Basic info Taxonomic history Classification Relationships
Morphology Ecology and taphonomy External Literature Search Age range and collections


Synonymy list
YearName and author
2004Embryophyta Crane et al.

Is something missing? Join the Paleobiology Database and enter the data


If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.

No diagnoses are available
No measurements are available
No ecological data are available

Age range

Maximum range based only on fossils: base of the Homerian to the top of the Holocene or 430.50000 to 0.00000 Ma
Minimum age of oldest fossil (stem group age): 427.4 Ma

Collections (6895 total)

Oldest occurrences

Time interval Ma Country or state Original ID and collection number
Llandovery - Ludlow443.4 - 423.0Peru Harringtonia acutiplicata (25078)
Silurian443.4 - 419.2Czech Republic (Bohemia) Cooksonia sp. (20759)
Silurian443.4 - 419.2Norway (Svalbard) Leiotriletes parvus, Granulatisporites muninensis, Emphanisporites minutus, Calamospora nigrata (20839)
Wenlock433.4 - 427.4Argentina Harringtonia sp. (25096)
Homerian430.5 - 427.4Ireland Cooksonia sp., Cooksonia cambrensis, Cooksonia pertoni, Cooksonia hemisphaerica (10594)
Gorstian427.4 - 425.6United Kingdom (Wales) Cooksonia sp., Cooksonia cambrensis, Cooksonia pertoni (10661)
Ludlow427.4 - 423.0Australia (Victoria) Hedeia gracilis (53660) Zosterophyllales indet., Hedeia sp. (20754)
Gorstian - Pridoli427.4 - 419.2Bolivia (Tarija dept.) Cooksonia caledonica, Cooksonia sp., Cooksonia pertoni, Cooksonia hemisphaerica (11158)
Gorstian - Pridoli427.4 - 419.2Bolivia (Tarija Dept.) Cooksonia pertoni, Cooksonia hemisphaerica (11172)
Ludlow - Pridoli427.4 - 419.2Canada (Nunavut) Distichophytum sp. (10599)
Ludfordian - Pridoli425.6 - 419.2Ukraine Cooksonia pertonii (14147)
Ludfordian - Pridoli425.6 - 419.2United Kingdom (Wales) Cooksonia sp. (10635)
Ludfordian - Silurian425.6 - 419.2United Kingdom (Wales) Cooksonia caledonica, Cooksonia downtonensis, Cooksonia hemisphaerica (10634)
Pridoli423.0 - 419.2USA (New York) Cooksonia hemisphaerica (53827) Cooksonia sp. (20664 26087)
Pridoli423.0 - 419.2China (Xinjiang) Thallites indet., Junggaria spinosa (10931)
Pridoli423.0 - 419.2United Kingdom (Wales) Cooksonia cambrensis, Cooksonia caledonica, Cooksonia sp., Cooksonia pertoni, Cooksonia hemisphaerica (10666)
Pridoli423.0 - 419.2Czech Republic Cooksonia sp. (14043 14131 52698) Cooksonia sp., Cooksonia hemisphaerica (14045)
Pridoli423.0 - 419.2Ukraine (Podolia) Cooksonia sp. 1, Cooksonia sp. 2, Cooksonia pertoni, Cooksonia hemisphaerica (52702)
Pridoli423.0 - 419.2Poland (Lysogory Region) Cooksonia sp. (52667)
Pridoli423.0 - 419.2Libya (Hammedah Basin) Cooksonia sp. (53669)