PBDB Taxon

Basic info	Taxonomic history	Classification	Included Taxa
Morphology	Ecology and taphonomy	External Literature Search	Age range and collections

Pancrustacea

Taxonomy

Pancrustacea was named by Zrzavý and Štys (1997). It is extant.

It was assigned to Crustacea by Schweitzer et al. (2020); and to Mandibulata by Nielsen (2012), Izquierdo-López and Caron (2022).

Subtaxa

Altocrustacea, Bredocaris, Crustacea, Hymenostraca, Skara, Wujicaris, Yicaris

Sister taxa

Altocrustacea, Hymenostraca

Synonyms

Maxillopoda was named by Dahl (1956). It is extant.

It was considered an invalid subgroup of Crustacea by Oakley et al. (2013).

It was assigned to Crustacea by Martin and Davis (2001).
Crustacea was named by Brünnich (1772). It is extant.

It was reranked as the class Crustacea by Latreille (1802), Phleger (1933), Mason (1935), Riccio (1952); it was reranked as the unranked clade Crustacea by Martens et al. (2005); it was considered an invalid subgroup of Pancrustacea by Lozano-Fernandez et al. (2019).

It was assigned to Animalia by Latreille (1802); to Branchiata by Mason (1935); to Arthropoda by Martin (1904), Phleger (1933), Riccio (1952), Bertels (1968), Martens et al. (2005), Carignano (2012); and to Pancrustacea by Nielsen (2012).

Synonymy list

Year	Name and author
1772	Crustacea Brünnich
1802	Crustacea Latreille p. 13
1904	Crustacea Martin p. 94
1933	Crustacea Phleger, Jr. p. 11
1935	Crustacea Mason p. 102
1952	Crustacea Riccio p. 29
1956	Maxillopoda Dahl
1968	Crustacea Bertels
1997	Pancrustacea Zrzavý and Štys
2001	Maxillopoda Martin and Davis p. 58
2005	Crustacea Martens et al.
2012	Crustacea Carignano
2012	Pancrustacea Nielsen p. 244 figs. Fig. 44.2
2012	Crustacea Nielsen p. 245
2020	Pancrustacea Schweitzer et al. p. 246
2022	Pancrustacea Izquierdo-López and Caron pp. 220933-14

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Rank	Name	Author
kingdom	Animalia	()
—	Bilateria
—	Eubilateria	Ax 1987
—	Protostomia
—	Ecdysozoa	Aguinaldo et al. 1997
—	Panarthropoda

Rank	Name	Author
phylum	Arthropoda	Latreille 1829
—	Deuteropoda
—	Mandibulata
—	Pancrustacea	Zrzavý and Štys 1997
—	Pancrustacea	Zrzavý and Štys 1997

If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.

Unr. Pancrustacea Zrzavý 1997

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Unr. Altocrustacea Regier et al. 2010

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Supercl. Allotriocarida Oakley et al. 2013

Unr. Athalassocarida Lozano-Fernandez et al. 2019

Cl. Branchiopoda Latreille 1817 [branchiopod]

Or. Anostraca Sars 1867

Fm. †Ketmeniidae Novozhilov 1957

Subcl. Phyllopoda

Invalid names: Sarsostraca Tasch 1969 [empty]

Unr. Labiocarida Schwentner et al. 2017

Supercl. Hexapoda Latreille 1825

Cl. Remipedia Yager 1981

Cl. Cephalocarida Sanders 1955

Or. †Brachypoda Birshteyn 1960

G. †Dala Muller 1983

G. †Rehbachiella Mueller 1983

†Rehbachiella kinnekullensis Mueller 1983

G. †Bredocaris Mueller 1983

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†Bredocaris admirabilis Mueller 1983

Unr. †Hymenostraca Rolfe 1969

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G. †Branchiocaris Briggs 1976

†Branchiocaris malongensis Luo et al. 2008

†Branchiocaris pretiosa Resser 1929

†Branchiocaris xundianensis Luo et al. 2008

†Branchiocaris yiliangensis Luo et al. 2008

†Branchiocaris yunnanensis Hou 1987

G. †Ercaicunia Luo et al. 1999

†Ercaicunia multinodosa Luo et al. 1999

G. †Hymenocaris Salter 1853

†Hymenocaris dubia Resser and Howell 1938

Supercl. Multicrustacea Regier et al. 2010

Infracl. †Halicyna Gall and Grauvogel 1967

Or. †Cyclida Glaessner 1928

Fm. †Alsasuacaridae van Bakel et al. 2011

Fm. †Americlidae Dzik 2008

G. †Anhelkocephalon Bill 1914

Fm. †Cyclidae Packard 1885

Fm. †Halicynidae Gall and Grauvogel 1967

Fm. †Hemitrochiscidae Trauth 1918

Fm. †Schraminidae Dzik 2008

Cl. Hexanauplia Oakley et al. 2013

Subcl. Copepoda Milne-Edwards,Milne-Edwards 1840 [copepod]

G. Cletocamptus Schmankevitch 1875

Or. Harpacticoida Sars 1903

G. †Kabatarina

Infracl. Neocopepoda Huys and Boxshall 1991

Infracl. Progymnoplea Lang 1948

G. Sphaerothylacus Sluiter 1884

Cl. Malacostraca Latreille 1802

G. †Amphipeltis Salter 1863

†Amphipeltis paradoxus Salter 1863

Subcl. Eumalacostraca Grobben 1892

Unr. Caridoida Hessler 1982

G. †Circularva Shcherbakov and Ponomarenko 2023

Superor. †Eocarida Brooks 1962

Infracl. Hoplocarida Calman 1904

G. †Joanellia Schram 1979

Fm. †Mesoprosopidae Glaessner 1928

G. †Schramocaris Clark et al. 2016

Invalid names: Percarida Calman 1904 [empty]

G. †Naranda Muenster 1842

Subcl. Phyllocarida Packard 1879

Or. †Archaeostraca Claus 1888

G. †Coreocaris Kobayashi 1937

G. †Dioxycaris Gurich 1929

Subor. †Discinocarina Clarke 1900

G. †Galenocaris Wells 1944

Or. †Hoplostraca Schram 1973

Or. Leptostraca Claus 1880

G. Nebaliella Thiele 1904

G. †Nothozoe Barrande 1868

G. †Phasganocaris Novak 1886

G. †Protocimex

G. †Saccocaris Salter 1868

G. †Shafferia

Invalid names: Aristocaris [invalid subgroup]

Invalid names: Belotelsonidea Schram 1981 [empty], Waterstonellidea [empty]

G. †Nereocaris Legg et al. 2012

†Nereocaris exilis Legg et al. 2012

Supercl. Oligostraca Zrzavý et al. 1997

Cl. Ichthyostraca Zrzavý et al. 1997

Subcl. Branchiura Thorell 1864

Invalid names: Arguloida Yamaguti 1963 [empty]

Subcl. Pentastomida Diesing 1836

G. †Boeckelericambria

Or. Cephalobaenida Heymons 1935

G. †Haffnericambria

G. †Heymonsicambria

Invalid names: Porocephalida Heymons 1935 [empty]

Invalid names: Mystacocarida Pennak and Zinn 1943 [empty]

Cl. Ostracoda Latreille 1802 [ostracod]

G. †Acanthomeridion Hou et al. 1989

G. †Acrossula

G. †Anabarochilina

G. †Anfedatia

G. †Aristaluta

G. †Aurilia

G. †Australopsis

G. †Bennviaspis

Or. †Beyrichiocopa Pokorný 1954

Subor. †Binodicopa Schallreuter 1972

G. †Biaurina

G. †Borodiella

G. †Boucekites

G. †Brachycytheropteron

G. †Brasacypris Krömmelbein 1965

†Brasacypris fulfaroi Dias-Brito et al. 2001

†Brasacypris morigerata Musacchio 1973

G. †Bullaluta

Fm. †Buregiidae Polenova 1953

G. †Buregia

G. †Bursulella Jones 1961

G. †Cambraechmina

G. †Carnarvonia

G. †Cedocamia

G. †Craspedopyxion

G. †Crescenticythere

G. Cyclocypris Brady and Norman 1889

G. Cyprinotus Brady 1886

Cyprinotus salinus Brady 1868

G. †Cytherina

†Cytherina spinosa Hall 1852

Invalid names: Gibba spinosa Fuchs 1920 [synonym]

G. Cytheropsis Sars 1866

G. †Damaoella

G. †Dhrumaella Manivit and Vaslet 1987

G. †Dorispina

G. †Duibianella

G. †Eucytheridea Bronstein 1930

G. †Fengtaiella

G. †Flemingia

G. †Flemingopsis

G. †Galeopsis Hinz-Schallreuter 1993

G. †Gelidicella

G. †Gerdalia Belousova 1961

G. †Gerubiella

G. †Gladioscutum

G. Glossicythereis Benson 1974

G. †Hameaschmidtella

G. †Hemiaechminoides

G. Heterocypris Claus 1893

G. †Huanghuaella

G. †Hupehella

G. †Ilmenoindivisia

G. †Infracytheropteron

G. †Ivaria

G. †Jixinglingella

G. †Koilocythere Deltel 1963

G. †Kuiperiana Bassiouni 1962

†Kuiperiana latissima Brady 1878

G. †Kuresaaria

G. †Liagnshanella

Or. †Limbatulocopida Abushik 1990

Fm. †Dolborellidae Melnikova 1976

G. †Lublinella

G. †Ludvigsenites

G. †Maghrebeis

G. †Majiashanella

G. †Mannosocmia

G. †Marginotimorites

G. †Microcosmia

G. †Moorea

Subcl. Myodocopa Sars 1866

Or. Halocyprida Dana 1853

Or. †Leperditicopida Scott 1961

Invalid names: Leperditiocopa Pokorny 1953 [synonym]

Or. Myodocopida Sars 1866

Superor. Myodocopomorpha Kozur 1972

Or. †Cypridinida Skogsberg 1920

G. †Navicularina

G. Neoatlanticythere Benson 1974

G. †Netzkaina

G. †Oepikaluta

Subor. †Paleocopa Henningsmoen 1953

Superfm. †Beyrichiacea Matthew 1886

Invalid names: Leperditiacea Bassler and Kellett 1934 [empty]

G. †Paleocythere

G. †Paleodarwinula Molostovskaya 1990

G. †Parashergoldopsis

G. †Paratsunyiella

G. †Phasoia

G. †Plesidielymella

Subcl. Podocopa Sars 1865

G. †Cetollina Shi 2002

G. †Eumiraculum Chen 1987

Or. Palaeocopida Henningsmoen 1953

Invalid names: Palaeocopamorphes Abushik 1990 [synonym]

Or. Platycopida Sars 1866

Invalid names: Platycopamorphes Abushik 1990 [synonym]

Or. Podocopida Sars 1866

Superor. Podocopamorpha Kozur 1972

Or. †Metacopida Sylvester-Bradley 1961

Superor. Podocopomorpha Sars 1865

Or. Punciocopida Schallreuter 1968

G. †Ponthocythere Dubowsky 1939

G. †Prasuchonella Molostovskaya 1980

G. †Pseudonodellina

G. †Pseudoperissocytheridea

G. Puriana Coryell and Fields 1953

†Puriana carolinensis Hazel 1983

Puriana fissispinata Benson and Coleman 1963

Invalid names: Favella Coryell and Fields 1937 [synonym]

G. †Reginacypris

G. †Renicytherura Gruendel 1981

G. †Reticulocambria

G. †Reversoparaparchites

G. †Rockallia Whatley et al. 1978

G. †Schallreuterina

G. †Septadella Smith and Stubblefield 1933

†Septadella jackmanae Smith and Stubblefield 1933

G. †Shergoldopsis

G. †Sigillium

G. †Spytihnevites

G. †Suchonellina Spizharsky 1937

†Suchonellina globosa Jones 1862

†Suchonellina stricta Jones 1894

G. †Supratoarcina

G. †Svealuta

G. †Tatariella

G. †Tethysia Donze 1975

G. †Tetrasulcata

G. †Torusilina

G. †Trapezisylthere

G. †Utsatia

G. †Uvonhactia

G. †Venzavella

G. †Vitella

Superfm. †Volganelloidea Mandelstam 1956

Fm. †Volganellidae Mandelstam 1956

G. †Yaxianella

G. †Yulinella

G. †Zerqacythere Basha 1980

Invalid names: Beichuania Xie 1989 [nomen nudum], Beyrichiida Pokorny 1953 [empty], Cuselina Ammon 1910 [nomen nudum], Darwinulacea Brady and Norman 1889 [empty], Golcondella [nomen dubium]

G. †Tokummia Aria and Caron 2017

†Tokummia katalepsis Aria and Caron 2017

Invalid names: Hymenocarididae Haeckel 1896 [empty]

G. †Skara Mueller 1983

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†Skara anulata Mueller 1983

G. †Wujicaris Zhang et al. 2010

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†Wujicaris muelleri Zhang et al. 2010

G. †Yicaris Zhang et al. 2007

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†Yicaris dianensis Zhang et al. 2007

Invalid names: Crustacea Brünnich 1772 [invalid subgroup]

Diagnosis

No diagnoses are available

Measurements

No measurements are available

Locomotion:

actively mobilep

Created:

2004-02-29 09:38:49

Modified:

2004-02-29 11:38:49

Source: p = phylum

Reference: Kiessling 2004

Age range

Maximum range based only on fossils: base of the Ediacaran to the top of the Holocene or 635.00000 to 0.00000 Ma
Minimum age of oldest fossil (stem group age): 538.8 Ma

Collections (17170 total)

Time interval	Ma	Country or state	Original ID and collection number
Oldest occurrences
Ediacaran	635.0 - 538.8	Mexico	Platella sp. (10662)
Meishucunian	530.7 - 521.0	China (Shaanxi)	Ostracoda indet. (6851)
Atdabanian	521.0 - 516.0	Australia (South Australia)	Phasoia spicata (155233)
Atdabanian	521.0 - 516.0	Germany (Brandenburg)	Diplostraca indet. (24190)
Atdabanian	521.0 - 516.0	China (Yunnan)	Branchiocaris yunnanensis (90163 90166 90494) Canadaspis laevigata (96403 96405 96416) Chuandianella ovata (86847 90126 90129 90149 90150 90151 90152 90161 90162) Chuandianella ovata, Branchiocaris yunnanensis (86845 86846 90138 90164) Chuandianella ovata, Canadaspis laevigata (90165 96402) Chuandianella ovata, Canadaspis laevigata, Branchiocaris yunnanensis (86851) Chuandianella ovata, Ercaicunia multinodosa (90137) Waptia sp. (86931) Waptia sp., Branchiocaris sp. (86929) Waptia sp., Branchiocaris sp., Acanthomeridion sp. (86930) Waptia sp., Canadaspis sp., Acanthomeridion sp. (86933) Waptia sp., Chuandianella ovata, Branchiocaris yunnanensis, Ercaicunia multinodosa (68083)
Nangaoian	521.0 - 514.5	China (Yunnan)	Waptia ovata (132328) Waptia ovata, Branchiocaris yunnanensis (132329 194140 194162 194163) Waptia ovata, Branchiocaris yunnanensis, Ercaicunia multinodosa (194141 194150 194153) Waptia ovata, Canadaspis laevigata, Branchiocaris yunnanensis (132327 194160 194161) Waptia ovata, Canadaspis laevigata, Branchiocaris yunnanensis, Ercaicunia multinodosa (132330 194143 194155)
Nangaoian	521.0 - 514.5	China (Hubai)	Branchiocaris sp. (219687)
Branchian	516.2 - 506.5	United Kingdom (England)	Hesslandona n. sp. B (79939)
Botomian	516.0 - 512.1	China (Yunnan)	Branchiocaris sp. (90353)
Botomian - Lenian	516.0 - 508.2	Russian Federation (Transbaikalia)	Ostracoda indet. (10007)
Botomian - Lenian	516.0 - 508.2	Russian Federation	Duibianella sp. (10018)
Dyeran	515.3 - 509.7	Canada (Newfoundland and Labrador)	Leperditicopa sp. (9900)
Dyeran	515.3 - 509.7	USA (Nevada)	Canadaspis sp. (10560)
Dyeran	515.3 - 509.7	Canada (Northwest Territories)	Branchiopoda indet. (6850)
Dyeran	515.3 - 509.7	USA (Pennsylvania)	Crustacea indet. (226194) Roddyia typa, Crustacea indet., Hymenocaris dubia (226192) Roddyia typa, Hymenocaris sp. (192)
Dyeran	515.3 - 509.7	USA (California)	Ostracoda indet. (219693)
Duyunian	514.5 - 506.5	China (Hubei)	Cycloconchoides venustus (201164)
Duyunian	514.5 - 506.5	China (Guizhou)	Waptia sp., Canadaspis sp. (237148)
Delamaran	509.7 - 505.3	USA (Utah)	Canadaspis perfecta, Branchiocaris pretiosa, Dioxycaris argenta (77218)
Delamaran	509.7 - 505.3	USA (Nevada)	Canadaspis perfecta (205001)