source: c = class, p = phylum
References: Bush and Bambach 2015, Kiessling 2004

Age range

Maximum range based only on fossils: base of the Turonian to the top of the Early/Lower Miocene or 93.90000 to 15.97000 Ma
Minimum age of oldest fossil (stem group age): 93.5 Ma

Collections (56 total)


Time interval Ma Country or state Original ID and collection number
Early/Lower Cenomanian99.6 - 93.5Myanmar Nelohelea neli (187929)
Turonian93.9 - 89.8USA (New Jersey) Palaeobrachypogon remmi, Palaeobrachypogon grandiforceps, Stilobezzia kurthi (110895) Palaeobrachypogon sp. (123016)
Early/Lower Santonian85.8 - 83.5France Metahelea roggeroi (152462)
Late/Upper Santonian85.8 - 83.5Russian Federation Ceratopogon macronyx, Ceratopogon frigidus, Washingtonhelea taimyrica, Baeohelea taimyrica, Peronehelea sibirica (35056)
Late/Upper Campanian83.5 - 70.6Canada (Manitoba) Ceratopogon aquilonius, Palaeobrachypogon remmi, Palaeobrachypogon vetus (122356) Palaeobrachypogon remmi, Palaeobrachypogon vetus, Peronehelea chrimikalydia (122354) Peronehelea chrimikalydia (122353)
Middle Campanian83.5 - 70.6Canada (Alberta) Palaeobrachypogon aquilonius, Peronehelea chrimikalydia (122388) Palaeobrachypogon remmi, Palaeobrachypogon vetus (122384) Palaeobrachypogon sp., Palaeobrachypogon vetus, Peronehelea sp., Peronehelea chrimikalydia, Stilobezzia pikei (122389) Palaeobrachypogon vetus, Peronehelea chrimikalydia (122383) Peronehelea chrimikalydia (34896 104932 122358)
Ypresian56.0 - 47.8India (Gujarat) Mantohelea sp., Stilobezzia sp., Brachypogon sp., Serromyia sp., Camptopterohelea odora, Eohelea indica, Gedanohelea gerdesorum, Meunierohelea cambayana, Meunierohelea borkenti, Meunierohelea orientalis (124463) Meunierohelea sp., Gedanohelea gerdesorum (183742)
Ypresian56.0 - 47.8China (Liaoning) Mantohelea sinica, Gedanohelea fushunensis, Gedanohelea liaoningensis (125934)
Lutetian47.8 - 41.3USA (Montana) Alluaudomyia sp. (224195)
Lutetian47.8 - 41.3Russian Federation Stilobezzia sp., Eohelea sakhalinica (107340)
Bartonian41.3 - 38.0Australia (Victoria) Monohelea sp., Meunierohelea anglesensis (224365)
Priabonian38.0 - 33.9United Kingdom (England) Stilobezzia sp. (21976 123959)
Priabonian38.0 - 33.9Poland Ceratopogon sp., Eohelea sinuosa, Meunierohelea sp., Serromyia sp. (154301) Ceratopogon sp., Fossihelea gracilitarsis, Meunierohelea sp. C, Ceratopogon clunipes, Serromyia sp., Neurohelea cothurnata, Ceratopogon spiniger, Brachypogon balticus, Brachypogon prominulus, Ceratopogon anomalicornis (109625) Ceratopogon sp., Fossihelea gracilitarsis, Monohelea clunipes, Brachypogon sp., Brachypogon balticus, Brachypogon polonicus, Brachypogon prominulus (153481) Ceratopogon sp., Mantohelea lacus, Mantohelea gedanica, Stilobezzia sp., Stilobezzia sp. C, Stilobezzia sp. D, Stilobezzia sp. E, Ceratopogonini indet., Ceratopogon eminens, Ceratopogon forcipiformis, Ceratopogon grogani, Ceratopogon hennigi, Ceratopogon margaritae, Ceratopogon tertiaricus, Eohelea gedanica, Eohelea grogani, Eohelea petrunkevitchi, Fossihelea gracilitarsis, Gedanohelea loewi, Meunierohelea sp., Meunierohelea sp. B, Meunierohelea sp. C, Meunierohelea nielseni, Monohelea clunipes, Nannohelea sp., Serromyia sp., Serromyia sp. A, Serromyia sp. B, Serromyia spinigera, Serromyia succinea, Stilobezzia falcata, Neurohelea cothurnata, Eohelea stridulans, Brachypogon sp., Brachypogon sp. A, Alluaudomyia succinea, Brachypogon balticus, Brachypogon eocenicus, Brachypogon gedanicus, Brachypogon henningseni, Brachypogon prominulus, Ceratopogon ceranowiczi, Ceratopogon crypticus, Ceratopogon gedanicus, Ceratopogon ritzkowskii, Gedanohelea succinea, Meunierohelea gedanicola, Monohelea baltica, Serromyia anomalicornis, Serromyia polonica, Serromyia ryszardi, Serromyia sinuosa, Wirthohelea trifida, Ceratopogon remmicolus, Eohelea sp. A, Eohelea sp. B (120813) Mantohelea gedanica, Monogedania clunipes, Schizohelea baltica (110737)
Priabonian38.0 - 33.9Russian Federation Ceratopogon lacus (123419)
Priabonian38.0 - 33.9Denmark Ceratopogon sp., Stilobezzia sp., Stilobezzia sp. A, Ceratopogonini indet., Ceratopogon eminens, Ceratopogon forcipiformis, Ceratopogon hennigi, Ceratopogon tertiaricus, Eohelea gedanica, Eohelea petrunkevitchi, Eohelea sinuosa, Fossihelea gracilitarsis, Meunierohelea sp., Meunierohelea sp. A, Meunierohelea nielseni, Monohelea clunipes, Nannohelea sp., Nannohelea grogani, Serromyia sp., Stilobezzia falcata, Neurohelea cothurnata, Eohelea stridulans, Brachypogon sp. A, Alluaudomyia sp. A, Alluaudomyia succinea, Brachypogon balticus, Brachypogon eocenicus, Brachypogon gedanicus, Brachypogon henningseni, Brachypogon prominulus, Ceratoculicoides danicus, Ceratopalpomyia eocenica, Ceratopogon crypticus, Ceratopogon piotrowskii, Ceratopogon ritzkowskii, Gedanohelea wirthi, Meunierohelea wirthi, Monohelea baltica, Nannohelea eocenica, Stilobezziini indet., Ceratopogon sp. B, Ceratopogon sp. A, Ceratopogon sp. C (123762)
Priabonian38.0 - 33.9Ukraine Ceratopogon sp. (122999) Ceratopogon sp., Mantohelea laca, Stilobezzia sp., Brachypogon sp., Brachypogon balticus, Brachypogon prominulus, Ceratopogon bitterfeldi, Ceratopogon forcipiformis, Ceratopogon grogani, Ceratopogon hennigi, Ceratopogon margaritae, Ceratopogon tertiaricus, Eohelea fossicola, Eohelea gedanica, Eohelea miocaenea, Eohelea petrunkevitchi, Eohelea sinuosa, Fossihelea sp., Fossihelea gracilitarsis, Fossihelea miocaenica, Gedanohelea loewi, Meunierohelea sp., Meunierohelea nielseni, Monohelea sp., Monohelea clunipes, Nannohelea sp., Nannohelea grogani, Serromyia sp., Serromyia spinigera, Serromyia succinea, Stilobezzia falcata, Neurohelea cothurnata (122997) Ceratopogon sp., Stilobezzia sp., Brachypogon sp., Brachypogon balticus, Brachypogon prominulus, Ceratopogon eminens, Ceratopogon forcipiformis, Eohelea gedanica, Eohelea grogani, Eohelea sinuosa, Meunierohelea sp., Monohelea clunipes, Stilobezzia falcata (113621) Ceratopogon sp., Stilobezzia sp., Brachypogon sp., Eohelea sinuosa, Meunierohelea sp., Monohelea clunipes (122996) Stilobezzia sp., Eohelea sinuosa, Meunierohelea sp. (122995)
Priabonian38.0 - 33.9Germany (Halle) Ceratopogon sp., Ceratopogon bitterfeldi, Ceratopogon forcipiformis, Ceratopogon hennigi, Eohelea fossicola, Eohelea miocaenea, Eohelea sinuosa, Fossihelea sp., Fossihelea miocaenica, Monohelea clunipes, Serromyia sp., Serromyia spinigera, Stilobezzia succinea, Brachypogon miocaenicus, Ceratopogon miocaenicus, Ceratopogon subeminens, Ceratopogon succinicolus, Chimaerohelea miocaenica, Serromyia alphea (129227) Ceratopogon sp., Mantohelea gedanica, Ceratopogon forcipiformis, Ceratopogon hennigi, Eohelea sinuosa, Fossihelea sp., Fossihelea sp. B, Monohelea clunipes, Nannohelea sp., Serromyia sp., Brachypogon sp., Brachypogon sp., Brachypogon prominulus, Ceratopogon kotejai, Stilobezzia falcata, Stilobezzia kutscheri, Stilobezzia saxonica, Chimaerohelea nielseni, Serromyia alphea (129228)
Priabonian38.0 - 33.9Russian Federation (Kaliningrad) Ceratopogon lacus, Ceratopogon prominulus, Ceratopogon eminens, Ceratopogon forcipiformis, Fossihelea sp. A, Ceratopogon gracilitarsis, Ceratopogon clunipes, Ceratopogon falcatus, Ceratopogon cothurnatus, Ceratopogon defectus, Ceratopogon flagellus, Ceratopogon elongatus, Ceratopogon spiniger, Ceratopogon sinuosus, Ceratopogon spinosus (123927) Ceratopogon sp., Ceratopogonini indet., Ceratopogon tertiaricus, Monohelea sp. A, Monohelea clunipes, Brachypogon balticus, Brachypogon prominulus (123993) Ceratopogon sp., Eohelea sinuosa, Monohelea clunipes, Brachypogon sp., Alluaudomyia succinea (123221) Ceratopogon sp., Stilobezzia sp. A, Stilobezzia sp. B, Ceratopogonini indet., Ceratopogon forcipiformis, Eohelea sinuosa, Fossihelea gracilitarsis, Monohelea clunipes, Nannohelea sp., Serromyia sp., Brachypogon prominulus (123962)
MP 3028.4 - 23.03Germany Ceratopogon alpheus (123945)
MP 3028.4 - 23.03Germany (Nordrhein-Westfalen) Tetragoneura veterana (138696)
Chattian28.1 - 23.03New Zealand (Otago) Austrohelea sp. (191651)
Chattian28.1 - 23.03Germany (North Rhine-Westphalia) Ceratopogon pupa 13, Ceratopogon pupa 12, Ceratopogon pupa 11, Ceratopogon pupa 10, Ceratopogon pupa 9, Ceratopogon pupa 8, Ceratopogon pupa 7, Ceratopogon pupa 6, Ceratopogon pupa 5, Ceratopogon pupa 4, Ceratopogon pupa 3, Ceratopogon pupa 2, Ceratopogon pupa 1, Ceratopogon sp., Serromyia sp., Serromyia austera, Serromyia colorata, Serromyia spinofemorata, Stilobezzia goetghebueri, Stilobezzia veterana (122010)
Chattian28.1 - 23.03France (Provence-Alpes-Côte d'Azur) Ceratopogon sp. (122488)
Early/Lower Miocene23.03 - 15.97Mexico Brachypogon sp. (21977)
Early/Lower Miocene23.03 - 15.97Mexico (Chiapas) Brachypogon sp. (124474) Nannohelea sp. (140212 140217)
Burdigalian - Langhian20.44 - 13.82Dominican Republic Brachypogon dominicanus (125970) Brachypogon dominicanus, Stilobezzia dominicana, Stilobezzia antilleana, Baeodasymyia dominicana, Brachypogon sp. (122982) Meunierohelea fudalai (124427) Nannohelea sp. A, Brachypogon dominicanus, Brachypogon prominuloides, Stilobezzia dominicana, Stilobezzia antilleana, Baeodasymyia dominicana, Brachypogon sp., Stilobezzia sp. A (122473) Nannohelea sp., Brachypogon americanus, Brachypogon sp., Brachypogon sp. (123857) Stilobezzia dominicana, Brachypogon sp. (140835)