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Plesiochelys etalloni

Reptilia - Testudines - Plesiochelyidae

Taxonomy
Emys etalloni was named by Pictet and Humbert (1857). Its type specimen is MAJ 2005-11-1, a shell (a shell missing a large part of the carapace medially.), and it is a 3D body fossil. Its type locality is Forêt de Lect, pres de Moirans (Lect Forest), which is in a Tithonian marine horizon in France.

It was recombined as Plesiochelys etalloni by Rütimeyer (1873), Bräm (1965), Gaffney (1975), Perea et al. (2014), Anquetin et al. (2014) and Puntener et al. (2017).

Synonyms
Synonymy list
YearName and author
1857Emys etalloni Pictet and Humbert p. 1
1869Stylemys hannoverana Maack p. 7 figs. Plate 23, fig. 6; Plate 25, figs 31-34, 36, 39-40; Plate 26, figs 41-43, 45, 47; Plate 27, figs 48-5
1873Plesiochelys sanctaverenae Rutimeyer
1873Plesiochelys solodurensis Rütimeyer p. 50
1873Plesiochelys etalloni Rütimeyer p. 72
1873Plesiochelys sanctaeverenae Rütimeyer p. 80
1873Plesiochelys langii Rütimeyer p. 82
1878Plesiochelys hannoverana Portis p. 9 figs. Plate 2, figs 6-7
1889Plesiochelys solodurensis Lydekker pp. 197-198
1889Plesiochelys hannoverana Lydekker p. 198
1902Plesiochelys solodurensis Hay p. 439
1965Plesiochelys etalloni Bräm p. 60
1965Plesiochelys solodurensis Bräm p. 81
1965Plesiochelys sanctaverenae Bräm p. 126
1975Plesiochelys etalloni Gaffney p. 7
1996Plesiochelys solodurensis Lapparent de Broin et al. pp. 553-554
2008Plesiochelys solodurensis Danilov and Parham
2008Plesiochelys solodurensis Pérez-García et al.
2013Plesiochelys solodurensis Pérez-García and Murelaga
2014Plesiochelys etalloni Anquetin et al.
2014Plesiochelys etalloni Perea et al.
2014Plesiochelys solodurensis Puntener et al.
2017Plesiochelys etalloni Puntener et al.

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RankNameAuthor
kingdomAnimalia()
Triploblastica
Nephrozoa
Deuterostomia
phylumChordataHaeckel 1847
subphylumVertebrata
superclassGnathostomata
Osteichthyes()
Sarcopterygii
subclassDipnotetrapodomorpha(Nelson 2006)
subclassTetrapodomorpha()
Tetrapoda()
Reptiliomorpha
RankNameAuthor
Anthracosauria
Batrachosauria()
Cotylosauria()
Amniota
Sauropsida
classReptilia
Testudinata(Oppel 1811)
orderTestudinesBatsch 1788
Thalassochelydia
familyPlesiochelyidaeRütimeyer 1873
genusPlesiochelys
speciesetalloni(Pictet and Humbert 1857)

If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.

Diagnosis
ReferenceDiagnosis
E. S. Gaffney 1975Member of the Plesiochelyidae (sensu Gaffney, present paper) known from nearly all the skeleton, distinguishable from Plesiochelys planiceps by the following features: lingual ridge of maxilla usually lower than in P. planiceps, anterior portion of lingual ridge on lower jaw curving anteriorly (as opposed to medially in P. planiceps), at level of vomer-premaxilla suture the distance between lingual ridges of maxillae is narrower than in P. planiceps.
C. Puntener et al. 2017Plesiochelys etalloni differs from other Plesiochelys spp. in a more extensive flooring of the cavum acustico–jugulare by the pterygoid, the complete ossification of the pila prootica, and a narrow, slit-like foramen nervi trigemini. In addition, Plesiochelys etalloni differs from Plesiochelys planiceps in a smaller size, a lower lingual ridge on the maxilla, a narrower distance between the lingual ridges of the maxilla at the level of the pterygoid–vomer suture, a more rounded foramen palatinum posterius, a parietal–quadrate contact posterior to the foramen nervi trigemini, a less developed processus trochlearis oticum, a superficial canalis caroticus internus often remaining partly open ventrally, a reduced contribution of the exoccipital to the condylus occipitalis, and the anterior portion of the lingual ridge on the dentary curving medially, and from Plesiochelys bigleri in a higher temporal skull roof, a deeper pterygoid fossa, a more developed processus trochlearis oticum, an anterior foramen nervi abducentis opening more posteriorly relative to the base of the processus clinoideus, foramina anterius canalis carotici cerebralis opening almost vertically below the dorsum sellae and usually more closely set, a processus paroccipitalis extending mainly posteriorly, an increased neural and costal bone thickness, the presence of epiplastral bulbs, and a more rounded or pointed anterior margin of the anterior plastral lobe.