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Iocheirata

Arachnida - Pseudoscorpiones

Taxonomy
Iocheirata was named by Harvey (1992). It is extant.

It was assigned to Chelonethi by Dunlop et al. (2013); and to Homosphyronida by Benavides et al. (2019).

Synonymy list
YearName and author
1992Iocheirata Harvey
2013Iocheirata Dunlop et al. p. 52
2019Iocheirata Benavides et al.

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RankNameAuthor
kingdomAnimalia()
—Bilateria
—EubilateriaAx 1987
—Protostomia
—Ecdysozoa
—Panarthropoda
phylumArthropodaLatreille 1829
—Deuteropoda
subphylumChelicerataHeymons 1901
—EuchelicerataWeygoldt and Paulus 1979
RankNameAuthor
—Prosomapoda
—Planaterga
—Dekatriata
—SclerophorataKamenz et al. 2011
classArachnidaLamarck 1801
—HaplocnemataBorner 1904
orderPseudoscorpionesLatreille 1817
—HomosphyronidaChamberlin 1929
suborderIocheirataHarvey 1992
suborderIocheirataHarvey 1992

If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.

Subor. Iocheirata Harvey 1992
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Infraor. Hemictenata Harvey 1992
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Superfm. Neobisioidea Chamberlin 1930
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Fm. Ideoroncidae Chamberlin 1930
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G. †Proalbiorix Geißler et al. 2021
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†Proalbiorix gracilis Geißler et al. 2021
Fm. Neobisiidae Chamberlin 1930
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G. †Electrobisium Cockerell 1917
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†Electrobisium acutum Cockerell 1917
G. Microcreagris Balzan 1892
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†Microcreagris koellnerorum Schawaller 1978
G. Neobisium Chamberlin 1930
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†Neobisium exstinctum Beier 1955
†Neobisium henderickxi Judson 2003
G. Roncus Koch 1873
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†Roncus succineus Beier 1955
Invalid names: Bochicidae Chamberlin 1930 [empty], Gymnobisiidae Beier 1947 [empty], Hyidae Chamberlin 1930 [empty], Parahyidae Harvey 1992 [empty], Syarinidae Chamberlin 1930 [empty]
Infraor. Panctenata Harvey 1992
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Superfm. Cheiridioidea Hansen 1894
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Fm. Cheiridiidae Hansen 1894
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G. Cheiridium Menge 1855
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†Cheiridium hartmanni Menge 1854
G. Cryptocheiridium Chamberlin 1931
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†Cryptocheiridium antiquum Schawaller 1981
Fm. Pseudochiridiidae Chamberlin 1923
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G. Pseudochiridium With 1906
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†Pseudochiridium lindae Judson 2007
Superfm. Cheliferoidea Risso 1826
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Fm. Atemnidae Kishida 1929
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Subfm. Atemninae Kishida 1929
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G. Paratemnoides Harvey 1991
G. †Progonatemnus Beier 1955
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Fm. Cheliferidae Risso 1826
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Subfm. Cheliferinae Risso 1827
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Invalid names: Cheliferini Risso 1827 [empty]
G. †Dichela Menge 1854
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†Dichela berendtii Menge 1854
†Dichela gracilis Beier 1937
†Dichela granulatus Beier 1937
Invalid names: Oligochelifer Beier 1937 [synonym]
G. †Electrochelifer Beier 1937
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†Electrochelifer mengei Beier 1937
G. †Heurtaultia Judson 2009
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G. Parawithius Chamberlin 1931
Subfm. Philomaoriinae Chamberlin 1931
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G. Philomaoria Chamberlin 1931
G. †Pycnochelifer Beier 1937
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†Pycnochelifer kleemanni Koch and Berendt 1854
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Invalid names: Obisium rathkii Koch and Berendt 1854 [synonym]
G. †Trachychelifer Hong 1983
Fm. Chernetidae Menge 1855
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G. Americhernes Muchmore 1976
G. Byrsochernes Beier 1959
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†Byrsochernes maatiatus Riquelme et al. 2014
Invalid names: Mayachernes Riquelme et al. 2014 [synonym]
G. Lustrochernes Beier 1932
G. †Oligochernes Beier 1937
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†Oligochernes bachofeni Beier 1937
†Oligochernes wigandi Menge 1854
G. Pachychernes Beier 1932
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†Pachychernes effossus Schawaller 1980
Fm. Withiidae Chamberlin 1931
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G. †Beierowithius Mahnert 1979
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G. Chelifer Geoffroy 1762
G. Dolichowithius Beier 1932
G. Withius Kew 1911
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†Withius eucarpus Dalman 1826
Superfm. Garypinoidea Daday 1888
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Fm. Garypinidae Daday 1888
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G. Amblyolpium Simon 1898
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†Amblyolpium burmiticum Cockerell 1920
Subfm. Garypininae Daday 1889
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G. †Ajkagarypinus Novák et al. 2023
G. Garypinus Daday 1888
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†Garypinus electri Beier 1937
Invalid names: Larcidae Chamberlin 1930 [empty]
Superfm. Garypoidea Simon 1879
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Fm. Garypidae Simon 1879
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G. †Garypus Koch 1873
Fm. Geogarypidae Chamberlin 1930
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G. Geogarypus Chamberlin 1930
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†Geogarypus gorskii Henderickx 2005
†Geogarypus major Beier 1937
Invalid names: Hesperolpiidae Chamberlin 1930 [empty], Menthidae Chamberlin 1930 [empty], Olpiidae Banks 1895 [empty]
Superfm. Sternophoroidea Chamberlin 1923
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Fm. Sternophoridae Chamberlin 1923
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G. Idiogaryops Hoff 1963
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Diagnosis
No diagnoses are available
Measurements
No measurements are available
Composition: chitinc
Grouping: solitaryc
Environment: terrestrialc
Locomotion: actively mobilec
Created: 2017-04-17 09:55:58
Modified: 2017-04-17 09:55:58
Source: c = class
Reference: Bush and Bambach 2015

Age range

Maximum range based only on fossils: base of the Late/Upper Albian to the top of the Holocene or 105.30000 to 0.00000 Ma
Minimum age of oldest fossil (stem group age): 99.6 Ma

Collections (24 total)


Time interval Ma Country or state Original ID and collection number
Late/Upper Albian105.3 - 99.6France (Poitou-Charentes) Cheliferidae indet., Heurtaultia rossiorum (99114)
Early/Lower Cenomanian99.6 - 93.5Myanmar Electrobisium acutum, Garypus burmiticus (123020) Proalbiorix gracilis (172900)
Santonian86.3 - 83.6Hungary Ajkagarypinus stephani (224054)
Middle Campanian83.5 - 70.6Canada (Alberta) Chernetidae indet. (34896)
Priabonian38.0 - 33.9Poland Chelifer kleemanni, Obisium rathkii (109625) Geogarypus gorskii (124519) Neobisium sp., Chelifer kleemanni, Obisium rathkii, Chelifer wigandi, Chelifer hartmanni, Dichela berendtii, Obisium sieboldtii, Garypinus electri, Geogarypus macrodactylus, Geogarypus major, Oligochelifer gracilis, Oligochelifer serratidentatus, Oligochelifer granulatus, Electrochelifer mengei (120811)
Priabonian38.0 - 33.9Russian Federation (Kaliningrad) Chernetidae indet., Electrochelifer bachofeni, Electrochelifer rapulitarsatus, Cheiridium hartmanni, Oligochernes bachofeni, Oligochelifer berendtii (123870) Microcreagris koellneri, Cheiridium sp. (120814) Neobisium henderickxi (123936) Pycnochelifer sp., Cheiridium hartmanni, Neobisium exstinctum, Roncus succineus, Geogarypus macrodactylus, Progonatemnus succineus, Oligowithius sieboldi, Electrochelifer balticus (123771)
Priabonian38.0 - 33.9Ukraine Geogarypus gorskii (154710)
Priabonian38.0 - 33.9Russian Federation Garypinus electri (123419)
Early/Lower Miocene23.03 - 15.97Mexico (Chiapas) Cheliferini indet. (178679) Mayachernes maatiatus (163578)
Early/Lower Miocene23.03 - 15.97Mexico Chernetidae indet., Lustrochernes sp. 1 (21977) Paratemnoides sp., Lustrochernes sp. 2 (146607)
Burdigalian20.44 - 15.97New Zealand (Otago) Philomaoria sp. (191650)
Burdigalian - Langhian20.44 - 13.82Dominican Republic Idiogaryops pumilus (123422) Pachychernes sp., Pachychernes effossus, Americhernes sp., Cryptocheiridium antiquum, Parawithius sp. (122474) Pseudochiridium lindae (156229)
Holocene0.0117 - 0.0Colombia (Santander) Paratemnoides nidificator, Pachychernes subrobustus (147613)
Holocene0.0117 - 0.0Tanzania Chelifer eucarpus (152160)