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Pantetrapulmonata

Arachnida

Synonymy list
YearName and author
2007Pantetrapulmonata Shultz
2014Pantetrapulmonata Garwood and Dunlop p. 22

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RankNameAuthor
kingdomAnimalia()
Bilateria
EubilateriaAx 1987
Protostomia
Ecdysozoa
Panarthropoda
phylumArthropodaLatreille 1829
Deuteropoda
subphylumChelicerataHeymons 1901
RankNameAuthor
EuchelicerataWeygoldt and Paulus 1979
Prosomapoda
Planaterga
Dekatriata
SclerophorataKamenz et al. 2011
classArachnidaLamarck 1801
Pantetrapulmonata
Pantetrapulmonata

If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.

Unr. Pantetrapulmonata Shultz 2007
show all | hide all
G. †Ecchosis Selden and Shear 1991
+
Ecchosis pulchribothrium Selden and Shear 1991
Subcl. Tetrapulmonata Schultz 1990
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G. †Eocteniza Pocock 1911
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Eocteniza silvicola Pocock 1911
G. †Protocteniza Petrunkevitch 1949
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Protocteniza britannica Petrunkevitch 1949
G. †Rakovnicia Kušta 1885
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Rakovnicia antiqua Kušta 1885
Unr. Schizotarsata Shultz 2007
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Or. †Haptopoda Pocock 1911
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Fm. †Plesiosironidae Pocock 1911
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G. †Plesiosiro Pocock 1911
Unr. Pedipalpi Borner 1904
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Or. Amblypygi Thorell 1883 [whip spider]
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Subor. Euamblypygi Weygoldt 1996
Subor. Paleoamblypygi Weygoldt 1996
G. †Thelyphrynus Petrunkevitch 1913
Invalid names: Phrynus fossilis Keferstein 1834 [nomen dubium]
Or. Thelyphonida Latreille 1804 [whip scorpion]
+
G. †Burmathelyphonia Wunderlich 2015
G. †Parageralinura Tetlie and Dunlop 2008
G. †Proschizomus Dunlop and Horrocks 1996
Or. Schizomida Petrunkevitch 1945 [short-tailed whip-scorpion]
Fm. Thelyphonidae Lucas 1835
Or. Uropygi Thorell 1882 [whip scorpion]
Unr. Serikodiastida Garwood and Dunlop 2014
+
Or. Araneae Clerck 1757 [spider]
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G. †Amphiclotho Gourret 1887
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Amphiclotho breviuscula Gourret 1887
G. †Amphithomisus Gourret 1887
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Amphithomisus barbatus Gourret 1887
G. †Araneaovoius Dunlop and Braddy 2011
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Araneaovoius columbiae Scudder 1878
G. †Argyrarachne Selden 1999
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Argyrarachne solitus Selden 1999
G. †Atocatle Feldmann et al. 1998
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Atocatle ranulfoi Feldmann et al. 1998
G. †Cercidiella Gourret 1887
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Cercidiella aquisextana Gourret 1887
G. †Clubionella Gourret 1887
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Clubionella antiqua Gourret 1887
G. †Eresoides Gourret 1887
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Eresoides orbicularis Gourret 1887
G. †Herselioides Gourret 1887
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Subor. Mesothelae
+
Fm. †Arthrolycosidae Fritsch 1904
Fm. †Arthromygalidae Petrunkevitch 1923
Fm. †Burmathelidae Wunderlich 2017
Fm. †Cretaceothelidae Wunderlich 2017
G. †Eolycosa Kušta 1886
+
Invalid names: Kustaria Petrunkevitch 1953 [synonym], Scudderia Kušta 1889 [synonym]
Fm. †Eomesothelidae Wunderlich 2019
Fm. †Palaeothelidae Selden 2021
Fm. †Parvithelidae Wunderlich 2017
Invalid names: Liphistiidae Thorell 1869 [empty]
G. Mygale Latreille 1802
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Mygale ambigua Gourret 1887
G. †Opisthophylax Menge 1856
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Opisthophylax exarata Menge 1856
Subor. Opisthothelae Pocock 1892
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Infraor. Araneomorphae Smith 1902
G. †Eoatypus McCook 1888
Infraor. Mygalomorphae Pocock 1892
G. †Palaranea Frič 1873
+
+
Invalid names: Arthrolycosa palaranea Fritsch 1904 [objective synonym]
G. †Paralycosa Dunlop and Jekel 2009
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Paralycosa attiformis Gourret 1887
Invalid names: Protolycosa Gourret 1887 [replaced]
G. †Prodysdera Gourret 1887
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Prodysdera intermedia Gourret 1887
G. †Protochersis Gourret 1887
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Protochersis spinosus Gourret 1887
G. †Protolachesis Gourret 1887
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Protolachesis annulata Gourret 1887
G. †Pseudothomisus Gourret 1887
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G. †Pyritaranea Fritsch 1899
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Pyritaranea tubifera Fritsch 1899
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Invalid names: Eopholcus pedatus Fritsch 1904 [synonym]
Invalid names: Eopholcus Fritsch 1904 [synonym]
G. †Schellenbergia Heer 1864
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G. †Timeropus Thorell 1891
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Timeropus hersiliformis Gourret 1887
Invalid names: Lycosoides Gourret 1887 [replaced]
Invalid names: Amaurobius faustus Koch and Berendt 1854 [nomen dubium], Amaurobius rimosus Koch and Berendt 1854 [nomen dubium], Amaurobius spinimanus Menge 1854 [nomen nudum], Anatone Menge 1854 [nomen nudum], Anatone marginata Menge 1854 [nomen nudum], Anatone spinipes Menge 1854 [nomen nudum], Archaea incomta Menge 1854 [nomen nudum], Archaea sphinx Menge 1854 [nomen nudum], Athera Menge 1854 [nomen nudum], Athera exilis Menge 1854 [nomen nudum], Auximus fossilis Petrunkevitch 1950 [nomen dubium], Auximus succini Petrunkevitch 1942 [nomen dubium], Clubiona eseri Heer 1864 [nomen nudum], Clubiona latifrons Menge 1854 [nomen nudum], Clythia Koch and Berendt 1854 [nomen dubium], Clythia alma Koch and Berendt 1854 [nomen dubium], Clythia gracilenta Menge 1854 [nomen nudum], Clythia leptocarena Menge 1854 [nomen nudum], Corynitoides Dunlop and Jekel 2009 [nomen dubium], Corynitoides spinosa Menge 1854 [nomen dubium], Corynitoides undulata Menge 1854 [nomen dubium], Dielacata Menge 1854 [nomen nudum], Dielacata superba Menge 1854 [nomen nudum], Drassus oblongus Menge 1854 [nomen nudum], Dysdera glabrata Menge 1854 [nomen nudum], Dysdera hippopodium Menge 1854 [nomen nudum], Dysdera scobiculata Menge 1854 [nomen nudum], Eocryphoeca distincta Petrunkevitch 1950 [nomen dubium], Eocryphoeca fossilis Petrunkevitch 1942 [nomen dubium], Eometa aberrans Petrunkevitch 1958 [nomen dubium], Eometa robusta Petrunkevitch 1958 [nomen dubium], Epeira eogena Menge 1854 [nomen nudum], Epeiridion Menge 1854 [nomen nudum], Epeiridion femoratum Menge 1854 [nomen nudum], Erithus Menge 1854 [nomen nudum], Erithus applanatus Menge 1854 [nomen nudum], Ero coronata Menge 1854 [nomen nudum], Ero exculpta Menge 1854 [nomen nudum], Ero quadripunctata Menge 1854 [nomen nudum], Ero setulosa Koch and Berendt 1854 [nomen dubium], Eyukselus Özdikmen 2007 [nomen nudum], Eyukselus argutus Menge 1854 [nomen nudum], Eyukselus felinus Menge 1854 [nomen nudum], Eyukselus griseus Menge 1854 [nomen nudum], Eyukselus latifrons Menge 1854 [nomen nudum], Eyukselus pumilus Menge 1854 [nomen nudum], Heteromma intersecta Menge 1856 [nomen nudum], Idmonia Menge 1854 [nomen nudum], Idmonia virginea Menge 1854 [nomen nudum], Laterigradae Thorell [empty], Melanophora lepida Menge 1854 [nomen nudum], Melanophora nitida Menge 1854 [nomen nudum], Memoratrix Petrunkevitch 1942 [nomen dubium], Memoratrix rydei Petrunkevitch 1942 [nomen dubium], Micaria ovata Menge 1854 [nomen nudum], Micaria squamata Menge 1854 [nomen nudum], Micaria tenuis Menge 1854 [nomen nudum], Micryphantes globulus Menge 1854 [nomen nudum], Micryphantes turritus Menge 1854 [nomen nudum], Miropholcus Petrunkevitch 1942 [nomen dubium], Miropholcus heteropus Petrunkevitch 1942 [nomen dubium], Mizalia truncata Menge 1854 [nomen nudum], Ocypete angustifrons Menge 1854 [nomen nudum], Ocypete marginata Menge 1854 [nomen nudum], Onca Menge 1854 [nomen nudum], Onca lepida Menge 1854 [nomen nudum], Onca pumila Menge 1854 [nomen nudum], Perturbator Petrunkevitch 1971 [nomen dubium], Perturbator corniger Petrunkevitch 1971 [nomen dubium], Phalangopus Menge 1854 [nomen dubium], Phalangopus subtilis Menge 1854 [nomen dubium], Philodromus griseus Menge 1856 [nomen nudum], Philodromus redrogradus Menge 1854 [nomen nudum], Philodromus reptans Menge 1854 [nomen nudum], Pyritaraneidae Petrunkevitch 1953 [invalid subgroup], Pythonissa bipunctata Menge 1854 [nomen nudum], Pythonissa discophora Menge 1854 [nomen nudum], Pythonissa glabra Menge 1854 [nomen nudum], Pythonissa villosa Menge 1854 [nomen nudum], Retitelariae Thorell [empty], Saltigradae Latreille [empty], Segestria exarata Menge 1854 [nomen nudum], Segestria undulata Menge 1854 [nomen nudum], Siga Menge 1854 [nomen nudum], Siga crinita Menge 1854 [nomen nudum], Spheconia Menge 1854 [nomen nudum], Spheconia brevipes Menge 1854 [nomen nudum], Syphax hirtus Menge 1854 [nomen nudum], Thomisus matutinus Menge 1856 [nomen nudum], Thyelia pectinata Menge 1854 [nomen nudum], Thyelia spinosa Menge 1854 [nomen nudum], Tubitelariae Thorell [empty], Zilla cornumana Menge 1854 [nomen nudum], Zilla spinipalpa Menge 1854 [nomen nudum]
Or. Araneida Clerck 1757
+
Subor. †Chimerarachnida Wunderlich 2018
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Fm. †Chimerarachnidae Wunderlich 2019
G. †Idmonarachne Garwood et al. 2016
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Idmonarachne brasieri Garwood et al. 2016
Or. †Uraraneida Selden and Shear 2008 [spider]
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G. †Attercopus Selden and Shear 1991
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Attercopus fimbriunguis Shear et al. 1987
Fm. †Permarachnidae Eskov and Selden 2005
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G. †Permarachne Eskov and Selden 2005
G. †Xenarachne Dunlop and Poschmann 1997
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Xenarachne willwerathensis Dunlop and Poschmann 1997
Or. †Trigonotarbida Petrunkevitch 1949
+
G. †Aenigmatarbus Poschmann et al. 2016
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Aenigmatarbus rasteli Poschmann et al. 2016
Fm. †Anthracomartidae Haase 1890
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G. †Anthracomartus Karsch 1882
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Anthracomartus bohemica Fritsch 1895
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Invalid names: Arthrolycosa salticoides Fritsch 1899 [synonym], Promygale rotundata Fritsch 1899 [synonym]
Anthracomartus elegans Fritsch 1899
Anthracomartus granulatus Fritsch 1904
Anthracomartus hindi Pocock 1911
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Invalid names: Cleptomartus planus Petrunkevitch 1949 [synonym], Cryptomartus meyeri Guthörl 1964 [synonym]
Anthracomartus janae Opluštil 1986
Anthracomartus kustae Petrunkevitch 1953
Anthracomartus minor Kušta 1885
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Invalid names: Anthracomartus socius Kušta 1889 [synonym]
Anthracomartus nyranensis Petrunkevitch 1953
Anthracomartus palatinus von Ammon 1901
Anthracomartus priesti Pocock 1911
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Invalid names: Anthracomartus denuiti Pruvost 1922 [synonym], Cleptomartus plautus Petrunkevitch 1949 [synonym]
Anthracomartus triangularis Petrunkevitch 1913
Anthracomartus trilobitus Scudder 1884
Invalid names: Brachylycosa Fritsch 1904 [synonym], Cleptomartus Petrunkevitch 1949 [synonym], Coryphomartus Petrunkevitch 1945 [synonym], Cryptomartus Petrunkevitch 1945 [synonym], Oomartus Petrunkevitch 1953 [synonym], Perneria Fritsch 1904 [synonym], Pleomartus Petrunkevitch 1945 [synonym], Promygale Fritsch 1899 [synonym]
G. †Brachypyge Woodward 1878
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Brachypyge carbonis Woodward 1878
G. †Maiocercus Pocock 1911
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Maiocercus celticus Pocock 1902
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Invalid names: Maiocercus orbicularis Gill 1911 [synonym]
Invalid names: Brachypygidae Pocock 1911 [synonym], Coryphomartidae Petrunkevitch 1945 [synonym], Eurymartus Matthew 1895 [nomen dubium], Eurymartus latus Matthew 1895 [nomen dubium], Pleomartidae Petrunkevitch 1945 [synonym], Promygalidae Fritsch 1904 [synonym]
G. †Anthracophrynus Andrée 1913
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Fm. †Anthracosironidae Pocock 1903
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G. †Anthracosiro Pocock 1903
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Anthracosiro fritschii Pocock 1903
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Invalid names: Anthracosiro elongatus Waterlot 1934 [synonym]
Anthracosiro woodwardi Pocock 1903
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Invalid names: Anthracosiro corsini Pruvost 1926 [synonym], Anthracosiro latipes Gill 1909 [synonym]
G. †Arianrhoda Dunlop and Selden 2004
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Arianrhoda bennetti Dunlop and Selden 2004
G. †Vratislavia Fritsch 1904
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Vratislavia silesiaca Römer 1878
Fm. †Aphantomartidae Petrunkevitch 1945
+
G. †Alkenia Størmer 1970
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Alkenia mirabilis Størmer 1970
G. †Aphantomartus Pocock 1911
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Aphantomartus areolatus Pocock 1911
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Invalid names: Aphantomartus pococki Pruvost 1912 [synonym], Eophrynus waechteri Guthörl 1938 [synonym], Trigonomartus dorlodoti Pruvost 1930 [synonym]
Aphantomartus ilfeldicus Scharf 1924
Aphantomartus pustulatus Scudder 1884
+
Invalid names: Cleptomartus ploetzensis Simon 1971 [synonym], Kreischeria villeti Pruvost 1912 [synonym]
Invalid names: Phrynomartus Petrunkevitch 1945 [synonym], Trigonomartus Petrunkevitch 1913 [synonym]
G. †Doubravatarbus Hradská et al. 2022
+
Doubravatarbus krafti Hradská et al. 2022
Invalid names: Trigonomartidae Petrunkevitch 1945 [synonym]
Fm. †Archaeomartidae Poschmann and Dunlop 2010
+
G. †Archaeomartus Størmer 1970
+
Archaeomartus levis Størmer 1970
+
Invalid names: Archaeomartus tuberculatus Størmer 1970 [synonym]
G. †Areomartus Petrunkevitch 1913
+
Areomartus ovatus Petrunkevitch 1913
Fm. †Eophrynidae Karsch 1882
+
G. †Eophrynus Woodward 1871
+
Eophrynus prestvicii Buckland 1837
Eophrynus udus Brauckmann et al. 1985
Euphrynus salmi Stur 1877
Invalid names: Duma Giebel 1856 [nomen oblitum]
G. †Nyranytarbus Harvey and Selden 1995
+
Nyranytarbus hofmanni Fritsch 1899
Nyranytarbus longipes Fritsch 1899
Invalid names: Hemiphrynus Fritsch 1899 [replaced]
G. †Petrovicia Fritsch 1904
+
Petrovicia proditoria Fritsch 1904
G. †Planomartus Petrunkevitch 1953
+
Planomartus krejcii Kušta 1884
+
Invalid names: Anthracomartus affinis Kušta 1885 [synonym]
G. †Pleophrynus Petrunkevitch 1945
+
Pleophrynus ensifer Petrunkevitch 1945
Pleophrynus hawesi Dunlop et al. 2014
Pleophrynus verrucosus Pocock 1911
+
Invalid names: Eophrynus jugatus Ambrose and Romano 1972 [synonym], Eophrynus warei Dix and Pringle 1930 [synonym]
Invalid names: Acrokreischeria Petrunkevitch 1953 [synonym]
G. †Pocononia Petrunkevitch 1953
+
Pocononia whitei Ewing 1930
G. †Somaspidion Jux 1982
+
G. †Stenotrogulus Fritsch 1904
+
Stenotrogulus salmii Stur 1877
+
Invalid names: Cyclotrogulus sturii Fritsch 1904 [synonym]
Invalid names: Cyclotrogulus Fritsch 1904 [synonym], Pseudoeophrynus Pribyl 1958 [synonym]
Invalid names: Hemiphrynidae Fritsch 1904 [synonym], Pleophrynidae Petrunkevitch 1945 [synonym]
Fm. †Kreischeriidae Haase 1890
+
G. †Anzinia Petrunkevitch 1953
+
Anzinia thevenini Pruvost 1919
G. †Gondwanarachne Pinto and Hunicken 1980
+
Gondwanarachne argentinensis Pinto and Hünicken 1980
G. †Hemikreischeria Fritsch 1904
+
Hemikreischeria geinitzi Thevenin 1902
+
Invalid names: Hemikreischeria thevenini Fritsch 1904 [objective synonym]
G. †Kreischeria Geinitz 1882
+
Kreischeria bohemica Fritsch 1895
Kreischeria wiedei Geinitz 1882
G. †Pseudokreischeria Petrunkevitch 1953
+
+
Invalid names: Eophrynus varius Petrunkevitch 1949 [synonym]
Fm. †Lissomartidae Dunlop 1995
+
G. †Lissomartus Petrunkevitch 1949
+
Lissomartus carbonarius Petrunkevitch 1913
Lissomartus schucherti Petrunkevitch 1913
G. †Namurotarbus Poschmann and Dunlop 2010
+
Namurotarbus roessleri Dunlop and Brauckmann 2006
Fm. †Palaeocharinidae Hirst 1923
+
G. †Aculeatarbus Shear et al. 1987
+
Aculeatarbus depressus Shear et al. 1987
G. †Gelasinotarbus Shear et al. 1987
+
Gelasinotarbus bifidus Shear et al. 1987
Gelasinotarbus bonamoae Shear et al. 1987
Gelasinotarbus fimbriunguis Shear et al. 1987
Gelasinotarbus heptops Shear et al. 1987
Gelasinotarbus reticulatus Shear et al. 1987
G. †Gigantocharinus Shear 2000
+
G. †Gilboarachne Shear et al. 1987
+
Gilboarachne griersoni Shear et al. 1987
G. †Palaeocharinus Hirst 1923
+
Palaeocharinus calmani Hirst 1923
Palaeocharinus hornei Hirst 1923
Palaeocharinus tuberculatus Fayers et al. 2005
Invalid names: Palaeocharinoides Hirst 1923 [synonym]
G. †Spinocharinus Poschmann and Dunlop 2011
+
Spinocharinus steinmeyeri Poschmann and Dunlop 2011
G. †Palaeocteniza Hirst 1923
+
G. †Palaeotarbus Dunlop 1999
+
Palaeotarbus jerami Dunlop 1996
Invalid names: Eotarbus Dunlop 1996 [replaced]
G. †Permotarbus Dunlop and Rößler 2013
+
Permotarbus schuberti Dunlop and Rößler 2013
Fm. †Trigonotarbidae Petrunkevitch 1949
+
G. †Trigonotarbus Pocock 1911
+
Trigonotarbus arnoldi Petrunkevitch 1955
Trigonotarbus carbonarius Petrunkevitch 1913
Trigonotarbus johnsoni Pocock 1911
Trigonotarbus stoermeri Schultka 1991
G. †Tynecotarbus Hradská and Dunlop 2013
+
Tynecotarbus tichaveki Hradská and Dunlop 2013
Invalid names: Anthracomarti Karsch 1882 [replaced], Anthracomartus hageni Goldenberg 1873 [nomen dubium], Elaverimartus Petrunkevitch 1953 [nomen dubium], Elaverimartus pococki Petrunkevitch 1953 [nomen dubium], Eophrynus scharfi Scharf 1924 [nomen dubium], Eurymarti Matthew 1895 [synonym], Eurymartidae Matthew 1895 [invalid subgroup], Eurymartus spinulosus Matthew 1895 [nomen dubium], Termes buchi Goldenberg 1873 [nomen dubium], Trigonomartus woodruffi Scudder 1893 [nomen dubium]
Diagnosis
No diagnoses are available
Measurements
No measurements are available
Composition: chitinc
Grouping: solitaryc
Environment: terrestrialc
Locomotion: actively mobilec
Created: 2017-04-17 09:55:58
Modified: 2017-04-17 09:55:58
Source: c = class
Reference: Bush and Bambach 2015

Age range

Maximum range based only on fossils: base of the Pridoli to the top of the Holocene or 423.00000 to 0.00000 Ma
Minimum age of oldest fossil (stem group age): 419.2 Ma

Collections (472 total)

Oldest occurrences

Time interval Ma Country or state Original ID and collection number
Pridoli423.0 - 419.2United Kingdom (England) Trigonotarbida indet. (11128)
Lochkovian419.2 - 410.8United Kingdom Arianrhoda bennetti (153478)
Pragian410.8 - 407.6Germany Archaeomartus levis, Spinocharinus steinmeyeri (155760) Trigonotarbida indet. (152107)
Pragian410.8 - 407.6United Kingdom (Scotland) Palaeocharinus kidstoni (10721) Palaeocharinus sp. (153498) Palaeocharinus sp., Palaeocharinus scourfieldi, Palaeocharinoides hornei, Palaeocharinus calmani, Palaeocharinus rhyniensis (172777) Palaeocharinus sp., Palaeocteniza crassipes (172778) Palaeocharinus tuberculatus (153492)
Early/Lower Emsian409.1 - 402.5Germany Alkenia mirabilis (155774) Archaeomartus levis, Alkenia mirabilis (155757) Trigonotarbus stoermeri (172343) Xenarachne willwerathensis (155775)
Early/Lower Emsian409.1 - 402.5Germany (Alken) Archaeomartus tuberculatus, Alkenia mirabilis (27049)
Early/Lower Givetian388.1 - 383.7USA (New York) Attercopus fimbriungus (28295) Gilboarachne griersoni, Gelasinotarbus reticulatus, Gelasinotarbus fimbriunguis, Gelasinotarbus bonamoae (153480)
Givetian387.7 - 382.7USA (New York) Gilboarachne griersoni, Aculeatarbus depressus, Gelasinotarbus reticulatus, Gelasinotarbus bifidus, Gelasinotarbus bonamoae, Gelasinotarbus heptops, Attercopus fimbriunguis, Ecchosis pulchribothrium (28246) Gilboarachne griersoni, Gelasinotarbus reticulatus, Gelasinotarbus fimbriunguis, Gelasinotarbus bonamoae (21172)
Late/Upper Famennian364.7 - 360.7USA (Pennsylvania) Gigantocharinus szatmaryi (35044)
Chadian346.7 - 340.0USA (Virginia) Trigonomartus whitei (155455)
Asbian340.0 - 336.0Germany Cryptomartus sp. (156764)
Brigantian336.0 - 330.9Germany Anthracomartus sp., Trigonomartus pustulatus (153594)
Brigantian - Serpukhovian336.0 - 323.2Canada (Nova Scotia) Anthracomartus sp. (155663)
Arnsbergian327.0 - 323.7Czech Republic Euphrynus salmi (38971)
Morrowan323.2 - 318.6USA (Arkansas) Anthracomartus trilobitus (153485)
Alportian322.8 - 321.8Austria Aphantomartus pustulatus (153488)
Marsdenian321.3 - 320.0Germany (Ruhr) Eophrynus udus (155259) Prothelyphonus naufragus (110939) Trigonotarbus johnsoni (152121)
Westphalian A318.7 - 316.9United Kingdom Aphantomartus areolatus (146813) Nemastomoides elaveris (146854)
Westphalian A318.7 - 316.9Belgium Anthracomartus denuiti (155639) Brachypyge carbonis (155637) Brachypyge orbicularis (155638) Trigonomartus dorlodoti (155635)
Westphalian A318.7 - 316.9United Kingdom (Staffordshire) Anthracosiro woodwardi (155452)